longtext: 5AH1-pdb

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HEADER    HYDROLASE                               04-FEB-15   5AH1
TITLE     STRUCTURE OF ESTA FROM CLOSTRIDIUM BOTULINUM
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: TRIACYLGLYCEROL LIPASE;
COMPND   3 CHAIN: A;
COMPND   4 FRAGMENT: RESIDUES 30-480;
COMPND   5 SYNONYM: ESTERASE A;
COMPND   6 EC: 3.1.1.3;
COMPND   7 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: CLOSTRIDIUM BOTULINUM;
SOURCE   3 ORGANISM_TAXID: 1491;
SOURCE   4 ATCC: 3502;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   8 EXPRESSION_SYSTEM_PLASMID: PET26B(PLUS)
KEYWDS    HYDROLASE, POLYESTERASE, POLYMER HYDROLYSIS, ZINC BINDING,
KEYWDS   2 ALPHA/BETA- HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    A.PAIRITSCH,A.LYSKOWSKI,A.HROMIC,G.STEINKELLNER,K.GRUBER,V.PERZ,
AUTHOR   2 A.BAUMSCHLAGER,K.BLEYMAIER,S.ZITZENBACHER,C.SINKEL,U.KUEPER,
AUTHOR   3 D.RIBITSCH,G.M.GUEBITZ
REVDAT   1   11-NOV-15 5AH1    0
JRNL        AUTH   V.PERZ,A.BAUMSCHLAGER,K.BLEYMAIER,S.ZITZENBACHER,A.HROMIC,
JRNL        AUTH 2 G.STEINKELLNER,A.PAIRITSCH,A.LYSKOWSKI,K.GRUBER,C.SINKEL,
JRNL        AUTH 3 U.KUEPER,D.RIBITSCH,G.M.GUEBITZ
JRNL        TITL   HYDROLYSIS OF SYNTHETIC POLYESTERS BY CLOSTRIDIUM BOTULINUM
JRNL        TITL 2 ESTERASES.
JRNL        REF    BIOTECHNOL.BIOENG.                         2015
JRNL        REFN                   ESSN 1097-0290
JRNL        PMID   26524601
JRNL        DOI    10.1002/BIT.25874
REMARK   2
REMARK   2 RESOLUTION.    1.20 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-
REMARK   3               : KUNSTLEVE,LI-WEI HUNG,ROBERT IMMORMINO,
REMARK   3               : TOM IOERGER,AIRLIE MCCOY,ERIK MCKEE,NIGEL
REMARK   3               : MORIARTY,REETAL PAI,RANDY READ,JANE
REMARK   3               : RICHARDSON,DAVID RICHARDSON,TOD ROMO,JIM
REMARK   3               : SACCHETTINI,NICHOLAS SAUTER,JACOB SMITH,
REMARK   3               : LAURENT STORONI,TOM TERWILLIGER,PETER
REMARK   3               : ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.20
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 34.954
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 0.00
REMARK   3   COMPLETENESS FOR RANGE        (%) : 82.75
REMARK   3   NUMBER OF REFLECTIONS             : 113119
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.1296
REMARK   3   R VALUE            (WORKING SET) : 0.1293
REMARK   3   FREE R VALUE                     : 0.1493
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 1.7
REMARK   3   FREE R VALUE TEST SET COUNT      : 1876
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 34.9694 -  2.8164    0.99    10470   175  0.1289 0.1423
REMARK   3     2  2.8164 -  2.2356    0.99    10234   169  0.1258 0.1307
REMARK   3     3  2.2356 -  1.9530    0.97    10063   168  0.1198 0.1397
REMARK   3     4  1.9530 -  1.7744    0.95     9814   171  0.1256 0.1522
REMARK   3     5  1.7744 -  1.6473    0.94     9699   166  0.1261 0.1369
REMARK   3     6  1.6473 -  1.5501    0.92     9534   154  0.1175 0.1500
REMARK   3     7  1.5501 -  1.4725    0.90     9246   163  0.1227 0.1613
REMARK   3     8  1.4725 -  1.4084    0.89     9210   145  0.1306 0.1607
REMARK   3     9  1.4084 -  1.3542    0.87     8896   164  0.1381 0.1942
REMARK   3    10  1.3542 -  1.3075    0.83     8519   136  0.1489 0.1869
REMARK   3    11  1.3075 -  1.2666    0.66     6761   133  0.1636 0.1927
REMARK   3    12  1.2666 -  1.2304    0.49     5043    72  0.1669 0.2357
REMARK   3    13  1.2304 -  1.1980    0.36     3754    60  0.1956 0.1770
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.09
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 16.23
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 8.85
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 15.2
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.008           3685
REMARK   3   ANGLE     :  1.182           5041
REMARK   3   CHIRALITY :  0.070            537
REMARK   3   PLANARITY :  0.007            670
REMARK   3   DIHEDRAL  : 13.093           1373
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: ANISOTROPIC ATOMIC DISPLACEMENT
REMARK   3  PARAMETERS WERE REFINED FOR ALL NON-H ATOMS EXCEPT FOR THE SOLVENT
REMARK   4
REMARK   4 5AH1 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 04-FEB-15.
