longtext: 4cib-pdb

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HEADER    HYDROLASE                               06-DEC-13   4CIB
TITLE     CRYSTAL STRUCTURE OF CATHEPSIN A, COMPLEXED WITH COMPOUND 2
CAVEAT     4CIB    NAG B 2 HAS WRONG CHIRALITY AT ATOM C1 NAG C 2 HAS WRONG
CAVEAT   2 4CIB    CHIRALITY AT ATOM C1 INCORRECT CHIRALITY AT C1 OF NAG A3011
CAVEAT   3 4CIB    INCORRECT CHIRALITY AT C1 OF NAG A3021
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LYSOSOMAL PROTECTIVE PROTEIN;
COMPND   3 CHAIN: A;
COMPND   4 SYNONYM: CATHEPSIN A, CARBOXYPEPTIDASE C, CARBOXYPEPTIDASE L,
COMPND   5 PROTECTIVE PROTEIN CATHEPSIN A, PPCA, PROTECTIVE PROTEIN FOR BETA-
COMPND   6 GALACTOSIDASE;
COMPND   7 EC: 3.4.16.5;
COMPND   8 ENGINEERED: YES;
COMPND   9 OTHER_DETAILS: ACTIVATED WITH TRYPSIN-SEPHAROSE
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 EXPRESSION_SYSTEM: SPODOPTERA FRUGIPERDA;
SOURCE   6 EXPRESSION_SYSTEM_COMMON: FALL ARMYWORM;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 7108;
SOURCE   8 EXPRESSION_SYSTEM_CELL_LINE: SF9;
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: BACULOVIRUS;
SOURCE  10 EXPRESSION_SYSTEM_VECTOR: BACULOVIRUS PVL1393
KEYWDS    HYDROLASE, DRUG DISCOVERY, SERINE CARBOXYPEPTIDASE, CARDIOVASCULAR
KEYWDS   2 DRUG, HEART FAILURE, ENDOTHELIN, TETRAHEDRAL INTERMEDIATE, COVALENT
KEYWDS   3 INHIBITOR
EXPDTA    X-RAY DIFFRACTION
AUTHOR    H.A.SCHREUDER,A.LIESUM,K.KROLL,B.BOEHNISCH,C.BUNING,S.RUF,C.BUNING,
AUTHOR   2 T.SADOWSKI
REVDAT   4   29-JUL-20 4CIB    1       CAVEAT COMPND REMARK HETNAM
REVDAT   4 2                   1       LINK   SITE   ATOM
REVDAT   3   13-MAR-19 4CIB    1       JRNL   REMARK LINK
REVDAT   2   02-APR-14 4CIB    1       JRNL
REVDAT   1   26-FEB-14 4CIB    0
JRNL        AUTH   H.A.SCHREUDER,A.LIESUM,K.KROLL,B.BOHNISCH,C.BUNING,S.RUF,
JRNL        AUTH 2 T.SADOWSKI
JRNL        TITL   CRYSTAL STRUCTURE OF CATHEPSIN A, A NOVEL TARGET FOR THE
JRNL        TITL 2 TREATMENT OF CARDIOVASCULAR DISEASES.
JRNL        REF    BIOCHEM. BIOPHYS. RES.        V. 445   451 2014
JRNL        REF  2 COMMUN.
JRNL        REFN                   ESSN 1090-2104
JRNL        PMID   24530914
JRNL        DOI    10.1016/J.BBRC.2014.02.014
REMARK   1
REMARK   1 REFERENCE 1
REMARK   1  AUTH   S.RUF,C.BUNING,H.SCHREUDER,G.HORSTICK,W.LINZ,T.OLPP,
REMARK   1  AUTH 2 J.PERNERSTORFER,K.HISS,K.KROLL,A.KANNT,M.KOHLMANN,D.LINZ,
REMARK   1  AUTH 3 T.HUBSCHLE,H.RUTTEN,K.WIRTH,T.SCHMIDT,T.SADOWSKI
REMARK   1  TITL   NOVEL BETA-AMINO ACID DERIVATIVES AS INHIBITORS OF CATHEPSIN
REMARK   1  TITL 2 A.
