longtext: 5ikk-pdb

content
HEADER    TRANSCRIPTION                           03-MAR-16   5IKK
TITLE     STRUCTURE OF THE HISTONE DEACETYLASE CLR3
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: HISTONE DEACETYLASE CLR3;
COMPND   3 CHAIN: A;
COMPND   4 SYNONYM: CRYPTIC LOCI REGULATOR 3;
COMPND   5 EC: 3.5.1.98;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: SCHIZOSACCHAROMYCES POMBE 972H-;
SOURCE   3 ORGANISM_COMMON: FISSION YEAST;
SOURCE   4 ORGANISM_TAXID: 284812;
SOURCE   5 GENE: CLR3, SPBC800.03;
SOURCE   6 EXPRESSION_SYSTEM: SPODOPTERA FRUGIPERDA;
SOURCE   7 EXPRESSION_SYSTEM_COMMON: FALL ARMYWORM;
SOURCE   8 EXPRESSION_SYSTEM_TAXID: 7108;
SOURCE   9 EXPRESSION_SYSTEM_CELL_LINE: SF9;
SOURCE  10 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  11 EXPRESSION_SYSTEM_VECTOR: PACEBAC1 MULTIBAC;
SOURCE  12 EXPRESSION_SYSTEM_PLASMID: P728
KEYWDS    HDAC DOMAIN, ALPHA/BETA HYDROLASE DOMAIN, DIMER, ALPHA/BETA SANDWICH,
KEYWDS   2 HYDROLASE, TRANSCRIPTION
EXPDTA    X-RAY DIFFRACTION
AUTHOR    C.BRUGGER,T.SCHALCH
REVDAT   2   04-MAY-16 5IKK    1       JRNL
REVDAT   1   20-APR-16 5IKK    0
JRNL        AUTH   G.JOB,C.BRUGGER,T.XU,B.R.LOWE,Y.PFISTER,C.QU,S.SHANKER,
JRNL        AUTH 2 J.I.BANOS SANZ,J.F.PARTRIDGE,T.SCHALCH
JRNL        TITL   SHREC SILENCES HETEROCHROMATIN VIA DISTINCT REMODELING AND
JRNL        TITL 2 DEACETYLATION MODULES.
JRNL        REF    MOL.CELL                      V.  62   207 2016
JRNL        REFN                   ISSN 1097-2765
JRNL        PMID   27105116
JRNL        DOI    10.1016/J.MOLCEL.2016.03.016
REMARK   2
REMARK   2 RESOLUTION.    2.40 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.8.4_1496
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.40
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 46.82
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.360
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.0
REMARK   3   NUMBER OF REFLECTIONS             : 31826
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.189
REMARK   3   R VALUE            (WORKING SET) : 0.187
REMARK   3   FREE R VALUE                     : 0.228
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.790
REMARK   3   FREE R VALUE TEST SET COUNT      : 1524
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 46.8236 -  5.3352    1.00     2944   148  0.1716 0.1949
REMARK   3     2  5.3352 -  4.2356    1.00     2831   141  0.1417 0.1865
REMARK   3     3  4.2356 -  3.7004    1.00     2807   141  0.1576 0.1941
REMARK   3     4  3.7004 -  3.3622    1.00     2750   138  0.1669 0.2114
REMARK   3     5  3.3622 -  3.1213    1.00     2780   139  0.1924 0.2554
REMARK   3     6  3.1213 -  2.9373    1.00     2780   139  0.2021 0.2165
REMARK   3     7  2.9373 -  2.7902    1.00     2736   138  0.2140 0.2606
REMARK   3     8  2.7902 -  2.6687    1.00     2749   137  0.2193 0.2796
REMARK   3     9  2.6687 -  2.5660    1.00     2745   138  0.2382 0.2814
REMARK   3    10  2.5660 -  2.4775    1.00     2740   138  0.2561 0.2767
REMARK   3    11  2.4775 -  2.4000    0.90     2440   127  0.2755 0.3360
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : NULL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.270
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 22.170
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 52.30
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.006           4859
REMARK   3   ANGLE     :  0.646           6551