REMARK 100 THE PDBE ID CODE IS EBI-62956.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 10-JUN-13
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : BM30A
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9777
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : NULL
REMARK 200  DETECTOR MANUFACTURER          : NULL
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 121016
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.20
REMARK 200  RESOLUTION RANGE LOW       (A) : 49.02
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.0
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 88.6
REMARK 200  DATA REDUNDANCY                : 4.9
REMARK 200  R MERGE                    (I) : 0.06
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 13.00
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.20
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.26
REMARK 200  COMPLETENESS FOR SHELL     (%) : 50.5
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.8
REMARK 200  R MERGE FOR SHELL          (I) : 0.41
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.80
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: PDB ENTRY 1KU0
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 43
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.1
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1 M TRIS, PH 8.5 AND 2 M AMMONIUM
REMARK 280  SULFATE
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       32.35500
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     ALA A     2
REMARK 465     GLU A     3
REMARK 465     PRO A     4
REMARK 465     LYS A     5
REMARK 465     ALA A     6
REMARK 465     GLN A     7
REMARK 465     GLY A     8
REMARK 465     THR A     9
REMARK 465     GLN A    10
REMARK 465     LYS A    11
REMARK 465     VAL A    12
REMARK 465     GLU A    13
REMARK 465     SER A    14
REMARK 465     SER A    15
REMARK 465     THR A    16
REMARK 465     THR A    17
REMARK 465     LYS A    18
REMARK 465     LYS A    19
REMARK 465     GLU A    20
REMARK 465     VAL A    21
REMARK 465     LYS A    22
REMARK 465     ASP A    23
REMARK 465     ALA A    24
REMARK 465     GLU A    25
REMARK 465     GLU A    26
REMARK 465     THR A    27
REMARK 465     HIS A   456
REMARK 465     HIS A   457
REMARK 465     HIS A   458
REMARK 465     HIS A   459
REMARK 465     HIS A   460
REMARK 465     HIS A   461
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     GLU A 455    CA   C    O    CB   CG   CD   OE1  OE2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OE2  GLU A   201     O    HOH A  2349              2.20
REMARK 500   O    HOH A  2033     O    HOH A  2089              2.19
REMARK 500   O    HOH A  2043     O    HOH A  2601              2.14
REMARK 500   O    HOH A  2051     O    HOH A  2141              2.11
REMARK 500   O    HOH A  2052     O    HOH A  2142              2.08
REMARK 500   O    HOH A  2119     O    HOH A  2262              2.16
REMARK 500   O    HOH A  2202     O    HOH A  2203              2.16
REMARK 500   O    HOH A  2299     O    HOH A  2550              2.13
REMARK 500   O    HOH A  2354     O    HOH A  2371              2.09
REMARK 500   O    HOH A  2472     O    HOH A  2474              2.13
REMARK 500   O    HOH A  2560     O    HOH A  2561              2.18
REMARK 500   O    HOH A  2594     O    HOH A  2595              2.20
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O    HOH A  2107     O    HOH A  2472     2555     2.16
