longtext: 4WY5-pdb

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HEADER    HYDROLASE                               15-NOV-14   4WY5
TITLE     STRUCTURAL ANALYSIS OF TWO FUNGAL ESTERASES FROM RHIZOMUCOR MIEHEI
TITLE    2 EXPLAINING THEIR SUBSTRATE SPECIFICITY
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ESTERASE;
COMPND   3 CHAIN: A, B;
COMPND   4 EC: 3.1.1.1;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: RHIZOMUCOR MIEHEI;
SOURCE   3 ORGANISM_TAXID: 4839;
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   5 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   6 EXPRESSION_SYSTEM_PLASMID: PET30A
KEYWDS    RHIZOMUCOR MIEHEI, ESTERASE, SUBSTRATE SPECIFICITY, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    Z.QIN,S.YANG,X.DUAN,Q.YAN,Z.JIANG
REVDAT   1   01-JUL-15 4WY5    0
JRNL        AUTH   Z.QIN,S.YANG,X.DUAN,Q.YAN,Z.JIANG
JRNL        TITL   STRUCTURAL INSIGHTS INTO THE SUBSTRATE SPECIFICITY OF TWO
JRNL        TITL 2 ESTERASES FROM THE THERMOPHILIC RHIZOMUCOR MIEHEI
JRNL        REF    J.LIPID RES.
JRNL        REFN                   ISSN 0022-2275
REMARK   2
REMARK   2 RESOLUTION.    2.43 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE: 1.8.2_1309)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.43
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 31.80
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.960
REMARK   3   COMPLETENESS FOR RANGE        (%) : 97.3
REMARK   3   NUMBER OF REFLECTIONS             : 26738
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.181
REMARK   3   R VALUE            (WORKING SET) : 0.178
REMARK   3   FREE R VALUE                     : 0.226
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 7.450
REMARK   3   FREE R VALUE TEST SET COUNT      : 1992
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 31.8550 -  5.8007    0.99     1806   144  0.1705 0.1879
REMARK   3     2  5.8007 -  4.6087    0.99     1800   143  0.1557 0.1935
REMARK   3     3  4.6087 -  4.0274    0.99     1793   141  0.1430 0.1896
REMARK   3     4  4.0274 -  3.6597    0.99     1840   147  0.1502 0.2057
REMARK   3     5  3.6597 -  3.3978    0.99     1745   146  0.1800 0.2103
REMARK   3     6  3.3978 -  3.1976    0.99     1812   143  0.1904 0.2532
REMARK   3     7  3.1976 -  3.0376    0.99     1799   151  0.1987 0.2244
REMARK   3     8  3.0376 -  2.9055    0.98     1791   143  0.2049 0.2489
REMARK   3     9  2.9055 -  2.7937    0.98     1759   142  0.1969 0.2880
REMARK   3    10  2.7937 -  2.6973    0.98     1760   143  0.2004 0.2523
REMARK   3    11  2.6973 -  2.6130    0.98     1843   147  0.1979 0.2702
REMARK   3    12  2.6130 -  2.5384    0.98     1769   140  0.2090 0.2943
REMARK   3    13  2.5384 -  2.4716    0.97     1718   140  0.2131 0.3155
REMARK   3    14  2.4716 -  2.4113    0.82     1511   122  0.2513 0.3216
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.280
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 24.560
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.010           4981
REMARK   3   ANGLE     :  1.242           6803
REMARK   3   CHIRALITY :  0.050            806
REMARK   3   PLANARITY :  0.007            872
REMARK   3   DIHEDRAL  : 13.816           1781
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: SF FILE CONTAINS FRIEDEL PAIRS UNDER
REMARK   3  I/F_MINUS AND I/F_PLUS COLUMNS.
REMARK   4
REMARK   4 4WY5 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 17-NOV-14.