REMARK   3   CHIRALITY :  0.028            719
REMARK   3   PLANARITY :  0.003            847
REMARK   3   DIHEDRAL  : 11.392           1757
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 2
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: (CHAIN 'A' AND RESID 54 THROUGH 420)
REMARK   3    ORIGIN FOR THE GROUP (A):  13.0244  65.1010 -21.9546
REMARK   3    T TENSOR
REMARK   3      T11:   0.1810 T22:   0.2709
REMARK   3      T33:   0.1736 T12:  -0.1439
REMARK   3      T13:  -0.1133 T23:  -0.0193
REMARK   3    L TENSOR
REMARK   3      L11:   2.4613 L22:   1.3436
REMARK   3      L33:   2.5269 L12:  -0.4452
REMARK   3      L13:  -0.3521 L23:   0.0725
REMARK   3    S TENSOR
REMARK   3      S11:   0.2747 S12:  -0.6336 S13:  -0.0272
REMARK   3      S21:   0.2356 S22:   0.0565 S23:  -0.2170
REMARK   3      S31:  -0.0872 S32:   0.3230 S33:  -0.0841
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: (CHAIN 'A' AND RESID 421 THROUGH 680)
REMARK   3    ORIGIN FOR THE GROUP (A):  14.3293  17.9836 -49.4468
REMARK   3    T TENSOR
REMARK   3      T11:   0.3043 T22:   0.2368
REMARK   3      T33:   0.2962 T12:   0.0228
REMARK   3      T13:   0.0793 T23:   0.0802
REMARK   3    L TENSOR
REMARK   3      L11:   2.2424 L22:   1.4612
REMARK   3      L33:   2.2357 L12:  -0.3123
REMARK   3      L13:  -0.9046 L23:   0.2532
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0265 S12:   0.2166 S13:   0.0703
REMARK   3      S21:  -0.3122 S22:  -0.0550 S23:  -0.3497
REMARK   3      S31:   0.1811 S32:   0.3509 S33:   0.1000
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 5IKK COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 03-MAR-16.
REMARK 100 THE DEPOSITION ID IS D_1000218969.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 16-MAY-14
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 7.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID29
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97924
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS3 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : AIMLESS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 31830
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.400
REMARK 200  RESOLUTION RANGE LOW       (A) : 46.815
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.9
REMARK 200  DATA REDUNDANCY                : 6.600
REMARK 200  R MERGE                    (I) : 0.07200
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 14.8600
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.40
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.46
REMARK 200  COMPLETENESS FOR SHELL     (%) : 87.7
REMARK 200  DATA REDUNDANCY IN SHELL       : 5.90
REMARK 200  R MERGE FOR SHELL          (I) : 0.60000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.500
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: 2VQW (HDAC4)
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 55.04
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.74
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 20% ETHYLENEGLYCOL 10% PEG 8000 0.1M
REMARK 280  MOPS/HEPES 0.04M SODIUM NITRATE 0.04M SODIUM MONOPHOSPHATE 0.04M
REMARK 280  AMMONIUM SULFATE, PH 7.0, VAPOR DIFFUSION, HANGING DROP,
REMARK 280  TEMPERATURE 291K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 2 2 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z+1/2
REMARK 290       3555   -X,Y,-Z+1/2
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z