REMARK 500   O    HOH A  2185     O    HOH A  2470     2555     2.20
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    THR A  32     -169.74   -122.49
REMARK 500    SER A 182     -139.31     59.84
REMARK 500    LEU A 271       33.72    -96.64
REMARK 500    ALA A 335       22.80   -143.96
REMARK 500    ASP A 377     -169.36   -126.09
REMARK 500    VAL A 386      -39.20   -132.27
REMARK 500    LYS A 396      -42.89   -130.32
REMARK 500
REMARK 500 REMARK: NULL
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                               K A1456   K
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASP A 425   OD2
REMARK 620 2 HOH A2010   O    84.7
REMARK 620 3 ASP A 428   OD2  81.8 131.3
REMARK 620 4 PRO A 351   O    85.0 144.0  80.8
REMARK 620 5 HOH A2012   O   115.8  76.4 150.2  77.4
REMARK 620 6 ASP A 428   OD1  84.9  80.6  51.8 132.5 147.0
REMARK 620 7 HOH A2582   O   153.9  85.5  86.7 116.3  85.1  69.7
REMARK 620 N                    1     2     3     4     5     6
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                               K A1457   K
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 VAL A  67   O
REMARK 620 2 VAL A  67   O     2.8
REMARK 620 3 ARG A  70   O    74.6  76.9
REMARK 620 4 HOH A2087   O    61.1  61.0  56.4
REMARK 620 5 SER A 237   O   141.4 139.6 141.2 141.9
REMARK 620 6 ASP A 238   O    85.6  86.6 102.9 143.6  73.8
REMARK 620 7 HOH A2090   O   152.7 155.2  78.4 105.9  64.1  97.2
REMARK 620 8 MET A 240   O    85.9  83.1 143.3  87.0  69.7 106.2 118.8
REMARK 620 N                    1     2     3     4     5     6     7
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              ZN A1455  ZN
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASP A 302   OD2
REMARK 620 2 HIS A 150   NE2 104.5
REMARK 620 3 HIS A 156   NE2  96.5 104.6
REMARK 620 4 ASP A 130   OD1 131.8  97.9 118.4
REMARK 620 N                    1     2     3
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  ZN A1455
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE   K A1456
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE   K A1457
DBREF  5AH1 A    2   452  UNP    A7FSB2   A7FSB2_CLOB1    30    480
SEQADV 5AH1 MET A    1  UNP  A7FSB2              CLONING ARTIFACT
SEQADV 5AH1 ALA A  453  UNP  A7FSB2              EXPRESSION TAG
SEQADV 5AH1 LEU A  454  UNP  A7FSB2              EXPRESSION TAG
SEQADV 5AH1 GLU A  455  UNP  A7FSB2              EXPRESSION TAG
SEQADV 5AH1 HIS A  456  UNP  A7FSB2              EXPRESSION TAG
SEQADV 5AH1 HIS A  457  UNP  A7FSB2              EXPRESSION TAG
SEQADV 5AH1 HIS A  458  UNP  A7FSB2              EXPRESSION TAG
SEQADV 5AH1 HIS A  459  UNP  A7FSB2              EXPRESSION TAG
SEQADV 5AH1 HIS A  460  UNP  A7FSB2              EXPRESSION TAG
SEQADV 5AH1 HIS A  461  UNP  A7FSB2              EXPRESSION TAG
SEQRES   1 A  461  MET ALA GLU PRO LYS ALA GLN GLY THR GLN LYS VAL GLU
SEQRES   2 A  461  SER SER THR THR LYS LYS GLU VAL LYS ASP ALA GLU GLU
SEQRES   3 A  461  THR ILE LYS ILE PRO THR LEU GLU ASP ILE ASP ASN LEU
SEQRES   4 A  461  ILE ASP SER ALA GLU GLU VAL LYS SER GLU GLU ASP ILE
SEQRES   5 A  461  ASN LYS MET PRO PRO LEU LYS PHE PRO VAL GLU PHE PRO
SEQRES   6 A  461  GLU VAL ASN THR ARG SER ILE ILE GLY GLY ASN ASN TYR
SEQRES   7 A  461  PRO ILE VAL LEU VAL HIS GLY PHE MET GLY PHE GLY ARG
SEQRES   8 A  461  ASP GLU LEU LEU GLY TYR LYS TYR TRP GLY GLY VAL VAL
SEQRES   9 A  461  ASP LEU GLN GLU LYS LEU ASN ALA SER GLY HIS GLU THR
