longtext: 4yk7-pdb

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HEADER    LYASE                                   04-MAR-15   4YK7
TITLE     CRYSTAL STRUCTURE OF S-HYDROXYNITRILE LYASE FROM MANIHOT ESCULENTA
TITLE    2 (HIS103LEU)
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: (S)-HYDROXYNITRILE LYASE;
COMPND   3 CHAIN: A, B, C, D;
COMPND   4 SYNONYM: (S)-ACETONE-CYANOHYDRIN LYASE,OXYNITRILASE;
COMPND   5 EC: 4.1.2.47;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: MANIHOT ESCULENTA;
SOURCE   3 ORGANISM_COMMON: CASSAVA;
SOURCE   4 ORGANISM_TAXID: 3983;
SOURCE   5 GENE: HNL;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 562
KEYWDS    S-HYDROXYNITRILE LYASE, HYDROLASE FOLD, HIS103LEU VARIANT, LYASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.DADASHIPOUR,S.NAKANO,Y.ASANO
REVDAT   1   13-APR-16 4YK7    0
JRNL        AUTH   M.DADASHIPOUR,S.NAKANO,Y.ASANO
JRNL        TITL   CRYSTAL STRUCTURE OF S-HYDROXYNITRILE LYASE FROM MANIHOT
JRNL        TITL 2 ESCULENTA (HIS103LEU)
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    2.60 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.7.0029
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.60
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 45.60
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.3
REMARK   3   NUMBER OF REFLECTIONS             : 30495
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.241
REMARK   3   R VALUE            (WORKING SET) : 0.240
REMARK   3   FREE R VALUE                     : 0.262
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.100
REMARK   3   FREE R VALUE TEST SET COUNT      : 1631
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.60
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.67
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 2014
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 90.49
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2710
REMARK   3   BIN FREE R VALUE SET COUNT          : 99
REMARK   3   BIN FREE R VALUE                    : 0.3090
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 8288
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 0
REMARK   3   SOLVENT ATOMS            : 60
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 17.00
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -2.36000
REMARK   3    B22 (A**2) : 5.47000
REMARK   3    B33 (A**2) : -3.11000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): NULL
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.364
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.247
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 11.329
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.869
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.840
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  8488 ; 0.007 ; 0.019
REMARK   3   BOND LENGTHS OTHERS               (A):  8140 ; 0.003 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES): 11512 ; 1.010 ; 1.952
REMARK   3   BOND ANGLES OTHERS          (DEGREES): 18760 ; 0.781 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):  1028 ; 7.306 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   384 ;38.911 ;24.583
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):  1488 ;18.393 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    32 ;17.980 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):  1288 ; 0.064 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  9472 ; 0.012 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  1936 ; 0.009 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NCS TYPE: LOCAL
REMARK   3   NUMBER OF DIFFERENT NCS PAIRS  : 6
REMARK   3  GROUP  CHAIN1    RANGE     CHAIN2     RANGE    COUNT RMS  WEIGHT
REMARK   3    1     A     1    258       B     1    258   16628  0.09  0.05
REMARK   3    2     A     1    258       C     1    258   17061  0.07  0.05
REMARK   3    3     A     1    258       D     1    258   16650  0.09  0.05
REMARK   3    4     B     1    258       C     1    258   16578  0.10  0.05
REMARK   3    5     B     1    258       D     1    258   17119  0.07  0.05
REMARK   3    6     C     1    258       D     1    258   16525  0.10  0.05
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS
REMARK   4
REMARK   4 4YK7 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 12-MAR-15.