REMARK 290       6555   -X+1/2,-Y+1/2,Z+1/2
REMARK 290       7555   -X+1/2,Y+1/2,-Z+1/2
REMARK 290       8555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       71.31650
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       71.31650
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       30.24350
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       93.62950
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       30.24350
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       93.62950
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       71.31650
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       30.24350
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       93.62950
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       71.31650
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       30.24350
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000       93.62950
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 10930 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 50970 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -127.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   2  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000      -71.31650
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     THR A    31
REMARK 465     ASN A    32
REMARK 465     ASN A    33
REMARK 465     LYS A    34
REMARK 465     PRO A    35
REMARK 465     MET A    36
REMARK 465     SER A    37
REMARK 465     GLY A    38
REMARK 465     SER A    39
REMARK 465     GLU A    40
REMARK 465     ASN A    41
REMARK 465     THR A    42
REMARK 465     LEU A    43
REMARK 465     ASN A    44
REMARK 465     ASN A    45
REMARK 465     GLU A    46
REMARK 465     SER A    47
REMARK 465     HIS A    48
REMARK 465     GLU A    49
REMARK 465     MET A    50
REMARK 465     SER A    51
REMARK 465     GLN A    52
REMARK 465     ILE A    53
REMARK 465     VAL A    74
REMARK 465     ASP A    75
REMARK 465     ASP A    76
REMARK 465     ARG A   131
REMARK 465     VAL A   132
REMARK 465     THR A   133
REMARK 465     ASN A   134
REMARK 465     THR A   135
REMARK 465     GLU A   136
REMARK 465     GLU A   197
REMARK 465     PRO A   198
REMARK 465     HIS A   199
REMARK 465     LYS A   200
REMARK 465     PRO A   201
REMARK 465     GLY A   202
REMARK 465     GLY A   203
REMARK 465     PHE A   204
REMARK 465     CYS A   205
REMARK 465     SER A   462
REMARK 465     VAL A   463
REMARK 465     SER A   464
REMARK 465     ASN A   465
REMARK 465     VAL A   466
REMARK 465     PHE A   467
REMARK 465     VAL A   538
REMARK 465     THR A   539
REMARK 465     ASP A   540
REMARK 465     GLY A   541
REMARK 465     GLU A   542
REMARK 465     ASN A   543
REMARK 465     ALA A   544
REMARK 465     PRO A   545
REMARK 465     VAL A   546
REMARK 465     LYS A   681
REMARK 465     ALA A   682
REMARK 465     ARG A   683
REMARK 465     PRO A   684
REMARK 465     THR A   685
REMARK 465     SER A   686
REMARK 465     GLN A   687
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    VAL A 188       60.99   -114.04
REMARK 500    VAL A 307      -62.76   -123.75
REMARK 500    GLU A 364     -130.77   -125.26
REMARK 500    ASN A 469       10.08     59.87
REMARK 500    TYR A 563      -53.27   -129.27
REMARK 500    GLU A 618      -63.57   -123.10
REMARK 500
REMARK 500 REMARK: NULL
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              ZN A 703  ZN
