longtext: 5bkm-pdb

content
HEADER    HYDROLASE                               20-MAR-21   5BKM
TITLE     CRYSTAL STRUCTURE OF HIP1 (RV2224C) MUTANT - S228DHA (DEHYDROALANINE)
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: CARBOXYLESTERASE A;
COMPND   3 CHAIN: A;
COMPND   4 SYNONYM: HIP1;
COMPND   5 EC: 3.1.1.-;
COMPND   6 ENGINEERED: YES;
COMPND   7 MUTATION: YES;
COMPND   8 OTHER_DETAILS: N-TERMINAL 6XHIS TAG WITH LINKER AND THROMBIN
COMPND   9 RECOGNITION CLEAVAGE SITE. ALSO RESIDUE SERINE 228 HAS BEEN MODIFIED
COMPND  10 TO DEHYDROALANINE (DHA)
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: MYCOBACTERIUM TUBERCULOSIS;
SOURCE   3 ORGANISM_TAXID: 83332;
SOURCE   4 STRAIN: ATCC 25618 / H37RV;
SOURCE   5 ATCC: 25618;
SOURCE   6 GENE: CAEA, RV2224C, MTCY427.05C;
SOURCE   7 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   8 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   9 EXPRESSION_SYSTEM_VARIANT: STAR;
SOURCE  10 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  11 EXPRESSION_SYSTEM_PLASMID: PET28A
KEYWDS    SERINE PROTEASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    J.L.NAFFIN-OLIVOS,A.DAAB,N.E.GOLDFARB,M.H.DORAN,J.BAIKOVITZ,D.LIU,
AUTHOR   2 S.SUN,A.WHITE,B.M.DUNN,J.RENGARAJAN,G.A.PETSKO,D.RINGE
REVDAT   1   23-MAR-22 5BKM    0
JRNL        AUTH   J.L.NAFFIN-OLIVOS,A.DAAB,N.GOLDFARB,M.H.DORAN,J.BAIKOVITZ,
JRNL        AUTH 2 D.LIU,S.SUN,A.WHITE,B.M.DUNN,J.RENGARAJAN,G.A.PETSKO,D.RINGE
JRNL        TITL   CRYSTAL STRUCTURE OF HIP1 (RV2224C) MUTANT - S228DHA
JRNL        TITL 2 (DEHYDROALANINE)
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    2.70 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.9_1692
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.70
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 45.23
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.360
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.9
REMARK   3   NUMBER OF REFLECTIONS             : 22484
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.203
REMARK   3   R VALUE            (WORKING SET) : 0.200
REMARK   3   FREE R VALUE                     : 0.242
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 8.840
REMARK   3   FREE R VALUE TEST SET COUNT      : 1987
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 45.2260 -  6.5083    0.99     1547   159  0.1827 0.2212
REMARK   3     2  6.5083 -  5.1682    1.00     1508   145  0.2024 0.2369
REMARK   3     3  5.1682 -  4.5156    1.00     1478   142  0.1662 0.2028
REMARK   3     4  4.5156 -  4.1030    1.00     1475   137  0.1688 0.2186
REMARK   3     5  4.1030 -  3.8091    1.00     1473   142  0.1824 0.2207
REMARK   3     6  3.8091 -  3.5846    1.00     1439   140  0.1853 0.2496
REMARK   3     7  3.5846 -  3.4052    1.00     1479   147  0.2159 0.2681
REMARK   3     8  3.4052 -  3.2570    1.00     1431   140  0.2254 0.2566
REMARK   3     9  3.2570 -  3.1316    1.00     1459   138  0.2364 0.2953
REMARK   3    10  3.1316 -  3.0236    1.00     1455   139  0.2388 0.2765
REMARK   3    11  3.0236 -  2.9291    1.00     1453   141  0.2483 0.2909
REMARK   3    12  2.9291 -  2.8453    1.00     1413   130  0.2653 0.2933
REMARK   3    13  2.8453 -  2.7705    1.00     1464   145  0.2668 0.3207
REMARK   3    14  2.7705 -  2.7030    1.00     1423   142  0.2635 0.2991
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.320
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 24.300
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 60.71
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 60.66
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.009           3586
REMARK   3   ANGLE     :  1.133           4882
REMARK   3   CHIRALITY :  0.043            536
REMARK   3   PLANARITY :  0.006            660
REMARK   3   DIHEDRAL  : 13.767           1313
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 5BKM COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 26-MAR-21.