REMARK 100 THE DEPOSITION ID IS D_1000207594.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 22-NOV-13
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : NULL
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : PHOTON FACTORY
REMARK 200  BEAMLINE                       : BL-17A
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315R
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 32186
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.600
REMARK 200  RESOLUTION RANGE LOW       (A) : 45.600
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 100.0
REMARK 200  DATA REDUNDANCY                : 14.10
REMARK 200  R MERGE                    (I) : 0.14400
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 34.6000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.60
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.67
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 14.10
REMARK 200  R MERGE FOR SHELL          (I) : 0.38200
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 12.10
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 43.86
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.19
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 2.1M DL-MALEIC ACID, VAPOR DIFFUSION,
REMARK 280  SITTING DROP, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       43.23450
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       67.60650
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       44.04900
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       67.60650
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       43.23450
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       44.04900
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2, 3
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: TETRAMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 1780 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 19230 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -14.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 3
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 1710 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 19250 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -15.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    CYS A  13      -30.73     85.47
REMARK 500    HIS A  14     -148.55    -91.17
REMARK 500    SER A  80     -114.79     56.19
REMARK 500    TYR A  93       45.29   -144.61
REMARK 500    ARG A 129     -125.66     74.48
REMARK 500    ILE A 210      -42.48   -132.47
REMARK 500    CYS B  13      -31.55     82.71
REMARK 500    HIS B  14     -152.35    -88.04
REMARK 500    SER B  80     -114.16     55.07
REMARK 500    TYR B  93       47.85   -146.49
REMARK 500    ARG B 129     -124.85     75.69
REMARK 500    ILE B 210      -44.33   -131.73
REMARK 500    LYS B 237       70.65   -101.07
REMARK 500    CYS C  13      -31.79     83.93
REMARK 500    HIS C  14     -148.20    -90.61
REMARK 500    SER C  80     -113.66     54.61
REMARK 500    TYR C  93       43.95   -144.19
REMARK 500    ARG C 129     -123.73     69.96
REMARK 500    ILE C 210      -42.64   -132.81
REMARK 500    LYS C 237       70.10   -100.94
REMARK 500    CYS D  13      -31.31     82.27
REMARK 500    HIS D  14     -152.24    -89.54
REMARK 500    SER D  80     -115.23     53.56
REMARK 500    TYR D  93       44.81   -146.42
REMARK 500    ARG D 129     -122.47     72.43