REMARK   1  REF    J.MED.CHEM.                   V.  55  7636 2012
REMARK   1  REFN                   ISSN 0022-2623
REMARK   1  PMID   22861813
REMARK   1  DOI    10.1021/JM300663N
REMARK   2
REMARK   2 RESOLUTION.    1.89 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.2.0019
REMARK   3   AUTHORS     : MURSHUDOV,SKUBAK,LEBEDEV,PANNU,STEINER,
REMARK   3               : NICHOLLS,WINN,LONG,VAGIN
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.89
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 44.02
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 100.0
REMARK   3   NUMBER OF REFLECTIONS             : 33271
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.145
REMARK   3   R VALUE            (WORKING SET) : 0.141
REMARK   3   FREE R VALUE                     : 0.211
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 1752
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.89
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.94
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 2414
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 100.0
REMARK   3   BIN R VALUE           (WORKING SET) : 0.1650
REMARK   3   BIN FREE R VALUE SET COUNT          : 127
REMARK   3   BIN FREE R VALUE                    : 0.2360
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 3292
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 75
REMARK   3   SOLVENT ATOMS            : 908
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 20.20
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 18.55
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -0.05000
REMARK   3    B22 (A**2) : 0.05000
REMARK   3    B33 (A**2) : -0.03000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : -0.09000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.140
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.145
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.087
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 2.815
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.964
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.919
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  3460 ; 0.013 ; 0.022
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  4710 ; 1.409 ; 1.975
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   410 ; 5.755 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   169 ;37.911 ;24.911
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   533 ;12.274 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    12 ;14.224 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   501 ; 0.097 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  2679 ; 0.006 ; 0.020
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  1809 ; 0.207 ; 0.200
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  2343 ; 0.312 ; 0.200
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):   687 ; 0.190 ; 0.200
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):    78 ; 0.181 ; 0.200
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):    87 ; 0.202 ; 0.200
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):     1 ; 0.011 ; 0.200
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  2108 ; 0.803 ; 1.500
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  3305 ; 1.300 ; 2.000
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  1558 ; 2.052 ; 3.000
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  1405 ; 3.105 ; 4.500
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.40
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS.
REMARK   4
REMARK   4 4CIB COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 06-DEC-13.
REMARK 100 THE DEPOSITION ID IS D_1290059179.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 29-AUG-07
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 4.6
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID14-1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.93400
