longtext: 5AO9-pdb

content
HEADER    HYDROLASE                               10-SEP-15   5AO9
TITLE     THE STRUCTURE OF A NOVEL THERMOPHILIC ESTERASE FROM THE
TITLE    2 PLANCTOMYCETES SPECIES, THERMOGUTTA TERRIFONTIS, EST2-NATIVE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ESTERASE;
COMPND   3 CHAIN: A;
COMPND   4 EC: 3.1.1.1;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: THERMOGUTTA TERRIFONTIS;
SOURCE   3 ORGANISM_TAXID: 1331910;
SOURCE   4 STRAIN: R1;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   9 EXPRESSION_SYSTEM_VECTOR: PLATE31
KEYWDS    HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    C.SAYER,Z.SZABO,M.N.ISUPOV,C.INGHAM,J.A.LITTLECHILD
REVDAT   1   09-DEC-15 5AO9    0
JRNL        AUTH   C.SAYER,Z.SZABO,M.N.ISUPOV,C.INGHAM,J.A.LITTLECHILD
JRNL        TITL   THE STRUCTURE OF A NOVEL THERMOPHILIC ESTERASE FROM THE
JRNL        TITL 2 PLANCTOMYCETES SPECIES, THERMOGUTTA TERRIFONTIS REVEALS AN
JRNL        TITL 3 OPEN ACTIVE SITE DUE TO A MINIMAL 'CAP' DOMAIN
JRNL        REF    FRONT.MICROBIOL.                           2015
JRNL        REFN                   ESSN 1664-302X
JRNL        DOI    10.3389/FMICB.2015.01294
REMARK   2
REMARK   2 RESOLUTION.    1.58 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0123
REMARK   3   AUTHORS     : MURSHUDOV,SKUBAK,LEBEDEV,PANNU,
REMARK   3                 STEINER,NICHOLLS,WINN,LONG,VAGIN
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.58
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 47.83
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NONE
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.87
REMARK   3   NUMBER OF REFLECTIONS             : 44425
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.15136
REMARK   3   R VALUE            (WORKING SET) : 0.15013
REMARK   3   FREE R VALUE                     : 0.17490
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.9
REMARK   3   FREE R VALUE TEST SET COUNT      : 2290
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.575
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.616
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 3260
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 99.82
REMARK   3   BIN R VALUE           (WORKING SET) : 0.286
REMARK   3   BIN FREE R VALUE SET COUNT          : 138
REMARK   3   BIN FREE R VALUE                    : 0.289
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 2428
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 108
REMARK   3   SOLVENT ATOMS            : 208
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 29.383
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.70
REMARK   3    B22 (A**2) : -1.28
REMARK   3    B33 (A**2) : 0.57
REMARK   3    B12 (A**2) : 0.00
REMARK   3    B13 (A**2) : 0.00
REMARK   3    B23 (A**2) : 0.00
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.069
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.070
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.049
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 1.431
REMARK   3
REMARK   3  CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.974
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.970
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  2618 ; 0.010 ; 0.019
REMARK   3   BOND LENGTHS OTHERS               (A):  2591 ; 0.002 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  3551 ; 1.486 ; 1.983
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  6015 ; 0.989 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   352 ; 5.731 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   132 ;34.211 ;22.955
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   452 ;15.258 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    29 ;18.973 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   362 ; 0.097 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  2969 ; 0.006 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):   624 ; 0.002 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  1205 ; 3.778 ; 6.490
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  1204 ; 3.765 ; 6.474
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  1523 ; 5.087 ;10.826
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  1413 ; 5.398 ; 8.324
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ANISOTROPIC THERMAL FACTOR RESTRAINTS.   COUNT   RMS    WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE
REMARK   3   RIDING POSITIONS.
REMARK   4
REMARK   4 5AO9 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 10-SEP-15.
