longtext: 5txe-pdb

content
HEADER    HYDROLASE                               16-NOV-16   5TXE
TITLE     ATXE2 ISOPEPTIDASE - S527A VARIANT WITH ASTEXIN3-DC4 BOUND
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ATXE2;
COMPND   3 CHAIN: A, B;
COMPND   4 ENGINEERED: YES;
COMPND   5 MUTATION: YES;
COMPND   6 MOL_ID: 2;
COMPND   7 MOLECULE: ASTEXIN3-DC4;
COMPND   8 CHAIN: C, D;
COMPND   9 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: ASTICCACAULIS EXCENTRICUS (STRAIN ATCC 15261 /
SOURCE   3 DSM 4724 / VKM B-1370 / CB 48);
SOURCE   4 ORGANISM_TAXID: 573065;
SOURCE   5 STRAIN: ATCC 15261 / DSM 4724 / VKM B-1370 / CB 48;
SOURCE   6 GENE: ASTEX_2444;
SOURCE   7 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   8 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   9 MOL_ID: 2;
SOURCE  10 ORGANISM_SCIENTIFIC: ASTICCACAULIS EXCENTRICUS (STRAIN ATCC 15261 /
SOURCE  11 DSM 4724 / VKM B-1370 / CB 48);
SOURCE  12 ORGANISM_TAXID: 573065;
SOURCE  13 STRAIN: ATCC 15261 / DSM 4724 / VKM B-1370 / CB 48;
SOURCE  14 GENE: ASTEX_2447;
SOURCE  15 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE  16 EXPRESSION_SYSTEM_TAXID: 562
KEYWDS    ISOPEPTIDASE, LASSO PEPTIDE, NATURAL PRODUCT, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    J.R.CHEKAN,S.K.NAIR
REVDAT   1   21-DEC-16 5TXE    0
JRNL        AUTH   J.R.CHEKAN,J.D.KOOS,C.ZONG,M.O.MAKSIMOV,J.A.LINK,S.K.NAIR
JRNL        TITL   STRUCTURE OF THE LASSO PEPTIDE ISOPEPTIDASE IDENTIFIES A
JRNL        TITL 2 TOPOLOGY FOR PROCESSING THREADED SUBSTRATES
JRNL        REF    J.AM.CHEM.SOC.                             2016
JRNL        REFN                   ESSN 1520-5126
JRNL        DOI    10.1021/JACS.6B10389
REMARK   2
REMARK   2 RESOLUTION.    2.20 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0073
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.20
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 29.82
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.8
REMARK   3   NUMBER OF REFLECTIONS             : 81764
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.157
REMARK   3   R VALUE            (WORKING SET) : 0.154
REMARK   3   FREE R VALUE                     : 0.198
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.900
REMARK   3   FREE R VALUE TEST SET COUNT      : 4255
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.20
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.26
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 5980
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 99.70
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2100
REMARK   3   BIN FREE R VALUE SET COUNT          : 331
REMARK   3   BIN FREE R VALUE                    : 0.2500
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 10555
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 0
REMARK   3   SOLVENT ATOMS            : 910
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 34.77
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 1.03000
REMARK   3    B22 (A**2) : -0.90000
REMARK   3    B33 (A**2) : -0.13000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.196
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.166
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.116
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 4.647
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.965
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.946
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A): 10868 ; 0.017 ; 0.019
REMARK   3   BOND LENGTHS OTHERS               (A): 10188 ; 0.006 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES): 14838 ; 1.748 ; 1.954
REMARK   3   BOND ANGLES OTHERS          (DEGREES): 23343 ; 1.191 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):  1349 ; 6.834 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   522 ;33.905 ;22.605
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):  1703 ;15.201 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):   124 ;19.764 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):  1621 ; 0.107 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A): 12458 ; 0.010 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  2608 ; 0.005 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  5372 ; 2.986 ; 3.156
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  5371 ; 2.985 ; 3.155
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  6714 ; 4.287 ; 4.719
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  6715 ; 4.287 ; 4.720
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  5495 ; 4.139 ; 3.674
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  5496 ; 4.139 ; 3.674
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  8120 ; 6.345 ; 5.308
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2): 12942 ; 8.274 ;26.341
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2): 12520 ; 8.219 ;26.029
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NCS TYPE: LOCAL
REMARK   3   NUMBER OF DIFFERENT NCS PAIRS  : 1
REMARK   3  GROUP  CHAIN1    RANGE     CHAIN2     RANGE    COUNT RMS  WEIGHT
REMARK   3    1     A    35   1016       B    35   1016   40295  0.10  0.05
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : NULL
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS
REMARK   4
REMARK   4 5TXE COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 17-NOV-16.