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC CCD
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 35026
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.890
REMARK 200  RESOLUTION RANGE LOW       (A) : 66.740
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.9
REMARK 200  DATA REDUNDANCY                : 3.800
REMARK 200  R MERGE                    (I) : 0.08000
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 14.1200
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.89
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.94
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.80
REMARK 200  R MERGE FOR SHELL          (I) : 0.35000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 3.760
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: PDB ENTRY 4AZ0
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 49.20
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.42
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: CATHEPSIN A WAS CRYSTALLIZED USING THE
REMARK 280  HANGING DROP METHOD: 1 UL OF PROTEIN SOLUTION, CONTAINING 6.5 MG/
REMARK 280  ML CATHEPSIN A, 25 MM TRIS-HCL (PH 8.0) AND 300 MM NACL, WAS
REMARK 280  MIXED WITH 1 UL RESERVOIR SOLUTION, CONTAINING 100 MM NAACETATE
REMARK 280  (PH 4.5), 18-20% PEG400 AND 100 MM CDCL2, AND SET TO EQUILIBRATE
REMARK 280  AT 4DEG.C. ROD-SHAPED CRYSTALS APPEARED IN ABOUT ONE WEEK., PH
REMARK 280  4.6, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 277K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 1 2 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y,-Z
REMARK 290       3555   X+1/2,Y+1/2,Z
REMARK 290       4555   -X+1/2,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   3  1.000000  0.000000  0.000000       44.82450
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       51.26350
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000       44.82450
REMARK 290   SMTRY2   4  0.000000  1.000000  0.000000       51.26350
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 6650 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 32770 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -53.7 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2 -1.000000  0.000000  0.000000      -89.64900
REMARK 350   BIOMT2   2  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH A2181  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A2308  LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     SER A    -1
REMARK 465     HIS A   260
REMARK 465     PHE A   261
REMARK 465     ARG A   262
REMARK 465     TYR A   263
REMARK 465     GLU A   264
REMARK 465     LYS A   265
REMARK 465     ASP A   266
REMARK 465     THR A   267
REMARK 465     VAL A   268
REMARK 465     VAL A   269
REMARK 465     VAL A   270
REMARK 465     GLN A   271
REMARK 465     ASP A   272
REMARK 465     LEU A   273
REMARK 465     GLY A   274
REMARK 465     ASN A   275
REMARK 465     ILE A   276
REMARK 465     PHE A   277
REMARK 465     THR A   278
REMARK 465     ARG A   279
REMARK 465     LEU A   280
REMARK 465     PRO A   281
REMARK 465     LEU A   282
REMARK 465     LYS A   283
REMARK 465     ARG A   284
REMARK 465     MET A   285
REMARK 465     TRP A   286
REMARK 465     HIS A   287
REMARK 465     GLN A   288
REMARK 465     ALA A   289
REMARK 465     LEU A   290
REMARK 465     LEU A   291
REMARK 465     ARG A   292
REMARK 465     SER A   293
REMARK 465     GLY A   294
REMARK 465     ASP A   295
REMARK 465     LYS A   296
REMARK 465     VAL A   297
REMARK 465     ARG A   298
REMARK 465     MET A   299
REMARK 465     ASP A   404
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     ARG A   0    CG   CD   NE   CZ   NH1  NH2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    ARG A     0     OE1  GLU A   138              1.90
REMARK 500   ND2  ASN A   202     O    HOH A  2539              2.01