SEQRES  10 A  461  TYR THR ALA THR VAL GLY PRO VAL SER SER ASN TRP ASP
SEQRES  11 A  461  ARG ALA CYS GLU LEU TYR ALA TYR ILE VAL GLY GLY THR
SEQRES  12 A  461  VAL ASP TYR GLY GLU ALA HIS ALA LYS LYS PHE LYS HIS
SEQRES  13 A  461  ASN ARG TYR GLY ARG THR TYR PRO GLY ILE TYR LYS ASN
SEQRES  14 A  461  ILE SER ASN GLU ASN LYS ILE HIS LEU ILE GLY HIS SER
SEQRES  15 A  461  MET GLY GLY GLN THR ILE ARG THR LEU THR GLN LEU LEU
SEQRES  16 A  461  SER GLU GLY SER GLU GLU GLU ILE ASN CYS GLY GLN GLU
SEQRES  17 A  461  ASN ILE SER PRO LEU PHE GLU GLY GLY LYS HIS TRP ILE
SEQRES  18 A  461  HIS SER VAL SER THR ILE SER THR PRO ASN ASP GLY THR
SEQRES  19 A  461  THR LEU SER ASP LEU MET PRO ALA LYS ASP LEU ILE SER
SEQRES  20 A  461  TYR THR PHE GLY VAL LEU GLY THR ILE THR GLY LYS ASN
SEQRES  21 A  461  LYS LEU PHE SER SER ILE TYR ASP LEU LYS LEU ASP GLN
SEQRES  22 A  461  TRP GLY LEU LYS LYS GLN ASN GLY GLU SER GLN ARG ASP
SEQRES  23 A  461  TYR ILE GLU ARG VAL LEU ASP SER ASN ILE TRP ASN SER
SEQRES  24 A  461  THR LYS ASP ILE ALA THR TYR ASP LEU SER THR GLU GLY
SEQRES  25 A  461  ALA GLN GLU LEU ASN THR TRP VAL LYS ALA GLN PRO ASP
SEQRES  26 A  461  VAL TYR TYR PHE SER TRP THR THR GLN ALA THR LYS GLU
SEQRES  27 A  461  SER ILE LEU THR GLY HIS SER VAL ALA GLN ILE GLY PRO
SEQRES  28 A  461  MET ASN PRO ILE PHE TYR PRO THR ALA ASN LEU MET GLY
SEQRES  29 A  461  ARG TYR SER ARG ASN GLN LYS ASP LEU PRO ILE ILE ASP
SEQRES  30 A  461  LYS LYS TRP PHE PRO ASN ASP GLY VAL VAL ASN CYS ILE
SEQRES  31 A  461  SER GLN ASP GLY PRO LYS LEU GLY SER ASN ASP VAL ILE
SEQRES  32 A  461  GLU GLN TYR ASN GLY GLY VAL LYS ILE GLY GLN TRP ASN
SEQRES  33 A  461  ALA MET PRO ARG ILE ILE ASN THR ASP HIS MET ASP ILE
SEQRES  34 A  461  VAL GLY THR PHE GLY ASN VAL LYS ASP TRP TYR MET ASP
SEQRES  35 A  461  TYR ALA SER PHE LEU SER ASN LEU SER ARG ALA LEU GLU
SEQRES  36 A  461  HIS HIS HIS HIS HIS HIS
HET     ZN  A1455       1
HET      K  A1456       1
HET      K  A1457       1
HETNAM       K POTASSIUM ION
HETNAM      ZN ZINC ION
FORMUL   2    K    2(K 1+)
FORMUL   3   ZN    ZN 2+
FORMUL   4  HOH   *602(H2 O)
HELIX    1   1 THR A   32  GLU A   34  5                                   3
HELIX    2   2 ASP A   35  ALA A   43  1                                   9
HELIX    3   3 SER A   48  ASN A   53  1                                   6
HELIX    4   4 GLU A   93  TYR A   97  5                                   5
HELIX    5   5 ASP A  105  SER A  113  1                                   9
HELIX    6   6 SER A  127  GLY A  141  1                                  15
HELIX    7   7 GLY A  147  LYS A  155  1                                   9
HELIX    8   8 SER A  182  GLY A  198  1                                  17
HELIX    9   9 SER A  199  CYS A  205  1                                   7
HELIX   10  10 SER A  211  GLY A  216  5                                   6
HELIX   11  11 THR A  234  MET A  240  1                                   7
HELIX   12  12 PRO A  241  GLY A  258  1                                  18
HELIX   13  13 ASN A  260  SER A  264  5                                   5
HELIX   14  14 LEU A  271  GLY A  275  5                                   5
HELIX   15  15 SER A  283  ASP A  293  1                                  11
HELIX   16  16 ASN A  295  THR A  300  1                                   6
HELIX   17  17 ILE A  303  SER A  309  1                                   7