REMARK 100 THE DEPOSITION ID IS D_1000204643.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 15-AUG-13
REMARK 200  TEMPERATURE           (KELVIN) : 90
REMARK 200  PH                             : 8.0
REMARK 200  NUMBER OF CRYSTALS USED        : NULL
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : BSRF
REMARK 200  BEAMLINE                       : 3W1A
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9791
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MAR CCD 165 MM
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : NULL
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 26738
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.430
REMARK 200  RESOLUTION RANGE LOW       (A) : 31.800
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 94.0
REMARK 200  DATA REDUNDANCY                : 2.000
REMARK 200  R MERGE                    (I) : 0.07900
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 19.9200
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: AUTOMAR
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: THE ENTRY CONTAINS FRIEDEL PAIRS IN F_PLUS/MINUS COLUMNS.
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 51.74
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.55
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 25% (W/V)PEG3350; 0.2M (NH4)2SO4; 0.1M
REMARK 280  MES BUFFER PH6.0, PH 8.0, VAPOR DIFFUSION, SITTING DROP,
REMARK 280  TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 2390 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 25980 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -35.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     THR A     2
REMARK 465     VAL A     3
REMARK 465     ALA A   324
REMARK 465     LEU A   325
REMARK 465     GLU A   326
REMARK 465     HIS A   327
REMARK 465     HIS A   328
REMARK 465     HIS A   329
REMARK 465     HIS A   330
REMARK 465     HIS A   331
REMARK 465     HIS A   332
REMARK 465     MET B     1
REMARK 465     THR B     2
REMARK 465     VAL B     3
REMARK 465     ALA B   324
REMARK 465     LEU B   325
REMARK 465     GLU B   326
REMARK 465     HIS B   327
REMARK 465     HIS B   328
REMARK 465     HIS B   329
REMARK 465     HIS B   330
REMARK 465     HIS B   331
REMARK 465     HIS B   332
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     LYS A  20    CG   CD   CE   NZ
REMARK 470     GLU A  21    CG   CD   OE1  OE2
REMARK 470     ARG A  22    CG   CD   NE   CZ   NH1  NH2
REMARK 470     SER A  36    OG
REMARK 470     SER A  37    OG
REMARK 470     ARG A  40    CG   CD   NE   CZ   NH1  NH2
REMARK 470     SER B  36    OG
REMARK 470     SER B  37    OG
REMARK 470     ASN B  45    CG   OD1  ND2
REMARK 470     LYS B  47    CG   CD   CE   NZ
REMARK 470     GLU B 249    CG   CD   OE1  OE2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    VAL A    32     CB   SER A    36              1.01
REMARK 500   C    VAL A    32     CB   SER A    36              2.10
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASN A   5       21.18   -155.58
REMARK 500    THR A  27       51.39   -114.49
REMARK 500    LEU A  30       11.32    -67.80
REMARK 500    ASN A 111       64.17     63.97
REMARK 500    SER A 161     -114.72     52.11
REMARK 500    LEU A 179       53.33   -113.41
REMARK 500    TYR A 190       68.81     36.44
REMARK 500    SER A 261       79.87   -105.92