REMARK 500   OD2  ASP A   110     O    HOH A  2160              2.02
REMARK 500   O    HOH A  2691     O    HOH A  2698              2.03
REMARK 500   O    HOH A  2026     O    HOH A  2690              2.05
REMARK 500   N    SER A   405     O    HOH A  2820              2.05
REMARK 500   O    HOH A  2277     O    HOH A  2278              2.06
REMARK 500   O    HOH A  2260     O    HOH A  2261              2.08
REMARK 500   O    HOH A  2506     O    HOH A  2507              2.09
REMARK 500   O    HOH A  2255     O    HOH A  2401              2.10
REMARK 500   O    HOH A  2401     O    HOH A  2733              2.12
REMARK 500   O    HOH A  2618     O    HOH A  2627              2.13
REMARK 500   O    HOH A  2026     O    HOH A  2054              2.13
REMARK 500   O    HOH A  2160     O    HOH A  2337              2.15
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    LEU A 174   CA  -  CB  -  CG  ANGL. DEV. =  25.5 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    GLU A  69      -77.77   -104.01
REMARK 500    PRO A 101     -168.63    -79.11
REMARK 500    SER A 150     -108.03     57.08
REMARK 500    ASN A 178       52.38     39.85
REMARK 500    GLN A 215     -125.81     46.92
REMARK 500    TYR A 221      -62.15   -103.08
REMARK 500    ASN A 248      104.15   -160.59
REMARK 500    CYS A 303       -0.83     69.70
REMARK 500    TYR A 402     -109.25    -90.62
REMARK 500    MET A 430       79.04   -108.56
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A2012        DISTANCE =  7.38 ANGSTROMS
REMARK 525    HOH A2014        DISTANCE =  6.47 ANGSTROMS
REMARK 525    HOH A2015        DISTANCE =  6.29 ANGSTROMS
REMARK 525    HOH A2016        DISTANCE =  6.92 ANGSTROMS
REMARK 525    HOH A2033        DISTANCE =  5.82 ANGSTROMS
REMARK 525    HOH A2080        DISTANCE =  6.07 ANGSTROMS
REMARK 525    HOH A2131        DISTANCE =  5.94 ANGSTROMS
REMARK 525    HOH A2136        DISTANCE =  6.34 ANGSTROMS
REMARK 525    HOH A2166        DISTANCE =  5.99 ANGSTROMS
REMARK 525    HOH A2210        DISTANCE =  6.76 ANGSTROMS
REMARK 525    HOH A2308        DISTANCE =  7.22 ANGSTROMS
REMARK 525    HOH A2310        DISTANCE =  6.06 ANGSTROMS
REMARK 525    HOH A2313        DISTANCE =  6.68 ANGSTROMS
REMARK 525    HOH A2314        DISTANCE =  6.21 ANGSTROMS
REMARK 525    HOH A2358        DISTANCE =  7.40 ANGSTROMS
REMARK 525    HOH A2383        DISTANCE =  6.26 ANGSTROMS
REMARK 525    HOH A2387        DISTANCE =  6.21 ANGSTROMS
REMARK 525    HOH A2393        DISTANCE =  6.08 ANGSTROMS
REMARK 525    HOH A2407        DISTANCE =  6.17 ANGSTROMS
REMARK 525    HOH A2408        DISTANCE =  5.98 ANGSTROMS
REMARK 525    HOH A2409        DISTANCE =  7.06 ANGSTROMS
REMARK 525    HOH A2432        DISTANCE =  5.92 ANGSTROMS
REMARK 525    HOH A2433        DISTANCE =  6.80 ANGSTROMS
REMARK 525    HOH A2455        DISTANCE =  6.68 ANGSTROMS
REMARK 525    HOH A2457        DISTANCE =  6.03 ANGSTROMS
REMARK 525    HOH A2796        DISTANCE =  7.91 ANGSTROMS
REMARK 525    HOH A2900        DISTANCE =  5.97 ANGSTROMS
REMARK 525    HOH A2901        DISTANCE =  7.33 ANGSTROMS
REMARK 525    HOH A2902        DISTANCE =  5.88 ANGSTROMS
REMARK 525    HOH A2903        DISTANCE =  7.51 ANGSTROMS
REMARK 525    HOH A2905        DISTANCE =  7.84 ANGSTROMS
REMARK 525    HOH A2906        DISTANCE =  6.46 ANGSTROMS
REMARK 525    HOH A2907        DISTANCE =  7.93 ANGSTROMS
REMARK 525    HOH A2908        DISTANCE =  8.44 ANGSTROMS
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              CD A1458  CD
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASP A   3   OD2
REMARK 620 2 ASP A   3   OD1  54.6
REMARK 620 3 HIS A 211   ND1 106.8  90.9
REMARK 620 4 ASP A 225   OD1  73.5 127.0  94.7
REMARK 620 5 ASP A 225   OD2  95.1 134.4 133.3  52.4
REMARK 620 6 HOH A2018   O    83.9  74.4 152.4 112.9  68.6
REMARK 620 7 HOH A2875   O   141.4  94.7  95.3 136.8  92.0  63.6