REMARK 100 THE DEPOSITION ID IS D_1000255327.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 21-NOV-17
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.6
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : APS
REMARK 200  BEAMLINE                       : 19-ID
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9794
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315R
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 22515
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.700
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.9
REMARK 200  DATA REDUNDANCY                : 13.20
REMARK 200  R MERGE                    (I) : 0.30300
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 2.6000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.70
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.80
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.7
REMARK 200  DATA REDUNDANCY IN SHELL       : 10.60
REMARK 200  R MERGE FOR SHELL          (I) : 0.83400
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: SOLVE
REMARK 200 STARTING MODEL: 5UNO
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 67.54
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.79
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 30% (W/V) PEG3350, 0.1 M SODIUM
REMARK 280  ACETATE PH 5.6, 0.2 M (NH4)2SO4, 10% (V/V) GLYCEROL, VAPOR
REMARK 280  DIFFUSION, HANGING DROP, TEMPERATURE 298K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 31 2 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z+1/3
REMARK 290       3555   -X+Y,-X,Z+2/3
REMARK 290       4555   Y,X,-Z
REMARK 290       5555   X-Y,-Y,-Z+2/3
REMARK 290       6555   -X,-X+Y,-Z+1/3
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       42.49233
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       84.98467
REMARK 290   SMTRY1   4 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   4  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   5  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       84.98467
REMARK 290   SMTRY1   6 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000       42.49233
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A    29
REMARK 465     GLY A    30
REMARK 465     SER A    31
REMARK 465     SER A    32
REMARK 465     HIS A    33
REMARK 465     HIS A    34
REMARK 465     HIS A    35
REMARK 465     HIS A    36
REMARK 465     HIS A    37
REMARK 465     HIS A    38
REMARK 465     SER A    39
REMARK 465     SER A    40
REMARK 465     GLY A    41
REMARK 465     LEU A    42
REMARK 465     VAL A    43
REMARK 465     PRO A    44
REMARK 465     ARG A    45
REMARK 465     GLY A    46
REMARK 465     SER A    47
REMARK 465     HIS A    48
REMARK 465     MET A    49
REMARK 465     SER A    57
REMARK 465     SER A    58
REMARK 465     ASN A    59
REMARK 465     PRO A    60
REMARK 465     GLN A    61
REMARK 465     VAL A    62
REMARK 465     LYS A    63
REMARK 465     ILE A    64
REMARK 480
REMARK 480 ZERO OCCUPANCY ATOM
REMARK 480 THE FOLLOWING RESIDUES HAVE ATOMS MODELED WITH ZERO
REMARK 480 OCCUPANCY. THE LOCATION AND PROPERTIES OF THESE ATOMS
REMARK 480 MAY NOT BE RELIABLE. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 480 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 480   M RES C SSEQI ATOMS
REMARK 480     GLU A   51   CB   CG   CD   OE1  OE2
REMARK 480     ARG A   56   CB   CG   CD   NE   CZ   NH1  NH2
REMARK 480     LYS A   72   CG   CD   CE   NZ
REMARK 480     ARG A   81   CD   NE   CZ   NH1  NH2
REMARK 480     ASP A   83   CB   CG   OD1  OD2
REMARK 480     LYS A  128   CG   CD   CE   NZ
REMARK 480     GLU A  132   CG   CD   OE1  OE2
REMARK 480     LYS A  296   CD   CE   NZ
REMARK 480     ARG A  321   CG   CD   NE   CZ   NH1  NH2
REMARK 480     LYS A  323   CG   CD   CE   NZ
REMARK 480     LYS A  404   CG   CD   CE   NZ
REMARK 480     LYS A  423   CG   CD   CE   NZ
REMARK 480     LYS A  519   CG   CD   CE   NZ
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    PRO A  54      172.18    -59.97