REMARK 500    ILE D 210      -44.84   -132.34
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 TYR B  257     ALA B  258                  132.60
REMARK 500 TYR D  257     ALA D  258                  132.74
REMARK 500
REMARK 500 REMARK: NULL
DBREF  4YK7 A    1   258  UNP    P52705   HNL_MANES        1    258
DBREF  4YK7 B    1   258  UNP    P52705   HNL_MANES        1    258
DBREF  4YK7 C    1   258  UNP    P52705   HNL_MANES        1    258
DBREF  4YK7 D    1   258  UNP    P52705   HNL_MANES        1    258
SEQADV 4YK7 LEU A  103  UNP  P52705    HIS   103 ENGINEERED MUTATION
SEQADV 4YK7 LEU B  103  UNP  P52705    HIS   103 ENGINEERED MUTATION
SEQADV 4YK7 LEU C  103  UNP  P52705    HIS   103 ENGINEERED MUTATION
SEQADV 4YK7 LEU D  103  UNP  P52705    HIS   103 ENGINEERED MUTATION
SEQRES   1 A  258  MET VAL THR ALA HIS PHE VAL LEU ILE HIS THR ILE CYS
SEQRES   2 A  258  HIS GLY ALA TRP ILE TRP HIS LYS LEU LYS PRO ALA LEU
SEQRES   3 A  258  GLU ARG ALA GLY HIS LYS VAL THR ALA LEU ASP MET ALA
SEQRES   4 A  258  ALA SER GLY ILE ASP PRO ARG GLN ILE GLU GLN ILE ASN
SEQRES   5 A  258  SER PHE ASP GLU TYR SER GLU PRO LEU LEU THR PHE LEU
SEQRES   6 A  258  GLU LYS LEU PRO GLN GLY GLU LYS VAL ILE ILE VAL GLY
SEQRES   7 A  258  GLU SER CYS ALA GLY LEU ASN ILE ALA ILE ALA ALA ASP
SEQRES   8 A  258  ARG TYR VAL ASP LYS ILE ALA ALA GLY VAL PHE LEU ASN
SEQRES   9 A  258  SER LEU LEU PRO ASP THR VAL HIS SER PRO SER TYR THR
SEQRES  10 A  258  VAL GLU LYS LEU LEU GLU SER PHE PRO ASP TRP ARG ASP
SEQRES  11 A  258  THR GLU TYR PHE THR PHE THR ASN ILE THR GLY GLU THR
SEQRES  12 A  258  ILE THR THR MET LYS LEU GLY PHE VAL LEU LEU ARG GLU
SEQRES  13 A  258  ASN LEU PHE THR LYS CYS THR ASP GLY GLU TYR GLU LEU
SEQRES  14 A  258  ALA LYS MET VAL MET ARG LYS GLY SER LEU PHE GLN ASN
SEQRES  15 A  258  VAL LEU ALA GLN ARG PRO LYS PHE THR GLU LYS GLY TYR
SEQRES  16 A  258  GLY SER ILE LYS LYS VAL TYR ILE TRP THR ASP GLN ASP
SEQRES  17 A  258  LYS ILE PHE LEU PRO ASP PHE GLN ARG TRP GLN ILE ALA
SEQRES  18 A  258  ASN TYR LYS PRO ASP LYS VAL TYR GLN VAL GLN GLY GLY
SEQRES  19 A  258  ASP HIS LYS LEU GLN LEU THR LYS THR GLU GLU VAL ALA
SEQRES  20 A  258  HIS ILE LEU GLN GLU VAL ALA ASP ALA TYR ALA
SEQRES   1 B  258  MET VAL THR ALA HIS PHE VAL LEU ILE HIS THR ILE CYS
SEQRES   2 B  258  HIS GLY ALA TRP ILE TRP HIS LYS LEU LYS PRO ALA LEU
SEQRES   3 B  258  GLU ARG ALA GLY HIS LYS VAL THR ALA LEU ASP MET ALA
SEQRES   4 B  258  ALA SER GLY ILE ASP PRO ARG GLN ILE GLU GLN ILE ASN
SEQRES   5 B  258  SER PHE ASP GLU TYR SER GLU PRO LEU LEU THR PHE LEU
SEQRES   6 B  258  GLU LYS LEU PRO GLN GLY GLU LYS VAL ILE ILE VAL GLY
SEQRES   7 B  258  GLU SER CYS ALA GLY LEU ASN ILE ALA ILE ALA ALA ASP
SEQRES   8 B  258  ARG TYR VAL ASP LYS ILE ALA ALA GLY VAL PHE LEU ASN
SEQRES   9 B  258  SER LEU LEU PRO ASP THR VAL HIS SER PRO SER TYR THR
SEQRES  10 B  258  VAL GLU LYS LEU LEU GLU SER PHE PRO ASP TRP ARG ASP
SEQRES  11 B  258  THR GLU TYR PHE THR PHE THR ASN ILE THR GLY GLU THR
SEQRES  12 B  258  ILE THR THR MET LYS LEU GLY PHE VAL LEU LEU ARG GLU
SEQRES  13 B  258  ASN LEU PHE THR LYS CYS THR ASP GLY GLU TYR GLU LEU
SEQRES  14 B  258  ALA LYS MET VAL MET ARG LYS GLY SER LEU PHE GLN ASN
SEQRES  15 B  258  VAL LEU ALA GLN ARG PRO LYS PHE THR GLU LYS GLY TYR
SEQRES  16 B  258  GLY SER ILE LYS LYS VAL TYR ILE TRP THR ASP GLN ASP
SEQRES  17 B  258  LYS ILE PHE LEU PRO ASP PHE GLN ARG TRP GLN ILE ALA