REMARK 620 N                    1     2     3     4     5     6
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              CD A1456  CD
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 GLY A 179   O
REMARK 620 2 ASP A 380   OD1  67.8
REMARK 620 3 ASP A 380   OD2  68.2  42.6
REMARK 620 4 HOH A2502   O    99.1  98.5 141.0
REMARK 620 5 HOH A2520   O   162.1 103.6  94.7  97.7
REMARK 620 N                    1     2     3     4
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              CD A1457  CD
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 GLU A 184   OE2
REMARK 620 2 GLU A 184   OE1  43.7
REMARK 620 3 HOH A2512   O    83.3  96.3
REMARK 620 4 HOH A2513   O    89.4 129.4  95.0
REMARK 620 5 HOH A2583   O    97.2  84.5 179.3  84.5
REMARK 620 6 HOH A2874   O   174.9 141.2  94.1  86.5  85.3
REMARK 620 N                    1     2     3     4     5
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              CD A1459  CD
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASN A 186   OD1
REMARK 620 2 CYS A 375   SG  135.4
REMARK 620 N                    1
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              CD A1455  CD
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASP A 222   OD1
REMARK 620 2 ASP A 222   OD2  56.2
REMARK 620 3 GLU A 326   OE1 162.8 134.1
REMARK 620 4 GLU A 326   OE2 136.0  85.3  49.0
REMARK 620 5 HOH A2513   O    88.2 141.6  84.2 133.1
REMARK 620 6 HOH A2575   O    86.0 104.5  78.0  84.0  84.7
REMARK 620 7 HOH A2583   O   102.1  84.0  93.4  93.0  91.0 170.7
REMARK 620 N                    1     2     3     4     5     6
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4CI9   RELATED DB: PDB
REMARK 900 CRYSTAL STRUCTURE OF CATHEPSIN A, APO-STRUCTURE
REMARK 900 RELATED ID: 4CIA   RELATED DB: PDB
REMARK 900 CRYSTAL STRUCTURE OF CATHEPSIN A, COMPLEXED WITH COMPOUND 1
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 2 EXTRA RESIDUES FROM PREPROMELLITIN SIGNAL SEQUENCE AT N-
REMARK 999 TERMINUS AND 1 EXTRA RESIDUE, LEFT OVER FROM MYC TAG AT C-
REMARK 999 TERMINUS
DBREF  4CIB A    1   452  UNP    P10619   PPGB_HUMAN      29    480
SEQADV 4CIB SER A   -1  UNP  P10619              EXPRESSION TAG
SEQADV 4CIB ARG A    0  UNP  P10619              EXPRESSION TAG
SEQADV 4CIB GLU A  453  UNP  P10619              EXPRESSION TAG
SEQRES   1 A  455  SER ARG ALA PRO ASP GLN ASP GLU ILE GLN ARG LEU PRO
SEQRES   2 A  455  GLY LEU ALA LYS GLN PRO SER PHE ARG GLN TYR SER GLY
SEQRES   3 A  455  TYR LEU LYS GLY SER GLY SER LYS HIS LEU HIS TYR TRP
SEQRES   4 A  455  PHE VAL GLU SER GLN LYS ASP PRO GLU ASN SER PRO VAL
SEQRES   5 A  455  VAL LEU TRP LEU ASN GLY GLY PRO GLY CYS SER SER LEU
SEQRES   6 A  455  ASP GLY LEU LEU THR GLU HIS GLY PRO PHE LEU VAL GLN
SEQRES   7 A  455  PRO ASP GLY VAL THR LEU GLU TYR ASN PRO TYR SER TRP
SEQRES   8 A  455  ASN LEU ILE ALA ASN VAL LEU TYR LEU GLU SER PRO ALA
SEQRES   9 A  455  GLY VAL GLY PHE SER TYR SER ASP ASP LYS PHE TYR ALA
SEQRES  10 A  455  THR ASN ASP THR GLU VAL ALA GLN SER ASN PHE GLU ALA
SEQRES  11 A  455  LEU GLN ASP PHE PHE ARG LEU PHE PRO GLU TYR LYS ASN
SEQRES  12 A  455  ASN LYS LEU PHE LEU THR GLY GLU SER TYR ALA GLY ILE
SEQRES  13 A  455  TYR ILE PRO THR LEU ALA VAL LEU VAL MET GLN ASP PRO
SEQRES  14 A  455  SER MET ASN LEU GLN GLY LEU ALA VAL GLY ASN GLY LEU
SEQRES  15 A  455  SER SER TYR GLU GLN ASN ASP ASN SER LEU VAL TYR PHE
SEQRES  16 A  455  ALA TYR TYR HIS GLY LEU LEU GLY ASN ARG LEU TRP SER
SEQRES  17 A  455  SER LEU GLN THR HIS CYS CYS SER GLN ASN LYS CYS ASN
SEQRES  18 A  455  PHE TYR ASP ASN LYS ASP LEU GLU CYS VAL THR ASN LEU
SEQRES  19 A  455  GLN GLU VAL ALA ARG ILE VAL GLY ASN SER GLY LEU ASN
SEQRES  20 A  455  ILE TYR ASN LEU TYR ALA PRO CYS ALA GLY GLY VAL PRO
SEQRES  21 A  455  SER HIS PHE ARG TYR GLU LYS ASP THR VAL VAL VAL GLN
SEQRES  22 A  455  ASP LEU GLY ASN ILE PHE THR ARG LEU PRO LEU LYS ARG
