longtext: 5e4j-pdb

content
HEADER    HYDROLASE                               06-OCT-15   5E4J
TITLE     ACETYCHOLINESTERASE METHYLENE BLUE NO PEG
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ACETYLCHOLINESTERASE;
COMPND   3 CHAIN: A;
COMPND   4 SYNONYM: ACHE;
COMPND   5 EC: 3.1.1.7
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: TORPEDO CALIFORNICA;
SOURCE   3 ORGANISM_COMMON: PACIFIC ELECTRIC RAY;
SOURCE   4 ORGANISM_TAXID: 7787
KEYWDS    INHIBITOR, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    O.DYM
REVDAT   1   30-MAR-16 5E4J    0
JRNL        AUTH   O.DYM,W.SONG,C.FELDER,E.ROTH,V.SHNYROV,Y.ASHANI,Y.XU,
JRNL        AUTH 2 R.P.JOOSTEN,L.WEINER,J.L.SUSSMAN,I.SILMAN
JRNL        TITL   THE IMPACT OF CRYSTALLIZATION CONDITIONS ON STRUCTURE-BASED
JRNL        TITL 2 DRUG DESIGN: A CASE STUDY ON THE METHYLENE
JRNL        TITL 3 BLUE/ACETYLCHOLINESTERASE COMPLEX.
JRNL        REF    PROTEIN SCI.                               2016
JRNL        REFN                   ESSN 1469-896X
JRNL        PMID   26990888
JRNL        DOI    10.1002/PRO.2923
REMARK   2
REMARK   2 RESOLUTION.    2.54 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0073
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.54
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 48.15
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.7
REMARK   3   NUMBER OF REFLECTIONS             : 31097
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.193
REMARK   3   R VALUE            (WORKING SET) : 0.191
REMARK   3   FREE R VALUE                     : 0.229
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.100
REMARK   3   FREE R VALUE TEST SET COUNT      : 1656
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.54
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.61
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 2241
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 100.0
REMARK   3   BIN R VALUE           (WORKING SET) : 0.3060
REMARK   3   BIN FREE R VALUE SET COUNT          : 124
REMARK   3   BIN FREE R VALUE                    : 0.3650
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4174
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 157
REMARK   3   SOLVENT ATOMS            : 15
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 59.39
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -0.67000
REMARK   3    B22 (A**2) : -0.67000
REMARK   3    B33 (A**2) : 2.18000
REMARK   3    B12 (A**2) : -0.34000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.280
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.222
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.188
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 19.863
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.952
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.931
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  4456 ; 0.007 ; 0.019
REMARK   3   BOND LENGTHS OTHERS               (A):  4059 ; 0.001 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  6072 ; 1.391 ; 1.977
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  9314 ; 0.778 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   531 ; 6.060 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   196 ;32.710 ;23.827
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   674 ;15.703 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    22 ;17.151 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   664 ; 0.164 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  4956 ; 0.005 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  1056 ; 0.001 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  2127 ; 1.262 ; 3.256
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  2126 ; 1.257 ; 3.255
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  2657 ; 2.133 ; 4.884
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  2658 ; 2.135 ; 4.885
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  2329 ; 1.537 ; 3.547
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  2290 ; 1.499 ; 3.486
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  3356 ; 2.455 ; 5.172
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  4953 ; 4.436 ;26.388
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  4953 ; 4.434 ;26.392
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 1
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A     4        A   535
REMARK   3    ORIGIN FOR THE GROUP (A):  52.5861 -28.2562  33.2983
REMARK   3    T TENSOR
REMARK   3      T11:   0.0921 T22:   0.2315
REMARK   3      T33:   0.0072 T12:  -0.1046
REMARK   3      T13:   0.0160 T23:  -0.0169
REMARK   3    L TENSOR
REMARK   3      L11:   1.4683 L22:   2.1497
REMARK   3      L33:   3.0133 L12:  -0.0876
REMARK   3      L13:  -0.4228 L23:   0.6247
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1937 S12:  -0.0531 S13:  -0.0587
REMARK   3      S21:  -0.1185 S22:   0.1274 S23:   0.0709
REMARK   3      S31:   0.2129 S32:  -0.3271 S33:   0.0664
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS
REMARK   4
REMARK   4 5E4J COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 06-OCT-15.