SEQRES  18 B  258  ASN TYR LYS PRO ASP LYS VAL TYR GLN VAL GLN GLY GLY
SEQRES  19 B  258  ASP HIS LYS LEU GLN LEU THR LYS THR GLU GLU VAL ALA
SEQRES  20 B  258  HIS ILE LEU GLN GLU VAL ALA ASP ALA TYR ALA
SEQRES   1 C  258  MET VAL THR ALA HIS PHE VAL LEU ILE HIS THR ILE CYS
SEQRES   2 C  258  HIS GLY ALA TRP ILE TRP HIS LYS LEU LYS PRO ALA LEU
SEQRES   3 C  258  GLU ARG ALA GLY HIS LYS VAL THR ALA LEU ASP MET ALA
SEQRES   4 C  258  ALA SER GLY ILE ASP PRO ARG GLN ILE GLU GLN ILE ASN
SEQRES   5 C  258  SER PHE ASP GLU TYR SER GLU PRO LEU LEU THR PHE LEU
SEQRES   6 C  258  GLU LYS LEU PRO GLN GLY GLU LYS VAL ILE ILE VAL GLY
SEQRES   7 C  258  GLU SER CYS ALA GLY LEU ASN ILE ALA ILE ALA ALA ASP
SEQRES   8 C  258  ARG TYR VAL ASP LYS ILE ALA ALA GLY VAL PHE LEU ASN
SEQRES   9 C  258  SER LEU LEU PRO ASP THR VAL HIS SER PRO SER TYR THR
SEQRES  10 C  258  VAL GLU LYS LEU LEU GLU SER PHE PRO ASP TRP ARG ASP
SEQRES  11 C  258  THR GLU TYR PHE THR PHE THR ASN ILE THR GLY GLU THR
SEQRES  12 C  258  ILE THR THR MET LYS LEU GLY PHE VAL LEU LEU ARG GLU
SEQRES  13 C  258  ASN LEU PHE THR LYS CYS THR ASP GLY GLU TYR GLU LEU
SEQRES  14 C  258  ALA LYS MET VAL MET ARG LYS GLY SER LEU PHE GLN ASN
SEQRES  15 C  258  VAL LEU ALA GLN ARG PRO LYS PHE THR GLU LYS GLY TYR
SEQRES  16 C  258  GLY SER ILE LYS LYS VAL TYR ILE TRP THR ASP GLN ASP
SEQRES  17 C  258  LYS ILE PHE LEU PRO ASP PHE GLN ARG TRP GLN ILE ALA
SEQRES  18 C  258  ASN TYR LYS PRO ASP LYS VAL TYR GLN VAL GLN GLY GLY
SEQRES  19 C  258  ASP HIS LYS LEU GLN LEU THR LYS THR GLU GLU VAL ALA
SEQRES  20 C  258  HIS ILE LEU GLN GLU VAL ALA ASP ALA TYR ALA
SEQRES   1 D  258  MET VAL THR ALA HIS PHE VAL LEU ILE HIS THR ILE CYS
SEQRES   2 D  258  HIS GLY ALA TRP ILE TRP HIS LYS LEU LYS PRO ALA LEU
SEQRES   3 D  258  GLU ARG ALA GLY HIS LYS VAL THR ALA LEU ASP MET ALA
SEQRES   4 D  258  ALA SER GLY ILE ASP PRO ARG GLN ILE GLU GLN ILE ASN
SEQRES   5 D  258  SER PHE ASP GLU TYR SER GLU PRO LEU LEU THR PHE LEU
SEQRES   6 D  258  GLU LYS LEU PRO GLN GLY GLU LYS VAL ILE ILE VAL GLY
SEQRES   7 D  258  GLU SER CYS ALA GLY LEU ASN ILE ALA ILE ALA ALA ASP
SEQRES   8 D  258  ARG TYR VAL ASP LYS ILE ALA ALA GLY VAL PHE LEU ASN
SEQRES   9 D  258  SER LEU LEU PRO ASP THR VAL HIS SER PRO SER TYR THR
SEQRES  10 D  258  VAL GLU LYS LEU LEU GLU SER PHE PRO ASP TRP ARG ASP
SEQRES  11 D  258  THR GLU TYR PHE THR PHE THR ASN ILE THR GLY GLU THR
SEQRES  12 D  258  ILE THR THR MET LYS LEU GLY PHE VAL LEU LEU ARG GLU
SEQRES  13 D  258  ASN LEU PHE THR LYS CYS THR ASP GLY GLU TYR GLU LEU
SEQRES  14 D  258  ALA LYS MET VAL MET ARG LYS GLY SER LEU PHE GLN ASN
SEQRES  15 D  258  VAL LEU ALA GLN ARG PRO LYS PHE THR GLU LYS GLY TYR
SEQRES  16 D  258  GLY SER ILE LYS LYS VAL TYR ILE TRP THR ASP GLN ASP
SEQRES  17 D  258  LYS ILE PHE LEU PRO ASP PHE GLN ARG TRP GLN ILE ALA
SEQRES  18 D  258  ASN TYR LYS PRO ASP LYS VAL TYR GLN VAL GLN GLY GLY
SEQRES  19 D  258  ASP HIS LYS LEU GLN LEU THR LYS THR GLU GLU VAL ALA
SEQRES  20 D  258  HIS ILE LEU GLN GLU VAL ALA ASP ALA TYR ALA
FORMUL   5  HOH   *60(H2 O)
HELIX    1 AA1 GLY A   15  HIS A   20  5                                   6
HELIX    2 AA2 LYS A   21  ALA A   29  1                                   9
HELIX    3 AA3 GLN A   47  ILE A   51  5                                   5
HELIX    4 AA4 SER A   53  SER A   58  1                                   6
HELIX    5 AA5 SER A   58  LEU A   68  1                                  11