REMARK 100 THE PDBE ID CODE IS EBI-64891.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : NULL
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : NULL
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : DIAMOND
REMARK 200  BEAMLINE                       : I04-1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.91741
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : NULL
REMARK 200  DATA SCALING SOFTWARE          : NULL
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 46778
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.58
REMARK 200  RESOLUTION RANGE LOW       (A) : 47.80
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 2.0
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.9
REMARK 200  DATA REDUNDANCY                : 6.7
REMARK 200  R MERGE                    (I) : 0.07
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 13.30
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.58
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.62
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.9
REMARK 200  DATA REDUNDANCY IN SHELL       : 7.0
REMARK 200  R MERGE FOR SHELL          (I) : 0.87
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.00
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: PDB ENTRY 1EVQ
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 53
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.64
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       30.82000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       37.92000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       35.53500
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       37.92000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       30.82000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       35.53500
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     ASP A   173
REMARK 465     HIS A   174
REMARK 465     VAL A   175
REMARK 465     SER A   284
REMARK 465     GLN A   285
REMARK 465     PRO A   286
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   NZ B LYS A    22     O    HOH A  2033              2.19
REMARK 500   NZ B LYS A    39     O    HOH A  2049              1.50
REMARK 500   O  A PRO A    40     O    HOH A  2056              1.48
REMARK 500   O  B PRO A    40     O    HOH A  2057              1.67
REMARK 500   NE B ARG A    43     O    HOH A  2064              2.20
REMARK 500   CZ B ARG A    43     O    HOH A  2064              1.99
REMARK 500   NH2B ARG A    43     O    HOH A  2064              1.33
REMARK 500   OG B SER A    57     O    HOH A  2079              1.99
REMARK 500   NH1B ARG A    64     O    HOH A  2009              1.07
REMARK 500   OD1B ASN A    89     O1   EDO A  1297              2.04
REMARK 500   OE2B GLU A   112     O    HOH A  2112              1.49
REMARK 500   OE2B GLU A   181     O    HOH A  2140              1.68
REMARK 500   CZ B ARG A   183     O    HOH A  2144              1.47
REMARK 500   NH2B ARG A   183     O    HOH A  2144              0.81
REMARK 500   CD A GLU A   186     O    HOH A  2145              2.01
REMARK 500   OE2A GLU A   186     O    HOH A  2145              1.99
REMARK 500   OE2B GLU A   186     O    HOH A  2097              1.62
REMARK 500   CD B ARG A   202     O    HOH A  2154              1.77
REMARK 500   CD B GLU A   229     O    HOH A  2167              1.59
REMARK 500   OE1B GLU A   229     O    HOH A  2167              1.45
REMARK 500   OE2B GLU A   229     O    HOH A  2169              1.87
REMARK 500   OE2B GLU A   229     O    HOH A  2167              2.01
REMARK 500   NZ B LYS A   233     O    HOH A  2174              1.89
REMARK 500   NH1A ARG A   237     O    HOH A  2176              2.06
REMARK 500   OE1B GLU A   244     O    HOH A  2181              1.99
REMARK 500   O2   EDO A  1297     O    HOH A  2096              2.11
REMARK 500  CL     CL A  1299     O    HOH A  2176              1.76
REMARK 500   O    HOH A  2017     O    HOH A  2018              1.66