SEQRES  23 A  455  MET TRP HIS GLN ALA LEU LEU ARG SER GLY ASP LYS VAL
SEQRES  24 A  455  ARG MET ASP PRO PRO CYS THR ASN THR THR ALA ALA SER
SEQRES  25 A  455  THR TYR LEU ASN ASN PRO TYR VAL ARG LYS ALA LEU ASN
SEQRES  26 A  455  ILE PRO GLU GLN LEU PRO GLN TRP ASP MET CYS ASN PHE
SEQRES  27 A  455  LEU VAL ASN LEU GLN TYR ARG ARG LEU TYR ARG SER MET
SEQRES  28 A  455  ASN SER GLN TYR LEU LYS LEU LEU SER SER GLN LYS TYR
SEQRES  29 A  455  GLN ILE LEU LEU TYR ASN GLY ASP VAL ASP MET ALA CYS
SEQRES  30 A  455  ASN PHE MET GLY ASP GLU TRP PHE VAL ASP SER LEU ASN
SEQRES  31 A  455  GLN LYS MET GLU VAL GLN ARG ARG PRO TRP LEU VAL LYS
SEQRES  32 A  455  TYR GLY ASP SER GLY GLU GLN ILE ALA GLY PHE VAL LYS
SEQRES  33 A  455  GLU PHE SER HIS ILE ALA PHE LEU THR ILE LYS GLY ALA
SEQRES  34 A  455  GLY HIS MET VAL PRO THR ASP LYS PRO LEU ALA ALA PHE
SEQRES  35 A  455  THR MET PHE SER ARG PHE LEU ASN LYS GLN PRO TYR GLU
MODRES 4CIB ASN A  117  ASN  GLYCOSYLATION SITE
MODRES 4CIB ASN A  305  ASN  GLYCOSYLATION SITE
HET    NAG  B   1      14
HET    NAG  B   2      14
HET    NAG  C   1      14
HET    NAG  C   2      14
HET    7UZ  A1454      14
HET     CD  A1455       1
HET     CD  A1456       1
HET     CD  A1457       1
HET     CD  A1458       1
HET     CD  A1459       1
HETNAM     NAG 2-ACETAMIDO-2-DEOXY-BETA-D-GLUCOPYRANOSE
HETNAM     7UZ 2-(CYCLOHEXYLMETHYL)PROPANEDIOIC ACID
HETNAM      CD CADMIUM ION
FORMUL   2  NAG    4(C8 H15 N O6)
FORMUL   4  7UZ    C10 H16 O4
FORMUL   5   CD    5(CD 2+)
FORMUL  10  HOH   *908(H2 O)
HELIX    1   1 PRO A    2  ASP A    5  5                                   4
HELIX    2   2 SER A   62  THR A   68  1                                   7
HELIX    3   3 SER A   88  ILE A   92  5                                   5
HELIX    4   4 ASN A  117  PHE A  136  1                                  20
HELIX    5   5 PRO A  137  LYS A  140  5                                   4
HELIX    6   6 TYR A  151  MET A  164  1                                  14
HELIX    7   7 SER A  182  HIS A  197  1                                  16
HELIX    8   8 GLY A  201  CYS A  212  1                                  12
HELIX    9   9 ASP A  225  ASN A  241  1                                  17
HELIX   10  10 THR A  306  ASN A  314  1                                   9
HELIX   11  11 ASN A  315  LEU A  322  1                                   8
HELIX   12  12 ASN A  335  TYR A  342  1                                   8
HELIX   13  13 MET A  349  SER A  359  1                                  11
HELIX   14  14 ASN A  376  LEU A  387  1                                  12
HELIX   15  15 MET A  430  LYS A  435  1                                   6
HELIX   16  16 LYS A  435  ASN A  448  1                                  14
SHEET    1  AA 2 GLN A  21  LYS A  27  0
SHEET    2  AA 2 LYS A  32  VAL A  39 -1  O  LEU A  34   N  LEU A  26
SHEET    1  AB 2 TYR A 108  SER A 109  0
SHEET    2  AB 2 LYS A  32  VAL A  39  1  O  HIS A  33   N  TYR A 108
SHEET    1  AC10 ARG A 396  LYS A 401  0
SHEET    2  AC10 GLU A 407  PHE A 416 -1  O  GLN A 408   N  VAL A 400
SHEET    3  AC10 ILE A 419  ILE A 424 -1  O  ILE A 419   N  PHE A 416
SHEET    4  AC10 GLN A 363  GLY A 369  1  O  ILE A 364   N  ALA A 420
SHEET    5  AC10 LEU A 171  GLY A 177  1  O  GLN A 172   N  GLN A 363
SHEET    6  AC10 LEU A 144  GLU A 149  1  O  LEU A 144   N  GLN A 172
SHEET    7  AC10 VAL A  50  LEU A  54  1  O  VAL A  50   N  PHE A 145
SHEET    8  AC10 ASN A  94  LEU A  98  1  O  ASN A  94   N  VAL A  51
SHEET    9  AC10 LYS A  32  VAL A  39 -1  O  TRP A  37   N  TYR A  97
SHEET   10  AC10 TYR A 108  SER A 109  1  O  TYR A 108   N  HIS A  33
SHEET    1  AD10 ARG A 396  LYS A 401  0
SHEET    2  AD10 GLU A 407  PHE A 416 -1  O  GLN A 408   N  VAL A 400
SHEET    3  AD10 ILE A 419  ILE A 424 -1  O  ILE A 419   N  PHE A 416
SHEET    4  AD10 GLN A 363  GLY A 369  1  O  ILE A 364   N  ALA A 420