REMARK 100 THE DEPOSITION ID IS D_1000214251.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 20-SEP-12
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID23-1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9763
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315R
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 38533
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.540
REMARK 200  RESOLUTION RANGE LOW       (A) : 48.150
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.8
REMARK 200  DATA REDUNDANCY                : 6.900
REMARK 200  R MERGE                    (I) : 0.14600
REMARK 200  R SYM                      (I) : 0.14000
REMARK 200   FOR THE DATA SET  : 3.5000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.54
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.68
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.8
REMARK 200  DATA REDUNDANCY IN SHELL       : 7.10
REMARK 200  R MERGE FOR SHELL          (I) : 0.56000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 1.200
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 69.83
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 4.08
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 60% AS/360 MM NA,K PHOSPHATE/100 MM NA
REMARK 280  PHOSPHATE, PH 6.5, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE
REMARK 280  279K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 31 2 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z+1/3
REMARK 290       3555   -X+Y,-X,Z+2/3
REMARK 290       4555   Y,X,-Z
REMARK 290       5555   X-Y,-Y,-Z+2/3
REMARK 290       6555   -X,-X+Y,-Z+1/3
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       45.67667
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       91.35333
REMARK 290   SMTRY1   4 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   4  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   5  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       91.35333
REMARK 290   SMTRY1   6 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000       45.67667
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 3540 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 21340 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -104.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     LYS A  52    CE   NZ
REMARK 470     GLU A 140    CG   CD   OE1  OE2
REMARK 470     ARG A 250    CZ   NH1  NH2
REMARK 470     GLU A 260    CG   CD   OE1  OE2
REMARK 470     GLU A 261    CG   CD   OE1  OE2
REMARK 470     GLU A 268    OE1  OE2
REMARK 470     LYS A 270    CG   CD   CE   NZ
REMARK 470     GLU A 344    OE1  OE2
REMARK 470     GLU A 350    OE1  OE2
REMARK 470     GLU A 434    CG   CD   OE1  OE2
REMARK 470     LYS A 454    CD   CE   NZ
REMARK 470     GLU A 455    CG   CD   OE1  OE2
REMARK 470     GLU A 461    CD   OE1  OE2
REMARK 470     HIS A 486    CG   ND1  CD2  CE1  NE2
REMARK 470     SER A 487    OG
REMARK 470     GLN A 488    CG   CD   OE1  NE2
REMARK 470     GLU A 489    CG   CD   OE1  OE2
REMARK 470     LYS A 498    CG   CD   CE   NZ
REMARK 470     GLU A 508    CD   OE1  OE2
REMARK 470     LYS A 511    CD   CE   NZ
REMARK 470     ARG A 515    CZ   NH1  NH2
REMARK 470     GLN A 526    OE1  NE2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    PHE A  45       -7.82     77.67
REMARK 500    PHE A 120        2.98     59.09
REMARK 500    LEU A 158       76.58   -104.83