REMARK 100 THE DEPOSITION ID IS D_1000224986.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 17-JUN-16
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : APS
REMARK 200  BEAMLINE                       : 21-ID-D
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97853
REMARK 200  MONOCHROMATOR                  : SI(111)
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MARMOSAIC 300 MM CCD
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : AIMLESS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 86021
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.200
REMARK 200  RESOLUTION RANGE LOW       (A) : 29.817
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.9
REMARK 200  DATA REDUNDANCY                : 8.400
REMARK 200  R MERGE                    (I) : 0.10700
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 16.2000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.20
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.21
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.9
REMARK 200  DATA REDUNDANCY IN SHELL       : 8.40
REMARK 200  R MERGE FOR SHELL          (I) : 0.73700
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.800
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 52.81
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.61
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 18% PEG3350, 0.3 M AMMONIUM CITRATE
REMARK 280  DIBASIC, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 296K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 2 2 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z
REMARK 290       3555   -X,Y,-Z
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z
REMARK 290       6555   -X+1/2,-Y+1/2,Z
REMARK 290       7555   -X+1/2,Y+1/2,-Z
REMARK 290       8555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       75.62650
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000      101.42700
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       75.62650
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000      101.42700
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       75.62650
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000      101.42700
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       75.62650
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000      101.42700
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2, 3
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 2000 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 27290 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -17.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 2070 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 27410 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -17.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 3
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: TETRAMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 7200 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 51570 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -52.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, C, B, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     ARG A     2
REMARK 465     SER A     3
REMARK 465     SER A     4
REMARK 465     LYS A     5
REMARK 465     ILE A     6
REMARK 465     ARG A     7
REMARK 465     CYS A     8
REMARK 465     PRO A     9
REMARK 465     GLY A    10
REMARK 465     ALA A    11
REMARK 465     ILE A    12
REMARK 465     ARG A    13
REMARK 465     VAL A    14
REMARK 465     GLY A    15
REMARK 465     THR A    16
REMARK 465     LEU A    17
REMARK 465     VAL A    18
REMARK 465     THR A    19
REMARK 465     ALA A    20
REMARK 465     PHE A    21
REMARK 465     GLY A    22
REMARK 465     CYS A    23
REMARK 465     LEU A    24
REMARK 465     PRO A    25
REMARK 465     HIS A    26
REMARK 465     VAL A    27
REMARK 465     ALA A    28
REMARK 465     PHE A    29
REMARK 465     ALA A    30
REMARK 465     ALA A    31
REMARK 465     ALA A    32
REMARK 465     ARG A    33
REMARK 465     GLU A    34