SHEET    5  AD10 LEU A 171  GLY A 177  1  O  GLN A 172   N  GLN A 363
SHEET    6  AD10 LEU A 144  GLU A 149  1  O  LEU A 144   N  GLN A 172
SHEET    7  AD10 VAL A  50  LEU A  54  1  O  VAL A  50   N  PHE A 145
SHEET    8  AD10 ASN A  94  LEU A  98  1  O  ASN A  94   N  VAL A  51
SHEET    9  AD10 LYS A  32  VAL A  39 -1  O  TRP A  37   N  TYR A  97
SHEET   10  AD10 GLN A  21  LYS A  27 -1  O  TYR A  22   N  PHE A  38
SHEET    1  AE 2 PHE A  73  VAL A  75  0
SHEET    2  AE 2 LEU A  82  TYR A  84 -1  O  GLU A  83   N  LEU A  74
SHEET    1  AF 2 CYS A 213  SER A 214  0
SHEET    2  AF 2 LYS A 217  CYS A 218 -1  O  LYS A 217   N  SER A 214
SSBOND   1 CYS A   60    CYS A  334                          1555   1555  2.05
SSBOND   2 CYS A  212    CYS A  228                          1555   1555  2.03
SSBOND   3 CYS A  213    CYS A  218                          1555   1555  2.05
SSBOND   4 CYS A  253    CYS A  303                          1555   1555  2.04
LINK         ND2 ASN A 117                 C1  NAG B   1     1555   1555  1.45
LINK         ND2 ASN A 305                 C1  NAG C   1     1555   1555  1.44
LINK         O4  NAG B   1                 C1  NAG B   2     1555   1555  1.45
LINK         O4  NAG C   1                 C1  NAG C   2     1555   1555  1.45
LINK         OD2 ASP A   3                CD    CD A1458     1555   1555  2.31
LINK         OD1 ASP A   3                CD    CD A1458     1555   1555  2.44
LINK         O   GLY A 179                CD    CD A1456     1555   1555  2.24
LINK         OE2 GLU A 184                CD    CD A1457     1555   1555  2.30
LINK         OE1 GLU A 184                CD    CD A1457     1555   1555  3.20
LINK         OD1 ASN A 186                CD    CD A1459     1555   1555  3.21
LINK         ND1 HIS A 211                CD    CD A1458     3545   1555  2.35
LINK         OD1 ASP A 222                CD    CD A1455     1555   1555  2.35
LINK         OD2 ASP A 222                CD    CD A1455     1555   1555  2.32
LINK         OD1 ASP A 225                CD    CD A1458     3545   1555  2.66
LINK         OD2 ASP A 225                CD    CD A1458     3545   1555  2.19
LINK         OE1 GLU A 326                CD    CD A1455     4456   1555  2.83
LINK         OE2 GLU A 326                CD    CD A1455     4456   1555  2.35
LINK         SG  CYS A 375                CD    CD A1459     1555   1555  2.46
LINK         OD1 ASP A 380                CD    CD A1456     1555   1555  3.24
LINK         OD2 ASP A 380                CD    CD A1456     1555   1555  2.70
LINK        CD    CD A1455                 O   HOH A2513     1555   1555  2.50
LINK        CD    CD A1455                 O   HOH A2575     1555   1555  2.42
LINK        CD    CD A1455                 O   HOH A2583     1555   1555  2.57
LINK        CD    CD A1456                 O   HOH A2502     1555   1555  2.32
LINK        CD    CD A1456                 O   HOH A2520     1555   1555  2.48
LINK        CD    CD A1457                 O   HOH A2512     1555   1555  2.42
LINK        CD    CD A1457                 O   HOH A2513     1555   1555  2.73
LINK        CD    CD A1457                 O   HOH A2583     1555   1555  2.64
LINK        CD    CD A1457                 O   HOH A2874     1555   1555  2.46
LINK        CD    CD A1458                 O   HOH A2018     1555   1555  3.07
LINK        CD    CD A1458                 O   HOH A2875     1555   1555  2.57
CISPEP   1 GLY A   57    PRO A   58          0        -1.10
CISPEP   2 SER A  100    PRO A  101          0         0.44
CRYST1   89.649  102.527   49.395  90.00 100.94  90.00 C 1 2 1       4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.011155  0.000000  0.002156        0.00000
SCALE2      0.000000  0.009754  0.000000        0.00000
SCALE3      0.000000  0.000000  0.020620        0.00000
TER    3293      GLU A 453
MASTER      505    0   10   16   28    0    0    6 4275    1  108   35
END