REMARK 500    ALA A 164       65.51   -150.18
REMARK 500    SER A 200     -118.24     68.04
REMARK 500    GLU A 299      -74.22    -82.64
REMARK 500    THR A 317     -157.57   -153.89
REMARK 500    ASP A 380      -70.00    -87.68
REMARK 500    ASP A 381      -45.84     80.51
REMARK 500    VAL A 400      -60.81   -133.37
REMARK 500    HIS A 486      -96.52    -88.26
REMARK 500    SER A 487      113.72     57.33
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue DME A 608
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 609
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 610
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 611
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 612
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 613
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 614
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 615
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 616
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 617
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 618
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Mono-Saccharide NAG A 601 bound
REMARK 800  to ASN A 59
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Mono-Saccharide NAG A 602 bound
REMARK 800  to ASN A 416
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NAG A
REMARK 800  603 through MAN A 607 bound to ASN A 457
DBREF  5E4J A    4   535  UNP    P04058   ACES_TORCA      25    556
SEQRES   1 A  532  SER GLU LEU LEU VAL ASN THR LYS SER GLY LYS VAL MET
SEQRES   2 A  532  GLY THR ARG VAL PRO VAL LEU SER SER HIS ILE SER ALA
SEQRES   3 A  532  PHE LEU GLY ILE PRO PHE ALA GLU PRO PRO VAL GLY ASN
SEQRES   4 A  532  MET ARG PHE ARG ARG PRO GLU PRO LYS LYS PRO TRP SER
SEQRES   5 A  532  GLY VAL TRP ASN ALA SER THR TYR PRO ASN ASN CYS GLN
SEQRES   6 A  532  GLN TYR VAL ASP GLU GLN PHE PRO GLY PHE SER GLY SER
SEQRES   7 A  532  GLU MET TRP ASN PRO ASN ARG GLU MET SER GLU ASP CYS
SEQRES   8 A  532  LEU TYR LEU ASN ILE TRP VAL PRO SER PRO ARG PRO LYS
SEQRES   9 A  532  SER THR THR VAL MET VAL TRP ILE TYR GLY GLY GLY PHE
SEQRES  10 A  532  TYR SER GLY SER SER THR LEU ASP VAL TYR ASN GLY LYS
SEQRES  11 A  532  TYR LEU ALA TYR THR GLU GLU VAL VAL LEU VAL SER LEU
SEQRES  12 A  532  SER TYR ARG VAL GLY ALA PHE GLY PHE LEU ALA LEU HIS
SEQRES  13 A  532  GLY SER GLN GLU ALA PRO GLY ASN VAL GLY LEU LEU ASP
SEQRES  14 A  532  GLN ARG MET ALA LEU GLN TRP VAL HIS ASP ASN ILE GLN
SEQRES  15 A  532  PHE PHE GLY GLY ASP PRO LYS THR VAL THR ILE PHE GLY
SEQRES  16 A  532  GLU SER ALA GLY GLY ALA SER VAL GLY MET HIS ILE LEU
SEQRES  17 A  532  SER PRO GLY SER ARG ASP LEU PHE ARG ARG ALA ILE LEU
SEQRES  18 A  532  GLN SER GLY SER PRO ASN CYS PRO TRP ALA SER VAL SER
SEQRES  19 A  532  VAL ALA GLU GLY ARG ARG ARG ALA VAL GLU LEU GLY ARG
SEQRES  20 A  532  ASN LEU ASN CYS ASN LEU ASN SER ASP GLU GLU LEU ILE
SEQRES  21 A  532  HIS CYS LEU ARG GLU LYS LYS PRO GLN GLU LEU ILE ASP
SEQRES  22 A  532  VAL GLU TRP ASN VAL LEU PRO PHE ASP SER ILE PHE ARG
SEQRES  23 A  532  PHE SER PHE VAL PRO VAL ILE ASP GLY GLU PHE PHE PRO
SEQRES  24 A  532  THR SER LEU GLU SER MET LEU ASN SER GLY ASN PHE LYS
SEQRES  25 A  532  LYS THR GLN ILE LEU LEU GLY VAL ASN LYS ASP GLU GLY
SEQRES  26 A  532  SER PHE PHE LEU LEU TYR GLY ALA PRO GLY PHE SER LYS
SEQRES  27 A  532  ASP SER GLU SER LYS ILE SER ARG GLU ASP PHE MET SER