REMARK 500    ASN B   5       23.18   -142.66
REMARK 500    LYS B  31       32.56    -86.97
REMARK 500    SER B  36       73.52    -67.84
REMARK 500    ASN B 111       63.96     63.12
REMARK 500    SER B 161     -114.68     51.76
REMARK 500    LEU B 179       51.84   -112.04
REMARK 500    TYR B 190       68.70     36.76
REMARK 500    SER B 261       79.72   -106.00
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 GLU A   35     SER A   36                 -144.82
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 402
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 B 401
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4WY8   RELATED DB: PDB
DBREF  4WY5 A    1   324  UNP    V5J5W4   V5J5W4_RHIMI     1    324
DBREF  4WY5 B    1   324  UNP    V5J5W4   V5J5W4_RHIMI     1    324
SEQADV 4WY5 LEU A  325  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 GLU A  326  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS A  327  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS A  328  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS A  329  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS A  330  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS A  331  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS A  332  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 LEU B  325  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 GLU B  326  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS B  327  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS B  328  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS B  329  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS B  330  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS B  331  UNP  V5J5W4              EXPRESSION TAG
SEQADV 4WY5 HIS B  332  UNP  V5J5W4              EXPRESSION TAG
SEQRES   1 A  332  MET THR VAL GLY ASN PRO PRO ILE HIS PRO VAL TYR ALA
SEQRES   2 A  332  ALA ALA PHE ALA ALA MET LYS GLU ARG PRO PRO ILE HIS
SEQRES   3 A  332  THR LEU ASP LEU LYS VAL VAL ARG GLU SER SER GLU ALA
SEQRES   4 A  332  ARG GLN LEU ALA ALA ASN ILE LYS LEU PRO GLU VAL ILE
SEQRES   5 A  332  GLU GLU ASP LYS VAL VAL GLU SER ASP GLY LYS THR LEU
SEQRES   6 A  332  LYS LEU THR ILE VAL ARG PRO PRO GLY THR GLU ASP GLN
SEQRES   7 A  332  ILE LEU PRO VAL LEU ILE PHE LEU HIS GLY GLY GLY PHE
SEQRES   8 A  332  VAL PHE GLY SER LYS TYR THR HIS ILE LYS PRO VAL ARG
SEQRES   9 A  332  ASP LEU THR VAL LYS ALA ASN VAL VAL THR VAL PHE VAL
SEQRES  10 A  332  ASP TYR SER LEU SER PRO GLU ALA LYS PHE PRO THR ALA
SEQRES  11 A  332  ILE GLU GLU ILE TYR ALA ALA ILE LEU TRP VAL ARG GLU
SEQRES  12 A  332  ASN ALA SER SER LEU ASN ILE ASN ALA GLU ALA LEU ALA
SEQRES  13 A  332  VAL ALA GLY ASP SER ALA GLY ALA THR LEU SER ALA ALA
SEQRES  14 A  332  VAL SER ILE TYR ALA LYS GLU LYS GLY LEU SER ALA ALA
SEQRES  15 A  332  ILE LYS THR GLN VAL LEU ILE TYR PRO ALA THR ALA VAL
SEQRES  16 A  332  SER HIS ALA LYS TYR GLU SER TYR LYS LEU PHE GLY ASN
SEQRES  17 A  332  GLY ASP TYR ILE LEU SER ALA GLU ASP LEU LYS PHE PHE
SEQRES  18 A  332  SER ASN ALA TYR LEU PRO ALA PRO ALA SER GLU LEU ASN
SEQRES  19 A  332  ASP LYS LEU ALA THR LEU GLU LEU ALA THR LYS ALA ASP
SEQRES  20 A  332  LEU GLU GLY LEU PRO PRO ALA LEU LEU PHE THR ALA GLU
SEQRES  21 A  332  SER ASP VAL LEU ARG ASP GLU GLY GLU LYS TYR ALA GLN
SEQRES  22 A  332  GLN LEU ALA GLU ALA GLY VAL ASP VAL ALA ALA VAL ARG