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   NZ B LYS A    29     CD A GLU A   191     4465     1.91
REMARK 500   NZ B LYS A    29     OE1A GLU A   191     4465     1.96
REMARK 500   NE2B GLN A   108     OE2B GLU A   266     3555     1.73
REMARK 500   NH1B ARG A   225     O    HOH A  2056     3545     1.64
REMARK 500   O    HOH A  2022     O    HOH A  2141     4465     1.92
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ARG A   7   NE  -  CZ  -  NH1 ANGL. DEV. =   3.2 DEGREES
REMARK 500    ARG A  87   NE  -  CZ  -  NH1 ANGL. DEV. =   3.3 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ALA A  41      -17.56     96.84
REMARK 500    SER A 126     -119.06     58.58
REMARK 500    ASN A 161       64.65     27.10
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    ASN A 161        23.6      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EPE A1284
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PE8 A1285
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A1286
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A1287
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PGE A1288
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PGE A1289
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1290
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1291
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1292
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1293
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1294
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1295
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1296
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1297
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  CL A1298
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  CL A1299
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 5AOA   RELATED DB: PDB
REMARK 900  THE STRUCTURE OF A NOVEL THERMOPHILIC ESTERASE FROM
REMARK 900  THE PLANCTOMYCETES SPECIES, THERMOGUTTA TERRIFONTIS, EST2-
REMARK 900  PROPIONATE BOUND
REMARK 900 RELATED ID: 5AOB   RELATED DB: PDB
REMARK 900  THE STRUCTURE OF A NOVEL THERMOPHILIC ESTERASE FROM
REMARK 900  THE PLANCTOMYCETES SPECIES, THERMOGUTTA TERRIFONTIS, EST2-
REMARK 900  BUTYRATE BOUND
REMARK 900 RELATED ID: 5AOC   RELATED DB: PDB
REMARK 900  THE STRUCTURE OF A NOVEL THERMOPHILIC ESTERASE FROM
REMARK 900  THE PLANCTOMYCETES SPECIES, THERMOGUTTA TERRIFONTIS, EST2-
REMARK 900  VALERATE BOUND
DBREF  5AO9 A    1   286  PDB    5AO9     5AO9             1    286
SEQRES   1 A  286  ALA GLU VAL GLY ARG LEU ARG TYR PRO PRO GLU MET PRO
SEQRES   2 A  286  GLY ALA GLU VAL LYS VAL TYR LYS LYS VAL ASP ASN VAL
SEQRES   3 A  286  ASP LEU LYS LEU TYR ILE TYR LYS PRO ALA ASP TRP LYS
SEQRES   4 A  286  PRO ALA ASP ARG ARG SER ALA ILE VAL PHE PHE PHE GLY
SEQRES   5 A  286  GLY GLY TRP GLN SER GLY SER PRO ALA GLN PHE ARG PRO
SEQRES   6 A  286  GLN CYS GLU TYR PHE ALA GLY ARG GLY MET VAL ALA MET
SEQRES   7 A  286  ALA ALA ASP TYR ARG VAL GLY SER ARG HIS ASN VAL LYS
SEQRES   8 A  286  VAL ALA ASP CYS VAL ALA ASP ALA LYS SER ALA ILE ARG
SEQRES   9 A  286  TRP VAL ARG GLN HIS ALA ALA GLU LEU GLY VAL ASP PRO
SEQRES  10 A  286  GLN LYS ILE VAL ALA SER GLY GLY SER ALA GLY GLY HIS
SEQRES  11 A  286  LEU ALA ALA CYS THR VAL MET VAL PRO ASP LEU GLU ALA
SEQRES  12 A  286  PRO GLU GLU ASP HIS THR ILE SER SER GLN ALA ASN ALA
SEQRES  13 A  286  ALA ILE LEU PHE ASN PRO VAL LEU ILE LEU SER ARG GLU
SEQRES  14 A  286  GLY LEU LYS ASP HIS VAL PRO ARG GLN ASP TRP GLU GLU
SEQRES  15 A  286  ARG LEU ARG GLU ARG LEU GLY THR GLU PRO LYS ALA VAL
SEQRES  16 A  286  SER PRO TYR HIS HIS ILE ARG ALA GLY LEU PRO PRO MET
SEQRES  17 A  286  ILE ILE PHE HIS GLY THR ALA ASP ASN THR VAL PRO PHE
SEQRES  18 A  286  GLU THR ILE ARG LEU PHE ALA GLU ALA MET LYS LYS ALA
SEQRES  19 A  286  GLY ASN ARG CYS GLU LEU VAL PRO PHE GLU GLY ALA ALA
SEQRES  20 A  286  HIS GLY PHE PHE ASN PHE GLY ARG GLY ASP ASN LEU ALA
SEQRES  21 A  286  TYR GLN LYS THR LEU GLU LEU ALA ASP GLU PHE LEU VAL