SEQRES  28 A  532  GLY VAL LYS LEU SER VAL PRO HIS ALA ASN ASP LEU GLY
SEQRES  29 A  532  LEU ASP ALA VAL THR LEU GLN TYR THR ASP TRP MET ASP
SEQRES  30 A  532  ASP ASN ASN GLY ILE LYS ASN ARG ASP GLY LEU ASP ASP
SEQRES  31 A  532  ILE VAL GLY ASP HIS ASN VAL ILE CYS PRO LEU MET HIS
SEQRES  32 A  532  PHE VAL ASN LYS TYR THR LYS PHE GLY ASN GLY THR TYR
SEQRES  33 A  532  LEU TYR PHE PHE ASN HIS ARG ALA SER ASN LEU VAL TRP
SEQRES  34 A  532  PRO GLU TRP MET GLY VAL ILE HIS GLY TYR GLU ILE GLU
SEQRES  35 A  532  PHE VAL PHE GLY LEU PRO LEU VAL LYS GLU LEU ASN TYR
SEQRES  36 A  532  THR ALA GLU GLU GLU ALA LEU SER ARG ARG ILE MET HIS
SEQRES  37 A  532  TYR TRP ALA THR PHE ALA LYS THR GLY ASN PRO ASN GLU
SEQRES  38 A  532  PRO HIS SER GLN GLU SER LYS TRP PRO LEU PHE THR THR
SEQRES  39 A  532  LYS GLU GLN LYS PHE ILE ASP LEU ASN THR GLU PRO MET
SEQRES  40 A  532  LYS VAL HIS GLN ARG LEU ARG VAL GLN MET CYS VAL PHE
SEQRES  41 A  532  TRP ASN GLN PHE LEU PRO LYS LEU LEU ASN ALA THR
HET    NAG  A 601      14
HET    NAG  A 602      14
HET    NAG  A 603      14
HET    NAG  A 604      14
HET    BMA  A 605      11
HET    MAN  A 606      11
HET    MAN  A 607      11
HET    DME  A 608      18
HET    SO4  A 609       5
HET    SO4  A 610       5
HET    SO4  A 611       5
HET    SO4  A 612       5
HET    SO4  A 613       5
HET    SO4  A 614       5
HET    SO4  A 615       5
HET    SO4  A 616       5
HET    SO4  A 617       5
HET    SO4  A 618       5
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     BMA BETA-D-MANNOSE
HETNAM     MAN ALPHA-D-MANNOSE
HETNAM     DME DECAMETHONIUM ION
HETNAM     SO4 SULFATE ION
FORMUL   2  NAG    4(C8 H15 N O6)
FORMUL   4  BMA    C6 H12 O6
FORMUL   4  MAN    2(C6 H12 O6)
FORMUL   5  DME    C16 H38 N2 2+
FORMUL   6  SO4    10(O4 S 2-)
FORMUL  16  HOH   *15(H2 O)
HELIX    1 AA1 VAL A   40  ARG A   44  5                                   5
HELIX    2 AA2 PHE A   78  MET A   83  1                                   6
HELIX    3 AA3 LEU A  127  ASN A  131  5                                   5
HELIX    4 AA4 GLY A  132  GLU A  140  1                                   9
HELIX    5 AA5 VAL A  150  LEU A  156  1                                   7
HELIX    6 AA6 ASN A  167  ILE A  184  1                                  18
HELIX    7 AA7 GLN A  185  PHE A  187  5                                   3
HELIX    8 AA8 SER A  200  SER A  212  1                                  13
HELIX    9 AA9 SER A  215  PHE A  219  5                                   5
HELIX   10 AB1 SER A  237  LEU A  252  1                                  16
HELIX   11 AB2 SER A  258  LYS A  269  1                                  12
HELIX   12 AB3 LYS A  270  GLU A  278  1                                   9
HELIX   13 AB4 TRP A  279  LEU A  282  5                                   4
HELIX   14 AB5 SER A  304  SER A  311  1                                   8
HELIX   15 AB6 GLY A  328  ALA A  336  1                                   9
HELIX   16 AB7 SER A  348  VAL A  360  1                                  13
HELIX   17 AB8 ASN A  364  THR A  376  1                                  13
HELIX   18 AB9 ASN A  383  VAL A  400  1                                  18
HELIX   19 AC1 VAL A  400  GLY A  415  1                                  16
HELIX   20 AC2 PRO A  433  GLY A  437  5                                   5
HELIX   21 AC3 GLU A  443  PHE A  448  1                                   6