SEQRES  23 A  332  VAL LEU GLY ALA VAL HIS GLY PHE ILE THR VAL PRO VAL
SEQRES  24 A  332  GLU THR PRO GLN TYR ARG PHE THR ILE ASN THR ILE VAL
SEQRES  25 A  332  ALA HIS LEU ARG ASP ILE TYR ALA LYS TYR ASN ALA LEU
SEQRES  26 A  332  GLU HIS HIS HIS HIS HIS HIS
SEQRES   1 B  332  MET THR VAL GLY ASN PRO PRO ILE HIS PRO VAL TYR ALA
SEQRES   2 B  332  ALA ALA PHE ALA ALA MET LYS GLU ARG PRO PRO ILE HIS
SEQRES   3 B  332  THR LEU ASP LEU LYS VAL VAL ARG GLU SER SER GLU ALA
SEQRES   4 B  332  ARG GLN LEU ALA ALA ASN ILE LYS LEU PRO GLU VAL ILE
SEQRES   5 B  332  GLU GLU ASP LYS VAL VAL GLU SER ASP GLY LYS THR LEU
SEQRES   6 B  332  LYS LEU THR ILE VAL ARG PRO PRO GLY THR GLU ASP GLN
SEQRES   7 B  332  ILE LEU PRO VAL LEU ILE PHE LEU HIS GLY GLY GLY PHE
SEQRES   8 B  332  VAL PHE GLY SER LYS TYR THR HIS ILE LYS PRO VAL ARG
SEQRES   9 B  332  ASP LEU THR VAL LYS ALA ASN VAL VAL THR VAL PHE VAL
SEQRES  10 B  332  ASP TYR SER LEU SER PRO GLU ALA LYS PHE PRO THR ALA
SEQRES  11 B  332  ILE GLU GLU ILE TYR ALA ALA ILE LEU TRP VAL ARG GLU
SEQRES  12 B  332  ASN ALA SER SER LEU ASN ILE ASN ALA GLU ALA LEU ALA
SEQRES  13 B  332  VAL ALA GLY ASP SER ALA GLY ALA THR LEU SER ALA ALA
SEQRES  14 B  332  VAL SER ILE TYR ALA LYS GLU LYS GLY LEU SER ALA ALA
SEQRES  15 B  332  ILE LYS THR GLN VAL LEU ILE TYR PRO ALA THR ALA VAL
SEQRES  16 B  332  SER HIS ALA LYS TYR GLU SER TYR LYS LEU PHE GLY ASN
SEQRES  17 B  332  GLY ASP TYR ILE LEU SER ALA GLU ASP LEU LYS PHE PHE
SEQRES  18 B  332  SER ASN ALA TYR LEU PRO ALA PRO ALA SER GLU LEU ASN
SEQRES  19 B  332  ASP LYS LEU ALA THR LEU GLU LEU ALA THR LYS ALA ASP
SEQRES  20 B  332  LEU GLU GLY LEU PRO PRO ALA LEU LEU PHE THR ALA GLU
SEQRES  21 B  332  SER ASP VAL LEU ARG ASP GLU GLY GLU LYS TYR ALA GLN
SEQRES  22 B  332  GLN LEU ALA GLU ALA GLY VAL ASP VAL ALA ALA VAL ARG
SEQRES  23 B  332  VAL LEU GLY ALA VAL HIS GLY PHE ILE THR VAL PRO VAL
SEQRES  24 B  332  GLU THR PRO GLN TYR ARG PHE THR ILE ASN THR ILE VAL
SEQRES  25 B  332  ALA HIS LEU ARG ASP ILE TYR ALA LYS TYR ASN ALA LEU
SEQRES  26 B  332  GLU HIS HIS HIS HIS HIS HIS
HET    SO4  A 401       5
HET    SO4  A 402       5
HET    SO4  B 401       5
HETNAM     SO4 SULFATE ION
FORMUL   3  SO4    3(O4 S 2-)
FORMUL   6  HOH   *269(H2 O)
HELIX    1 AA1 HIS A    9  LYS A   20  1                                  12
HELIX    2 AA2 PRO A   24  LEU A   28  5                                   5
HELIX    3 AA3 ASP A   29  VAL A   33  5                                   5
HELIX    4 AA4 GLU A   38  ALA A   43  1                                   6
HELIX    5 AA5 HIS A   99  ASN A  111  1                                  13
HELIX    6 AA6 PRO A  128  ASN A  144  1                                  17
HELIX    7 AA7 ASN A  144  ASN A  149  1                                   6
HELIX    8 AA8 SER A  161  LYS A  177  1                                  17
HELIX    9 AA9 SER A  196  LYS A  199  5                                   4
HELIX   10 AB1 TYR A  200  GLY A  207  1                                   8
HELIX   11 AB2 SER A  214  LEU A  226  1                                  13
HELIX   12 AB3 PRO A  229  ASN A  234  1                                   6
HELIX   13 AB4 THR A  239  ALA A  243  5                                   5
HELIX   14 AB5 THR A  244  GLU A  249  1                                   6