SEQRES  22 A  286  GLU ILE GLY PHE LEU ALA PRO LYS GLY GLU SER GLN PRO
HET    EPE  A1284      15
HET    PE8  A1285      25
HET    PEG  A1286       7
HET    PEG  A1287       7
HET    PGE  A1288      10
HET    PGE  A1289      10
HET    EDO  A1290       4
HET    EDO  A1291       4
HET    EDO  A1292       4
HET    EDO  A1293       4
HET    EDO  A1294       4
HET    EDO  A1295       4
HET    EDO  A1296       4
HET    EDO  A1297       4
HET     CL  A1298       1
HET     CL  A1299       1
HETNAM     EDO 1,2-ETHANEDIOL
HETNAM     EPE 4-(2-HYDROXYETHYL)-1-PIPERAZINE
HETNAM   2 EPE  ETHANESULFONIC ACID
HETNAM     PE8 3,6,9,12,15,18,21-HEPTAOXATRICOSANE-1,23-DIOL
HETNAM      CL CHLORIDE ION
HETNAM     PGE TRIETHYLENE GLYCOL
HETNAM     PEG DI(HYDROXYETHYL)ETHER
HETSYN     EDO ETHYLENE GLYCOL
HETSYN     EPE HEPES
FORMUL   2  EDO    8(C2 H6 O2)
FORMUL   3  EPE    C8 H18 N2 O4 S
FORMUL   4  PE8    C16 H34 O9
FORMUL   5   CL    2(CL 1-)
FORMUL   6  PGE    2(C6 H14 O4)
FORMUL   7  PEG    2(C4 H10 O3)
FORMUL   8  HOH   *208(H2 O)
HELIX    1   1 GLU A    2  LEU A    6  5                                   5
HELIX    2   2 SER A   59  GLN A   62  5                                   4
HELIX    3   3 PHE A   63  ARG A   73  1                                  11
HELIX    4   4 VAL A   84  ASN A   89  1                                   6
HELIX    5   5 LYS A   91  HIS A  109  1                                  19
HELIX    6   6 HIS A  109  GLY A  114  1                                   6
HELIX    7   7 SER A  126  VAL A  138  1                                  13
HELIX    8   8 ARG A  177  GLY A  189  1                                  13
HELIX    9   9 GLU A  191  SER A  196  5                                   6
HELIX   10  10 SER A  196  ILE A  201  5                                   6
HELIX   11  11 PRO A  220  ALA A  234  1                                  15
HELIX   12  12 ARG A  255  ASP A  257  5                                   3
HELIX   13  13 ASN A  258  ILE A  275  1                                  18
SHEET    1  AA 8 GLU A  16  VAL A  23  0
SHEET    2  AA 8 VAL A  26  TYR A  33 -1  O  VAL A  26   N  VAL A  23
SHEET    3  AA 8 VAL A  76  ALA A  80 -1  O  ALA A  77   N  TYR A  33
SHEET    4  AA 8 ARG A  44  PHE A  50  1  O  SER A  45   N  VAL A  76
SHEET    5  AA 8 VAL A 115  GLY A 125  1  N  ASP A 116   O  ARG A  44
SHEET    6  AA 8 ALA A 156  PHE A 160  1  O  ALA A 156   N  ALA A 122
SHEET    7  AA 8 MET A 208  GLY A 213  1  O  ILE A 209   N  LEU A 159
SHEET    8  AA 8 CYS A 238  PHE A 243  1  O  GLU A 239   N  ILE A 210
CISPEP   1 TYR A    8    PRO A    9          0        -5.74
SITE     1 AC1 11 GLY A  52  GLN A  62  GLY A 125  SER A 126
SITE     2 AC1 11 ASN A 161  THR A 218  HIS A 248  GLY A 249
SITE     3 AC1 11 PHE A 251  ASN A 252  HOH A2075
SITE     1 AC2  9 MET A 137  VAL A 138  ASP A 140  GLN A 153
SITE     2 AC2  9 GLY A 204  LEU A 205  PGE A1288  HOH A2201
SITE     3 AC2  9 HOH A2202
SITE     1 AC3  4 GLN A  56  ARG A 187  HOH A2077  HOH A2098
SITE     1 AC4  1 VAL A 163
SITE     1 AC5  7 ARG A 107  GLN A 118  GLN A 153  ASN A 155
SITE     2 AC5  7 PE8 A1285  HOH A2115  HOH A2204
SITE     1 AC6  5 TYR A  20  LYS A  22  GLU A 145  GLU A 270
SITE     2 AC6  5 HOH A2028
SITE     1 AC7  2 LYS A  34  PRO A  35
SITE     1 AC8  1 ARG A 168
SITE     1 AC9  2 GLN A 262  GLU A 266
SITE     1 BC1  1 TYR A 198
SITE     1 BC2  4 PRO A  65  GLU A  68  TYR A 261  HOH A2206
SITE     1 BC3  3 VAL A   3  LEU A   6  PHE A 253
SITE     1 BC4  2 GLY A 276  PHE A 277
SITE     1 BC5  4 GLY A  85  SER A  86  ARG A  87  HOH A2096
SITE     1 BC6  3 ASN A 217  THR A 218  HIS A 248
SITE     1 BC7  2 ARG A 237  HOH A2176
CRYST1   61.640   71.070   75.840  90.00  90.00  90.00 P 21 21 21    4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.016223  0.000000  0.000000        0.00000
SCALE2      0.000000  0.014071  0.000000        0.00000
SCALE3      0.000000  0.000000  0.013186        0.00000
TER    2429      GLU A 283
MASTER      440    0   16   13    8    0   22    6 2744    1  106   22
END