HELIX    6 AA6 ALA A   82  ASP A   91  1                                  10
HELIX    7 AA7 TYR A   93  ASP A   95  5                                   3
HELIX    8 AA8 SER A  115  PHE A  125  1                                  11
HELIX    9 AA9 GLY A  150  ASN A  157  1                                   8
HELIX   10 AB1 THR A  163  MET A  174  1                                  12
HELIX   11 AB2 PHE A  180  GLN A  186  1                                   7
HELIX   12 AB3 GLY A  194  ILE A  198  5                                   5
HELIX   13 AB4 LEU A  212  TYR A  223  1                                  12
HELIX   14 AB5 LYS A  237  LYS A  242  1                                   6
HELIX   15 AB6 LYS A  242  ALA A  258  1                                  17
HELIX   16 AB7 GLY B   15  HIS B   20  5                                   6
HELIX   17 AB8 LYS B   21  ALA B   29  1                                   9
HELIX   18 AB9 GLN B   47  ILE B   51  5                                   5
HELIX   19 AC1 SER B   53  SER B   58  1                                   6
HELIX   20 AC2 SER B   58  LEU B   68  1                                  11
HELIX   21 AC3 ALA B   82  VAL B   94  1                                  13
HELIX   22 AC4 SER B  115  PHE B  125  1                                  11
HELIX   23 AC5 GLY B  150  ASN B  157  1                                   8
HELIX   24 AC6 THR B  163  MET B  174  1                                  12
HELIX   25 AC7 PHE B  180  GLN B  186  1                                   7
HELIX   26 AC8 GLY B  194  ILE B  198  5                                   5
HELIX   27 AC9 LEU B  212  TYR B  223  1                                  12
HELIX   28 AD1 LYS B  237  LYS B  242  1                                   6
HELIX   29 AD2 LYS B  242  ALA B  258  1                                  17
HELIX   30 AD3 GLY C   15  HIS C   20  5                                   6
HELIX   31 AD4 LYS C   21  ARG C   28  1                                   8
HELIX   32 AD5 GLN C   47  ILE C   51  5                                   5
HELIX   33 AD6 SER C   53  SER C   58  1                                   6
HELIX   34 AD7 SER C   58  LEU C   68  1                                  11
HELIX   35 AD8 ALA C   82  ASP C   91  1                                  10
HELIX   36 AD9 TYR C   93  ASP C   95  5                                   3
HELIX   37 AE1 SER C  115  PHE C  125  1                                  11
HELIX   38 AE2 GLY C  150  ASN C  157  1                                   8
HELIX   39 AE3 THR C  163  MET C  174  1                                  12
HELIX   40 AE4 PHE C  180  GLN C  186  1                                   7
HELIX   41 AE5 GLY C  194  ILE C  198  5                                   5
HELIX   42 AE6 LEU C  212  TYR C  223  1                                  12
HELIX   43 AE7 LYS C  237  LYS C  242  1                                   6
HELIX   44 AE8 LYS C  242  ALA C  258  1                                  17
HELIX   45 AE9 GLY D   15  HIS D   20  5                                   6
HELIX   46 AF1 LYS D   21  ALA D   29  1                                   9
HELIX   47 AF2 GLN D   47  ILE D   51  5                                   5
HELIX   48 AF3 SER D   53  SER D   58  1                                   6
HELIX   49 AF4 SER D   58  LEU D   68  1                                  11