HELIX   15 AB6 LEU A  264  ALA A  278  1                                  15
HELIX   16 AB7 GLY A  293  VAL A  297  5                                   5
HELIX   17 AB8 THR A  301  ALA A  320  1                                  20
HELIX   18 AB9 LYS A  321  ASN A  323  5                                   3
HELIX   19 AC1 HIS B    9  GLU B   21  1                                  13
HELIX   20 AC2 GLU B   38  ALA B   43  1                                   6
HELIX   21 AC3 HIS B   99  ASN B  111  1                                  13
HELIX   22 AC4 PRO B  128  ASN B  144  1                                  17
HELIX   23 AC5 ASN B  144  ASN B  149  1                                   6
HELIX   24 AC6 SER B  161  LYS B  177  1                                  17
HELIX   25 AC7 SER B  196  LYS B  199  5                                   4
HELIX   26 AC8 TYR B  200  GLY B  207  1                                   8
HELIX   27 AC9 SER B  214  LEU B  226  1                                  13
HELIX   28 AD1 PRO B  229  ASN B  234  1                                   6
HELIX   29 AD2 THR B  239  ALA B  243  5                                   5
HELIX   30 AD3 THR B  244  GLU B  249  1                                   6
HELIX   31 AD4 LEU B  264  ALA B  278  1                                  15
HELIX   32 AD5 GLY B  293  VAL B  297  5                                   5
HELIX   33 AD6 THR B  301  LYS B  321  1                                  21
SHEET    1 AA116 ILE A  52  SER A  60  0
SHEET    2 AA116 LYS A  63  ARG A  71 -1  O  ILE A  69   N  GLU A  54
SHEET    3 AA116 VAL A 113  VAL A 117 -1  O  PHE A 116   N  THR A  68
SHEET    4 AA116 LEU A  80  LEU A  86  1  N  LEU A  83   O  VAL A 113
SHEET    5 AA116 ILE A 150  ASP A 160  1  O  ALA A 156   N  ILE A  84
SHEET    6 AA116 THR A 185  ILE A 189  1  O  VAL A 187   N  VAL A 157
SHEET    7 AA116 ALA A 254  SER A 261  1  O  LEU A 255   N  LEU A 188
SHEET    8 AA116 VAL A 282  VAL A 291  1  O  VAL A 287   N  THR A 258
SHEET    9 AA116 VAL B 282  VAL B 291 -1  O  ALA B 284   N  LEU A 288
SHEET   10 AA116 ALA B 254  SER B 261  1  N  THR B 258   O  VAL B 287
SHEET   11 AA116 GLN B 186  ILE B 189  1  N  LEU B 188   O  LEU B 255
SHEET   12 AA116 ILE B 150  ASP B 160  1  N  VAL B 157   O  VAL B 187
SHEET   13 AA116 LEU B  80  LEU B  86  1  N  ILE B  84   O  ALA B 156
SHEET   14 AA116 VAL B 113  VAL B 117  1  O  VAL B 113   N  LEU B  83
SHEET   15 AA116 LYS B  63  ARG B  71 -1  N  THR B  68   O  PHE B 116
SHEET   16 AA116 ILE B  52  SER B  60 -1  N  GLU B  54   O  ILE B  69
CISPEP   1 SER A  122    PRO A  123          0         7.90
CISPEP   2 PHE A  127    PRO A  128          0         4.28
CISPEP   3 SER B  122    PRO B  123          0         8.22
CISPEP   4 PHE B  127    PRO B  128          0         4.78
SITE     1 AC1  7 ARG A 265  ARG A 286  HOH A 590  HOH A 634
SITE     2 AC1  7 HOH A 646  ARG B 265  ARG B 286
SITE     1 AC2  1 ARG A 104
SITE     1 AC3  1 ARG B 104
CRYST1   56.691   64.528   64.792 106.81 116.37 103.64 P 1           2
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.017639  0.004280  0.011977        0.00000
SCALE2      0.000000  0.015947  0.008092        0.00000
SCALE3      0.000000  0.000000  0.019317        0.00000
TER    2426      ASN A 323
TER    4861      ASN B 323
MASTER      315    0    3   33   16    0    4    6 5143    2   15   52
END