REMARK 500    ALA A 145     -108.48     50.70
REMARK 500    CYS A 153      -56.50   -128.27
REMARK 500    PRO A 314        0.90    -69.46
REMARK 500    ARG A 318       71.94   -114.87
REMARK 500    THR A 491      165.01     78.13
REMARK 500    VAL A 492      -17.86   -158.55
REMARK 500
REMARK 500 REMARK: NULL
DBREF  5BKM A   50   520  UNP    P9WHR3   CAEA_MYCTU      50    520
SEQADV 5BKM MET A   29  UNP  P9WHR3              INITIATING METHIONINE
SEQADV 5BKM GLY A   30  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM SER A   31  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM SER A   32  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM HIS A   33  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM HIS A   34  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM HIS A   35  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM HIS A   36  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM HIS A   37  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM HIS A   38  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM SER A   39  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM SER A   40  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM GLY A   41  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM LEU A   42  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM VAL A   43  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM PRO A   44  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM ARG A   45  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM GLY A   46  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM SER A   47  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM HIS A   48  UNP  P9WHR3              EXPRESSION TAG
SEQADV 5BKM MET A   49  UNP  P9WHR3              EXPRESSION TAG
SEQRES   1 A  492  MET GLY SER SER HIS HIS HIS HIS HIS HIS SER SER GLY
SEQRES   2 A  492  LEU VAL PRO ARG GLY SER HIS MET VAL GLU TRP THR PRO
SEQRES   3 A  492  CYS ARG SER SER ASN PRO GLN VAL LYS ILE PRO GLY GLY
SEQRES   4 A  492  ALA LEU CYS GLY LYS LEU ALA VAL PRO VAL ASP TYR ASP
SEQRES   5 A  492  ARG PRO ASP GLY ASP VAL ALA ALA LEU ALA LEU ILE ARG
SEQRES   6 A  492  PHE PRO ALA THR GLY ASP LYS ILE GLY SER LEU VAL ILE
SEQRES   7 A  492  ASN PRO GLY GLY PRO GLY GLU SER GLY ILE GLU ALA ALA
SEQRES   8 A  492  LEU GLY VAL PHE GLN THR LEU PRO LYS ARG VAL HIS GLU
SEQRES   9 A  492  ARG PHE ASP LEU VAL GLY PHE ASP PRO ARG GLY VAL ALA
SEQRES  10 A  492  SER SER ARG PRO ALA ILE TRP CYS ASN SER ASP ALA ASP
SEQRES  11 A  492  ASN ASP ARG LEU ARG ALA GLU PRO GLN VAL ASP TYR SER
SEQRES  12 A  492  ARG GLU GLY VAL ALA HIS ILE GLU ASN GLU THR LYS GLN
SEQRES  13 A  492  PHE VAL GLY ARG CYS VAL ASP LYS MET GLY LYS ASN PHE
SEQRES  14 A  492  LEU ALA HIS VAL GLY THR VAL ASN VAL ALA LYS ASP LEU
SEQRES  15 A  492  ASP ALA ILE ARG ALA ALA LEU GLY ASP ASP LYS LEU THR
SEQRES  16 A  492  TYR LEU GLY TYR DHA TYR GLY THR ARG ILE GLY SER ALA
SEQRES  17 A  492  TYR ALA GLU GLU PHE PRO GLN ARG VAL ARG ALA MET ILE
SEQRES  18 A  492  LEU ASP GLY ALA VAL ASP PRO ASN ALA ASP PRO ILE GLU
SEQRES  19 A  492  ALA GLU LEU ARG GLN ALA LYS GLY PHE GLN ASP ALA PHE
SEQRES  20 A  492  ASN ASN TYR ALA ALA ASP CYS ALA LYS ASN ALA GLY CYS
SEQRES  21 A  492  PRO LEU GLY ALA ASP PRO ALA LYS ALA VAL GLU VAL TYR
SEQRES  22 A  492  HIS SER LEU VAL ASP PRO LEU VAL ASP PRO ASP ASN PRO
SEQRES  23 A  492  ARG ILE SER ARG PRO ALA ARG THR LYS ASP PRO ARG GLY
SEQRES  24 A  492  LEU SER TYR SER ASP ALA ILE VAL GLY THR ILE MET ALA
SEQRES  25 A  492  LEU TYR SER PRO ASN LEU TRP GLN HIS LEU THR ASP GLY
SEQRES  26 A  492  LEU SER GLU LEU VAL ASP ASN ARG GLY ASP THR LEU LEU
SEQRES  27 A  492  ALA LEU ALA ASP MET TYR MET ARG ARG ASP SER HIS GLY
SEQRES  28 A  492  ARG TYR ASN ASN SER GLY ASP ALA ARG VAL ALA ILE ASN
SEQRES  29 A  492  CYS VAL ASP GLN PRO PRO VAL THR ASP ARG ASP LYS VAL
SEQRES  30 A  492  ILE ASP GLU ASP ARG ARG ALA ARG GLU ILE ALA PRO PHE
SEQRES  31 A  492  MET SER TYR GLY LYS PHE THR GLY ASP ALA PRO LEU GLY