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HIS A  68   NE2
REMARK 620 2 HIS A  77   NE2 103.2
REMARK 620 3 CYS A 162   SG   96.4 135.4
REMARK 620 N                    1     2
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              ZN A 701  ZN
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HIS A 195   NE2
REMARK 620 2 ASP A 234   OD1 139.7
REMARK 620 3 ASP A 234   OD2 113.9  63.2
REMARK 620 4 HIS A 236   ND1  88.3  95.7 156.8
REMARK 620 5 ASP A 327   OD2 103.9 114.8  82.5  99.4
REMARK 620 N                    1     2     3     4
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                               K A 704   K
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASP A 232   O
REMARK 620 2 ASP A 232   OD1  71.4
REMARK 620 3 ASP A 232   OD2  91.6  39.6
REMARK 620 4 ASP A 234   O    95.5  88.7 121.0
REMARK 620 5 HIS A 236   O   161.7  94.0  83.3  72.6
REMARK 620 6 SER A 255   OG   86.9 125.4  94.6 144.1 110.9
REMARK 620 7 LEU A 256   O    72.2 129.7 163.8  61.7 111.9  85.5
REMARK 620 N                    1     2     3     4     5     6
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                               K A 702   K
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 PHE A 245   O
REMARK 620 2 ASP A 248   O    71.6
REMARK 620 3 VAL A 251   O   117.7  82.7
REMARK 620 4 ARG A 284   O   155.7 122.0  85.4
REMARK 620 5 HOH A 822   O    85.3  96.4 154.9  73.7
REMARK 620 6 HOH A 856   O    73.2 144.7 116.4  90.3  78.2
REMARK 620 N                    1     2     3     4     5
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              MG A 712  MG
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 PRO A 427   O
REMARK 620 2 ASP A 429   O    98.9
REMARK 620 3 VAL A 432   O   101.3  97.9
REMARK 620 4 HOH A 906   O    90.5 170.2  82.5
REMARK 620 5 HOH A 934   O    85.5  94.7 164.6  83.6
REMARK 620 N                    1     2     3     4
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue ZN A 701
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue K A 702
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue ZN A 703
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue K A 704
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NO3 A 705
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NO3 A 706
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NO3 A 707
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 708
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 709
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 710
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 711
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue MG A 712
DBREF  5IKK A   31   687  UNP    P56523   CLR3_SCHPO      31    687
SEQRES   1 A  657  THR ASN ASN LYS PRO MET SER GLY SER GLU ASN THR LEU
SEQRES   2 A  657  ASN ASN GLU SER HIS GLU MET SER GLN ILE LEU LYS LYS
SEQRES   3 A  657  SER GLY LEU CYS TYR ASP PRO ARG MET ARG PHE HIS ALA
SEQRES   4 A  657  THR LEU SER GLU VAL ASP ASP HIS PRO GLU ASP PRO ARG
SEQRES   5 A  657  ARG VAL LEU ARG VAL PHE GLU ALA ILE LYS LYS ALA GLY
SEQRES   6 A  657  TYR VAL SER ASN VAL PRO SER PRO SER ASP VAL PHE LEU
SEQRES   7 A  657  ARG ILE PRO ALA ARG GLU ALA THR LEU GLU GLU LEU LEU
SEQRES   8 A  657  GLN VAL HIS SER GLN GLU MET TYR ASP ARG VAL THR ASN
SEQRES   9 A  657  THR GLU LYS MET SER HIS GLU ASP LEU ALA ASN LEU GLU
SEQRES  10 A  657  LYS ILE SER ASP SER LEU TYR TYR ASN ASN GLU SER ALA