HELIX   18  18 SER A  309  THR A  318  1                                  10
HELIX   19  19 ASN A  353  ILE A  355  5                                   3
HELIX   20  20 PHE A  356  GLY A  364  1                                   9
HELIX   21  21 ASP A  377  PHE A  381  5                                   5
HELIX   22  22 CYS A  389  GLN A  392  5                                   4
HELIX   23  23 MET A  427  GLY A  431  5                                   5
HELIX   24  24 VAL A  436  ASN A  449  1                                  14
SHEET    1  AA 7 THR A 117  ALA A 120  0
SHEET    2  AA 7 ILE A  80  VAL A  83  1  O  ILE A  80   N  TYR A 118
SHEET    3  AA 7 ILE A 176  HIS A 181  1  O  HIS A 177   N  VAL A  81
SHEET    4  AA 7 ILE A 221  ILE A 227  1  N  HIS A 222   O  ILE A 176
SHEET    5  AA 7 TYR A 327  TRP A 331  1  O  TYR A 327   N  VAL A 224
SHEET    6  AA 7 TRP A 415  ALA A 417  1  O  ASN A 416   N  SER A 330
SHEET    7  AA 7 ILE A 403  GLN A 405  1  O  GLU A 404   N  ALA A 417
SHEET    1  AB 2 GLY A 142  ASP A 145  0
SHEET    2  AB 2 TYR A 159  TYR A 163 -1  N  GLY A 160   O  VAL A 144
SHEET    1  AC 2 THR A 336  GLU A 338  0
SHEET    2  AC 2 SER A 345  ALA A 347 -1  O  VAL A 346   N  LYS A 337
LINK        ZN    ZN A1455                 OD2 ASP A 302     1555   1555  1.97
LINK        ZN    ZN A1455                 OD1 ASP A 130     1555   1555  1.94
LINK        ZN    ZN A1455                 NE2 HIS A 156     1555   1555  2.07
LINK        ZN    ZN A1455                 NE2 HIS A 150     1555   1555  2.08
LINK         K     K A1456                 O   HOH A2012     1555   1555  2.44
LINK         K     K A1456                 OD1 ASP A 428     1555   1555  2.57
LINK         K     K A1456                 O   HOH A2582     1555   1555  2.56
LINK         K     K A1456                 O   HOH A2010     1555   1555  2.44
LINK         K     K A1456                 OD2 ASP A 428     1555   1555  2.47
LINK         K     K A1456                 O   PRO A 351     1555   1555  2.42
LINK         K     K A1456                 OD2 ASP A 425     1555   1555  2.47
LINK         K     K A1457                 O   ASP A 238     1555   1555  2.69
LINK         K     K A1457                 O   HOH A2090     1555   1455  2.80
LINK         K     K A1457                 O   MET A 240     1555   1555  2.68
LINK         K     K A1457                 O   SER A 237     1555   1555  2.77
LINK         K     K A1457                 O   HOH A2087     1555   1455  2.95
LINK         K     K A1457                 O   ARG A  70     1555   1455  2.91
LINK         K     K A1457                 O  BVAL A  67     1555   1455  2.87
LINK         K     K A1457                 O  AVAL A  67     1555   1455  2.78
CISPEP   1 THR A  432    PHE A  433          0         7.01
SITE     1 AC1  4 ASP A 130  HIS A 150  HIS A 156  ASP A 302
SITE     1 AC2  7 PRO A 351  ASN A 353  ASP A 425  ASP A 428
SITE     2 AC2  7 HOH A2010  HOH A2012  HOH A2582
SITE     1 AC3  7 VAL A  67  ARG A  70  SER A 237  ASP A 238
SITE     2 AC3  7 MET A 240  HOH A2087  HOH A2090
CRYST1   51.240   64.710   69.790  90.00 106.93  90.00 P 1 21 1      2
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.019516  0.000000  0.005941        0.00000
SCALE2      0.000000  0.015454  0.000000        0.00000
SCALE3      0.000000  0.000000  0.014978        0.00000
TER    7077      GLU A 455
MASTER      358    0    3   24   11    0    5    6 7683    1   18   36
END