HELIX   50 AF5 ALA D   82  ASP D   91  1                                  10
HELIX   51 AF6 TYR D   93  ASP D   95  5                                   3
HELIX   52 AF7 SER D  115  PHE D  125  1                                  11
HELIX   53 AF8 GLY D  150  ASN D  157  1                                   8
HELIX   54 AF9 THR D  163  MET D  174  1                                  12
HELIX   55 AG1 PHE D  180  GLN D  186  1                                   7
HELIX   56 AG2 GLY D  194  ILE D  198  5                                   5
HELIX   57 AG3 LEU D  212  TYR D  223  1                                  12
HELIX   58 AG4 LYS D  237  LYS D  242  1                                   6
HELIX   59 AG5 LYS D  242  ALA D  258  1                                  17
SHEET    1 AA1 6 LYS A  32  LEU A  36  0
SHEET    2 AA1 6 HIS A   5  ILE A   9  1  N  LEU A   8   O  THR A  34
SHEET    3 AA1 6 VAL A  74  GLU A  79  1  O  VAL A  77   N  ILE A   9
SHEET    4 AA1 6 ILE A  97  LEU A 103  1  O  VAL A 101   N  ILE A  76
SHEET    5 AA1 6 LYS A 200  TRP A 204  1  O  ILE A 203   N  PHE A 102
SHEET    6 AA1 6 LYS A 227  GLN A 230  1  O  TYR A 229   N  TYR A 202
SHEET    1 AA2 2 GLU A 132  THR A 137  0
SHEET    2 AA2 2 THR A 143  LYS A 148 -1  O  ILE A 144   N  PHE A 136
SHEET    1 AA3 6 LYS B  32  LEU B  36  0
SHEET    2 AA3 6 HIS B   5  ILE B   9  1  N  LEU B   8   O  THR B  34
SHEET    3 AA3 6 VAL B  74  GLU B  79  1  O  VAL B  77   N  ILE B   9
SHEET    4 AA3 6 ILE B  97  LEU B 103  1  O  VAL B 101   N  ILE B  76
SHEET    5 AA3 6 LYS B 200  TRP B 204  1  O  ILE B 203   N  PHE B 102
SHEET    6 AA3 6 LYS B 227  GLN B 230  1  O  TYR B 229   N  TYR B 202
SHEET    1 AA4 3 GLU B 132  THR B 137  0
SHEET    2 AA4 3 THR B 143  LYS B 148 -1  O  ILE B 144   N  PHE B 136
SHEET    3 AA4 3 GLY B 177  SER B 178 -1  O  GLY B 177   N  MET B 147
SHEET    1 AA5 6 LYS C  32  LEU C  36  0
SHEET    2 AA5 6 HIS C   5  ILE C   9  1  N  LEU C   8   O  THR C  34
SHEET    3 AA5 6 VAL C  74  GLU C  79  1  O  VAL C  77   N  ILE C   9
SHEET    4 AA5 6 ILE C  97  LEU C 103  1  O  VAL C 101   N  ILE C  76
SHEET    5 AA5 6 LYS C 200  TRP C 204  1  O  ILE C 203   N  PHE C 102
SHEET    6 AA5 6 LYS C 227  GLN C 230  1  O  TYR C 229   N  TYR C 202
SHEET    1 AA6 3 GLU C 132  THR C 137  0
SHEET    2 AA6 3 THR C 143  LYS C 148 -1  O  ILE C 144   N  PHE C 136
SHEET    3 AA6 3 GLY C 177  SER C 178 -1  O  GLY C 177   N  MET C 147
SHEET    1 AA7 6 LYS D  32  LEU D  36  0
SHEET    2 AA7 6 HIS D   5  ILE D   9  1  N  LEU D   8   O  THR D  34
SHEET    3 AA7 6 VAL D  74  GLU D  79  1  O  VAL D  77   N  ILE D   9
SHEET    4 AA7 6 ILE D  97  LEU D 103  1  O  VAL D 101   N  ILE D  76
SHEET    5 AA7 6 LYS D 200  TRP D 204  1  O  ILE D 203   N  PHE D 102
SHEET    6 AA7 6 LYS D 227  GLN D 230  1  O  TYR D 229   N  TYR D 202
SHEET    1 AA8 3 GLU D 132  THR D 137  0
SHEET    2 AA8 3 THR D 143  LYS D 148 -1  O  ILE D 144   N  PHE D 136
SHEET    3 AA8 3 GLY D 177  SER D 178 -1  O  GLY D 177   N  MET D 147
CRYST1   86.469   88.098  135.213  90.00  90.00  90.00 P 21 21 21   16
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.011565  0.000000  0.000000        0.00000
SCALE2      0.000000  0.011351  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007396        0.00000
TER    2073      ALA A 258
TER    4146      ALA B 258
TER    6219      ALA C 258
TER    8292      ALA D 258
MASTER      319    0    0   59   35    0    0    6 8348    4    0   80
END