SEQRES  11 A  657  PHE CYS ALA ARG LEU ALA CYS GLY SER ALA ILE GLU THR
SEQRES  12 A  657  CYS THR ALA VAL VAL THR GLY GLN VAL LYS ASN ALA PHE
SEQRES  13 A  657  ALA VAL VAL ARG PRO PRO GLY HIS HIS ALA GLU PRO HIS
SEQRES  14 A  657  LYS PRO GLY GLY PHE CYS LEU PHE ASN ASN VAL SER VAL
SEQRES  15 A  657  THR ALA ARG SER MET LEU GLN ARG PHE PRO ASP LYS ILE
SEQRES  16 A  657  LYS ARG VAL LEU ILE VAL ASP TRP ASP ILE HIS HIS GLY
SEQRES  17 A  657  ASN GLY THR GLN MET ALA PHE TYR ASP ASP PRO ASN VAL
SEQRES  18 A  657  LEU TYR VAL SER LEU HIS ARG TYR GLU ASN GLY ARG PHE
SEQRES  19 A  657  TYR PRO GLY THR ASN TYR GLY CYS ALA GLU ASN CYS GLY
SEQRES  20 A  657  GLU GLY PRO GLY LEU GLY ARG THR VAL ASN ILE PRO TRP
SEQRES  21 A  657  SER CYS ALA GLY MET GLY ASP GLY ASP TYR ILE TYR ALA
SEQRES  22 A  657  PHE GLN ARG VAL VAL MET PRO VAL ALA TYR GLU PHE ASP
SEQRES  23 A  657  PRO ASP LEU VAL ILE VAL SER CYS GLY PHE ASP ALA ALA
SEQRES  24 A  657  ALA GLY ASP HIS ILE GLY GLN PHE LEU LEU THR PRO ALA
SEQRES  25 A  657  ALA TYR ALA HIS MET THR GLN MET LEU MET GLY LEU ALA
SEQRES  26 A  657  ASP GLY LYS VAL PHE ILE SER LEU GLU GLY GLY TYR ASN
SEQRES  27 A  657  LEU ASP SER ILE SER THR SER ALA LEU ALA VAL ALA GLN
SEQRES  28 A  657  SER LEU LEU GLY ILE PRO PRO GLY ARG LEU HIS THR THR
SEQRES  29 A  657  TYR ALA CYS PRO GLN ALA VAL ALA THR ILE ASN HIS VAL
SEQRES  30 A  657  THR LYS ILE GLN SER GLN TYR TRP ARG CYS MET ARG PRO
SEQRES  31 A  657  LYS HIS PHE ASP ALA ASN PRO LYS ASP ALA HIS VAL ASP
SEQRES  32 A  657  ARG LEU HIS ASP VAL ILE ARG THR TYR GLN ALA LYS LYS
SEQRES  33 A  657  LEU PHE GLU ASP TRP LYS ILE THR ASN MET PRO ILE LEU
SEQRES  34 A  657  ARG ASP SER VAL SER ASN VAL PHE ASN ASN GLN VAL LEU
SEQRES  35 A  657  CYS SER SER ASN PHE PHE GLN LYS ASP ASN LEU LEU VAL
SEQRES  36 A  657  ILE VAL HIS GLU SER PRO ARG VAL LEU GLY ASN GLY THR
SEQRES  37 A  657  SER GLU THR ASN VAL LEU ASN LEU ASN ASP SER LEU LEU
SEQRES  38 A  657  VAL ASP PRO VAL SER LEU TYR VAL GLU TRP ALA MET GLN
SEQRES  39 A  657  GLN ASP TRP GLY LEU ILE ASP ILE ASN ILE PRO GLU VAL
SEQRES  40 A  657  VAL THR ASP GLY GLU ASN ALA PRO VAL ASP ILE LEU SER
SEQRES  41 A  657  GLU VAL LYS GLU LEU CYS LEU TYR VAL TRP ASP ASN TYR
SEQRES  42 A  657  VAL GLU LEU SER ILE SER LYS ASN ILE PHE PHE ILE GLY
SEQRES  43 A  657  GLY GLY LYS ALA VAL HIS GLY LEU VAL ASN LEU ALA SER
SEQRES  44 A  657  SER ARG ASN VAL SER ASP ARG VAL LYS CYS MET VAL ASN
SEQRES  45 A  657  PHE LEU GLY THR GLU PRO LEU VAL GLY LEU LYS THR ALA
SEQRES  46 A  657  SER GLU GLU ASP LEU PRO THR TRP TYR TYR ARG HIS SER
SEQRES  47 A  657  LEU VAL PHE VAL SER SER SER ASN GLU CYS TRP LYS LYS
SEQRES  48 A  657  ALA LYS ARG ALA LYS ARG ARG TYR GLY ARG LEU MET GLN
SEQRES  49 A  657  SER GLU HIS THR GLU THR SER ASP MET MET GLU GLN HIS
SEQRES  50 A  657  TYR ARG ALA VAL THR GLN TYR LEU LEU HIS LEU LEU GLN
SEQRES  51 A  657  LYS ALA ARG PRO THR SER GLN
HET     ZN  A 701       1
HET      K  A 702       1
HET     ZN  A 703       1
HET      K  A 704       1
HET    NO3  A 705       4
HET    NO3  A 706       4
HET    NO3  A 707       4
HET    EDO  A 708      10
HET    EDO  A 709      10
HET    EDO  A 710      10
HET    SO4  A 711       5
HET     MG  A 712       1
HETNAM      ZN ZINC ION
HETNAM       K POTASSIUM ION
HETNAM     NO3 NITRATE ION
HETNAM     EDO 1,2-ETHANEDIOL
HETNAM     SO4 SULFATE ION
HETNAM      MG MAGNESIUM ION
HETSYN     EDO ETHYLENE GLYCOL
FORMUL   2   ZN    2(ZN 2+)