SEQRES  32 A  492  THR CYS ALA PHE TRP PRO VAL PRO PRO THR SER GLN PRO
SEQRES  33 A  492  HIS ALA VAL SER ALA PRO GLY LEU VAL PRO THR VAL VAL
SEQRES  34 A  492  VAL SER THR THR HIS ASP PRO ALA THR PRO TYR LYS ALA
SEQRES  35 A  492  GLY VAL ASP LEU ALA ASN GLN LEU ARG GLY SER LEU LEU
SEQRES  36 A  492  THR PHE ASP GLY THR GLN HIS THR VAL VAL PHE GLN GLY
SEQRES  37 A  492  ASP SER CYS ILE ASP GLU TYR VAL THR ALA TYR LEU ILE
SEQRES  38 A  492  GLY GLY THR THR PRO PRO SER GLY ALA LYS CYS
MODRES 5BKM DHA A  228  SER  MODIFIED RESIDUE
HET    DHA  A 228       5
HETNAM     DHA 2-AMINO-ACRYLIC ACID
HETSYN     DHA 2,3-DIDEHYDROALANINE
FORMUL   1  DHA    C3 H5 N O2
HELIX    1 AA1 SER A  114  LEU A  126  1                                  13
HELIX    2 AA2 PRO A  127  ARG A  133  1                                   7
HELIX    3 AA3 SER A  155  GLU A  165  1                                  11
HELIX    4 AA4 SER A  171  GLY A  194  1                                  24
HELIX    5 AA5 GLY A  194  HIS A  200  1                                   7
HELIX    6 AA6 GLY A  202  LEU A  217  1                                  16
HELIX    7 AA7 TYR A  229  PHE A  241  1                                  13
HELIX    8 AA8 ASP A  259  LYS A  284  1                                  26
HELIX    9 AA9 ASP A  293  ALA A  295  5                                   3
HELIX   10 AB1 LYS A  296  ASP A  306  1                                  11
HELIX   11 AB2 PRO A  307  VAL A  309  5                                   3
HELIX   12 AB3 SER A  329  LEU A  341  1                                  13
HELIX   13 AB4 TYR A  342  ASN A  345  5                                   4
HELIX   14 AB5 LEU A  346  ASP A  359  1                                  14
HELIX   15 AB6 GLY A  362  ARG A  374  1                                  13
HELIX   16 AB7 ASN A  383  GLN A  396  1                                  14
HELIX   17 AB8 ASP A  401  ALA A  416  1                                  16
HELIX   18 AB9 PRO A  417  SER A  420  5                                   4
HELIX   19 AC1 GLY A  431  TRP A  436  5                                   6
HELIX   20 AC2 PRO A  467  ARG A  479  1                                  13
HELIX   21 AC3 ASP A  497  GLY A  511  1                                  15
SHEET    1 AA110 THR A  53  PRO A  54  0
SHEET    2 AA110 LEU A  69  PRO A  76 -1  O  CYS A  70   N  THR A  53
SHEET    3 AA110 VAL A  86  PHE A  94 -1  O  LEU A  91   N  GLY A  71
SHEET    4 AA110 PHE A 134  PHE A 139 -1  O  LEU A 136   N  PHE A  94
SHEET    5 AA110 GLY A 102  ASN A 107  1  N  LEU A 104   O  VAL A 137
SHEET    6 AA110 LEU A 222  TYR A 227  1  O  THR A 223   N  VAL A 105
SHEET    7 AA110 VAL A 245  ASP A 251  1  O  ILE A 249   N  TYR A 224
SHEET    8 AA110 VAL A 456  THR A 460  1  O  VAL A 456   N  MET A 248
SHEET    9 AA110 SER A 481  PHE A 485  1  O  SER A 481   N  VAL A 457
SHEET   10 AA110 LYS A 519  CYS A 520  1  O  CYS A 520   N  THR A 484
SSBOND   1 CYS A   55    CYS A   70                          1555   1555  2.05
SSBOND   2 CYS A  153    CYS A  189                          1555   1555  2.05
SSBOND   3 CYS A  282    CYS A  288                          1555   1555  2.07
SSBOND   4 CYS A  393    CYS A  433                          1555   1555  2.03
SSBOND   5 CYS A  499    CYS A  520                          1555   1555  2.06
LINK         C   TYR A 227                 N   DHA A 228     1555   1555  1.33
LINK         C   DHA A 228                 N   TYR A 229     1555   1555  1.33
CISPEP   1 GLY A  110    PRO A  111          0         0.24
CISPEP   2 ARG A  148    PRO A  149          0        -3.14
CRYST1  104.444  104.444  127.477  90.00  90.00 120.00 P 31 2 1      6
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.009575  0.005528  0.000000        0.00000
SCALE2      0.000000  0.011056  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007845        0.00000
TER    3501      CYS A 520
MASTER      289    0    1   21   10    0    0    6 3500    1   17   38
END