HELIX   22 AC4 GLY A  449  ASN A  457  5                                   9
HELIX   23 AC5 THR A  459  GLY A  480  1                                  22
HELIX   24 AC6 ARG A  517  GLN A  526  1                                  10
HELIX   25 AC7 GLN A  526  THR A  535  1                                  10
SHEET    1 AA1 3 LEU A   7  THR A  10  0
SHEET    2 AA1 3 GLY A  13  MET A  16 -1  O  VAL A  15   N  VAL A   8
SHEET    3 AA1 3 VAL A  57  ASN A  59  1  O  TRP A  58   N  MET A  16
SHEET    1 AA211 THR A  18  PRO A  21  0
SHEET    2 AA211 HIS A  26  PRO A  34 -1  O  ILE A  27   N  VAL A  20
SHEET    3 AA211 TYR A  96  VAL A 101 -1  O  VAL A 101   N  SER A  28
SHEET    4 AA211 VAL A 142  SER A 145 -1  O  LEU A 143   N  TRP A 100
SHEET    5 AA211 THR A 109  ILE A 115  1  N  TRP A 114   O  VAL A 144
SHEET    6 AA211 GLY A 189  GLU A 199  1  O  THR A 195   N  VAL A 113
SHEET    7 AA211 ARG A 221  GLN A 225  1  O  GLN A 225   N  GLY A 198
SHEET    8 AA211 GLN A 318  ASN A 324  1  O  LEU A 320   N  LEU A 224
SHEET    9 AA211 GLY A 417  PHE A 423  1  O  GLY A 417   N  ILE A 319
SHEET   10 AA211 LYS A 501  LEU A 505  1  O  ILE A 503   N  PHE A 422
SHEET   11 AA211 VAL A 512  GLN A 514 -1  O  HIS A 513   N  PHE A 502
SSBOND   1 CYS A   67    CYS A   94                          1555   1555  2.06
SSBOND   2 CYS A  254    CYS A  265                          1555   1555  2.07
SSBOND   3 CYS A  402    CYS A  521                          1555   1555  2.08
LINK         ND2 ASN A  59                 C1  NAG A 601     1555   1555  1.46
LINK         ND2 ASN A 416                 C1  NAG A 602     1555   1555  1.46
LINK         ND2 ASN A 457                 C1  NAG A 603     1555   1555  1.47
LINK         O4  NAG A 603                 C1  NAG A 604     1555   1555  1.45
LINK         O4  NAG A 604                 C1  BMA A 605     1555   1555  1.45
LINK         O3  BMA A 605                 C1  MAN A 606     1555   1555  1.45
LINK         O2  MAN A 606                 C1  MAN A 607     1555   1555  1.44
CISPEP   1 SER A  103    PRO A  104          0        -1.83
SITE     1 AC1  8 TYR A  70  TRP A  84  GLY A 118  TYR A 121
SITE     2 AC1  8 GLU A 199  TRP A 279  TYR A 334  GLY A 441
SITE     1 AC2  3 HIS A 406  ASN A 525  GLN A 526
SITE     1 AC3  5 ASN A 424  HIS A 425  ASN A 506  THR A 507
SITE     2 AC3  5 GLU A 508
SITE     1 AC4  5 PRO A  21  VAL A  22  LEU A  23  LYS A 133
SITE     2 AC4  5 TYR A 134
SITE     1 AC5  6 ARG A 289  SER A 359  VAL A 360  PRO A 361
SITE     2 AC5  6 HIS A 362  HIS A 398
SITE     1 AC6  4 SER A 237  ALA A 239  GLU A 240  ARG A 243
SITE     1 AC7  2 HIS A 513  GLN A 514
SITE     1 AC8  3 LYS A 346  ASN A 382  GLY A 384
SITE     1 AC9  2 SER A 258  ASP A 259
SITE     1 AD1  4 GLN A  68  SER A  91  GLU A  92  PRO A 271
SITE     1 AD2  5 PRO A 213  PRO A 302  ASP A 342  SER A 343
SITE     2 AD2  5 GLU A 344
SITE     1 AD3  2 ASN A  59  SER A  61
SITE     1 AD4  2 ASN A 416  MAN A 607
SITE     1 AD5  5 LYS A 413  PHE A 414  GLY A 415  ASN A 457
SITE     2 AD5  5 NAG A 602
CRYST1  111.210  111.210  137.030  90.00  90.00 120.00 P 31 2 1      6
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008992  0.005192  0.000000        0.00000
SCALE2      0.000000  0.010383  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007298        0.00000
TER    4175      THR A 535
MASTER      382    0   18   25   14    0   20    6 4346    1  166   41
END