FORMUL   3    K    2(K 1+)
FORMUL   6  NO3    3(N O3 1-)
FORMUL   9  EDO    3(C2 H6 O2)
FORMUL  12  SO4    O4 S 2-
FORMUL  13   MG    MG 2+
FORMUL  14  HOH   *213(H2 O)
HELIX    1 AA1 ARG A   64  ALA A   69  1                                   6
HELIX    2 AA2 PRO A   81  ALA A   94  1                                  14
HELIX    3 AA3 THR A  116  ASP A  130  1                                  15
HELIX    4 AA4 SER A  139  LEU A  143  5                                   5
HELIX    5 AA5 ASN A  145  SER A  152  1                                   8
HELIX    6 AA6 SER A  159  THR A  179  1                                  21
HELIX    7 AA7 ASN A  208  PHE A  221  1                                  14
HELIX    8 AA8 GLY A  238  TYR A  246  1                                   9
HELIX    9 AA9 THR A  268  CYS A  272  5                                   5
HELIX   10 AB1 GLU A  278  LEU A  282  5                                   5
HELIX   11 AB2 GLY A  296  VAL A  307  1                                  12
HELIX   12 AB3 VAL A  307  ASP A  316  1                                  10
HELIX   13 AB4 THR A  340  MET A  352  1                                  13
HELIX   14 AB5 GLY A  353  GLY A  357  5                                   5
HELIX   15 AB6 ASN A  368  LEU A  384  1                                  17
HELIX   16 AB7 CYS A  397  SER A  412  1                                  16
HELIX   17 AB8 ARG A  434  LYS A  452  1                                  19
HELIX   18 AB9 ASN A  476  LYS A  480  5                                   5
HELIX   19 AC1 PRO A  514  GLN A  525  1                                  12
HELIX   20 AC2 ILE A  548  TYR A  563  1                                  16
HELIX   21 AC3 LYS A  579  ARG A  591  1                                  13
HELIX   22 AC4 GLU A  618  HIS A  627  1                                  10
HELIX   23 AC5 ASN A  636  LYS A  640  5                                   5
HELIX   24 AC6 LYS A  646  GLY A  650  5                                   5
HELIX   25 AC7 GLU A  659  HIS A  667  1                                   9
HELIX   26 AC8 HIS A  667  LEU A  679  1                                  13
SHEET    1 AA1 8 PHE A 107  ILE A 110  0
SHEET    2 AA1 8 SER A  57  TYR A  61  1  N  SER A  57   O  LEU A 108
SHEET    3 AA1 8 ASN A 184  VAL A 188  1  O  ASN A 184   N  GLY A  58
SHEET    4 AA1 8 VAL A 359  LEU A 363  1  O  ILE A 361   N  ALA A 185
SHEET    5 AA1 8 LEU A 319  CYS A 324  1  N  VAL A 322   O  PHE A 360
SHEET    6 AA1 8 VAL A 228  ASP A 232  1  N  LEU A 229   O  LEU A 319
SHEET    7 AA1 8 VAL A 251  ARG A 258  1  O  LEU A 252   N  ILE A 230
SHEET    8 AA1 8 THR A 285  TRP A 290  1  O  ILE A 288   N  SER A 255
SHEET    1 AA2 8 THR A 454  ASN A 455  0
SHEET    2 AA2 8 VAL A 471  CYS A 473 -1  O  CYS A 473   N  THR A 454
SHEET    3 AA2 8 GLY A 528  ASN A 533 -1  O  ASP A 531   N  LEU A 472
SHEET    4 AA2 8 ASN A 482  HIS A 488  1  N  ASN A 482   O  GLY A 528
SHEET    5 AA2 8 ASN A 571  GLY A 577  1  O  ILE A 575   N  VAL A 487
SHEET    6 AA2 8 VAL A 597  PHE A 603  1  O  VAL A 601   N  PHE A 574
SHEET    7 AA2 8 SER A 628  VAL A 632  1  O  LEU A 629   N  ASN A 602
SHEET    8 AA2 8 LEU A 652  GLN A 654  1  O  MET A 653   N  VAL A 630
SHEET    1 AA3 2 ARG A 492  LEU A 494  0
SHEET    2 AA3 2 LEU A 510  VAL A 512 -1  O  VAL A 512   N  ARG A 492
LINK         NE2 HIS A  68                ZN    ZN A 703     1555   1555  2.03
LINK         NE2 HIS A  77                ZN    ZN A 703     1555   1555  2.04
LINK         SG  CYS A 162                ZN    ZN A 703     1555   1555  2.34
LINK         NE2 HIS A 195                ZN    ZN A 701     1555   1555  1.99
LINK         O   ASP A 232                 K     K A 704     1555   1555  2.82
LINK         OD1 ASP A 232                 K     K A 704     1555   1555  2.82
LINK         OD2 ASP A 232                 K     K A 704     1555   1555  3.44
LINK         O   ASP A 234                 K     K A 704     1555   1555  2.77
LINK         OD1 ASP A 234                ZN    ZN A 701     1555   1555  2.06
LINK         OD2 ASP A 234                ZN    ZN A 701     1555   1555  2.09
LINK         O   HIS A 236                 K     K A 704     1555   1555  2.89
LINK         ND1 HIS A 236                ZN    ZN A 701     1555   1555  2.04
LINK         O   PHE A 245                 K     K A 702     1555   1555  2.66
LINK         O   ASP A 248                 K     K A 702     1555   1555  2.84
LINK         O   VAL A 251                 K     K A 702     1555   1555  2.61
LINK         OG  SER A 255                 K     K A 704     1555   1555  2.79
LINK         O   LEU A 256                 K     K A 704     1555   1555  3.04
LINK         O   ARG A 284                 K     K A 702     1555   1555  2.83
LINK         OD2 ASP A 327                ZN    ZN A 701     1555   1555  1.99
LINK         O   PRO A 427                MG    MG A 712     1555   1555  2.26
LINK         O   ASP A 429                MG    MG A 712     1555   1555  2.13
LINK         O   VAL A 432                MG    MG A 712     1555   1555  2.24
LINK         K     K A 702                 O   HOH A 822     1555   1555  2.81
LINK         K     K A 702                 O   HOH A 856     1555   1555  2.83
LINK        MG    MG A 712                 O   HOH A 906     1555   1555  2.08
LINK        MG    MG A 712                 O   HOH A 934     1555   1555  2.08
CISPEP   1 TYR A  265    PRO A  266          0         4.22
CISPEP   2 ARG A  419    PRO A  420          0        -1.06
SITE     1 AC1  4 HIS A 195  ASP A 234  HIS A 236  ASP A 327
SITE     1 AC2  6 PHE A 245  ASP A 248  VAL A 251  ARG A 284
SITE     2 AC2  6 HOH A 822  HOH A 856
SITE     1 AC3  3 HIS A  68  HIS A  77  CYS A 162
SITE     1 AC4  5 ASP A 232  ASP A 234  HIS A 236  SER A 255
SITE     2 AC4  5 LEU A 256
SITE     1 AC5  6 ARG A  64  MET A  65  ARG A  66  LYS A 452
SITE     2 AC5  6 ASN A 476  HOH A 808
SITE     1 AC6  6 TYR A 313  ASP A 316  HIS A 346  ARG A 390
SITE     2 AC6  6 LEU A 391  HOH A 967
SITE     1 AC7  4 SER A 412  GLN A 413  TYR A 414  ARG A 419
SITE     1 AC8  3 TYR A 302  ARG A 416  ARG A 419
SITE     1 AC9  8 ARG A 440  HIS A 488  ASP A 513  VAL A 515
SITE     2 AC9  8 SER A 516  ASP A 531  HOH A 925  HOH A 954
SITE     1 AD1  4 ASP A  80  GLY A 366  TYR A 367  LEU A 369
SITE     1 AD2  6 TYR A 129  GLU A 617  PRO A 621  LYS A 646
SITE     2 AD2  6 ARG A 647  ARG A 648
SITE     1 AD3  5 PRO A 427  ASP A 429  VAL A 432  HOH A 906
SITE     2 AD3  5 HOH A 934
CRYST1   60.487  187.259  142.633  90.00  90.00  90.00 C 2 2 21      8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.016532  0.000000  0.000000        0.00000
SCALE2      0.000000  0.005340  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007011        0.00000
TER    9290      GLN A 680
MASTER      450    0   12   26   18    0   19    6 4952    1   82   51
END