REMARK 465     LYS A   687
REMARK 465     SER A   688
REMARK 465     PRO A   689
REMARK 465     SER A   690
REMARK 465     PRO A   691
REMARK 465     THR A   692
REMARK 465     PRO A   693
REMARK 465     ALA A   694
REMARK 465     PRO A   695
REMARK 465     GLY A   696
REMARK 465     SER A   697
REMARK 465     ARG A   698
REMARK 465     SER A   699
REMARK 465     HIS A   700
REMARK 465     HIS A   701
REMARK 465     HIS A   702
REMARK 465     HIS A   703
REMARK 465     HIS A   704
REMARK 465     HIS A   705
REMARK 465     GLN C  1018
REMARK 465     HIS C  1019
REMARK 465     ALA C  1020
REMARK 465     PRO C  1021
REMARK 465     LEU C  1022
REMARK 465     ALA C  1023
REMARK 465     ASP C  1024
REMARK 465     MET B     1
REMARK 465     ARG B     2
REMARK 465     SER B     3
REMARK 465     SER B     4
REMARK 465     LYS B     5
REMARK 465     ILE B     6
REMARK 465     ARG B     7
REMARK 465     CYS B     8
REMARK 465     PRO B     9
REMARK 465     GLY B    10
REMARK 465     ALA B    11
REMARK 465     ILE B    12
REMARK 465     ARG B    13
REMARK 465     VAL B    14
REMARK 465     GLY B    15
REMARK 465     THR B    16
REMARK 465     LEU B    17
REMARK 465     VAL B    18
REMARK 465     THR B    19
REMARK 465     ALA B    20
REMARK 465     PHE B    21
REMARK 465     GLY B    22
REMARK 465     CYS B    23
REMARK 465     LEU B    24
REMARK 465     PRO B    25
REMARK 465     HIS B    26
REMARK 465     VAL B    27
REMARK 465     ALA B    28
REMARK 465     PHE B    29
REMARK 465     ALA B    30
REMARK 465     ALA B    31
REMARK 465     ALA B    32
REMARK 465     ARG B    33
REMARK 465     GLU B    34
REMARK 465     LYS B   687
REMARK 465     SER B   688
REMARK 465     PRO B   689
REMARK 465     SER B   690
REMARK 465     PRO B   691
REMARK 465     THR B   692
REMARK 465     PRO B   693
REMARK 465     ALA B   694
REMARK 465     PRO B   695
REMARK 465     GLY B   696
REMARK 465     SER B   697
REMARK 465     ARG B   698
REMARK 465     SER B   699
REMARK 465     HIS B   700
REMARK 465     HIS B   701
REMARK 465     HIS B   702
REMARK 465     HIS B   703
REMARK 465     HIS B   704
REMARK 465     HIS B   705
REMARK 465     HIS D  1019
REMARK 465     ALA D  1020
REMARK 465     PRO D  1021
REMARK 465     LEU D  1022
REMARK 465     ALA D  1023
REMARK 465     ASP D  1024
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    HOH B  1289     O    HOH B  1316              1.89
REMARK 500   C    THR B   348     O    HOH B   802              1.91
REMARK 500   O    HOH A  1039     O    HOH A  1113              1.99
REMARK 500   OE2  GLU B   202     O    HOH B   801              2.00
REMARK 500   O    HOH B  1099     O    HOH B  1256              2.01
REMARK 500   O    HOH B  1159     O    HOH B  1277              2.08
REMARK 500   OG   SER A   157     O    HOH A   801              2.09
REMARK 500   O    HOH A  1088     O    HOH A  1110              2.09
REMARK 500   OG1  THR B   348     O    HOH B   802              2.13
REMARK 500   O    HOH B   901     O    HOH B  1171              2.17
REMARK 500   O    HOH B   812     O    HOH B  1232              2.17
REMARK 500   O    HOH A   855     O    HOH B   805              2.18
REMARK 500   O    HOH B  1042     O    HOH B  1214              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   OE2  GLU B   403     OE2  GLU B   403     6565     2.18
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),1X,F6.3)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION
REMARK 500    ARG B 241   CD    ARG B 241   NE     -0.110
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ARG A 133   CG  -  CD  -  NE  ANGL. DEV. =  13.0 DEGREES
REMARK 500    ARG A 222   NE  -  CZ  -  NH1 ANGL. DEV. =   4.3 DEGREES
REMARK 500    ARG A 677   NE  -  CZ  -  NH1 ANGL. DEV. =   4.8 DEGREES
REMARK 500    ARG A 677   NE  -  CZ  -  NH2 ANGL. DEV. =  -5.2 DEGREES
REMARK 500    ARG A 679   NE  -  CZ  -  NH1 ANGL. DEV. =   3.1 DEGREES
REMARK 500    ARG A 679   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.2 DEGREES
REMARK 500    TRP C1016   CB  -  CA  -  C   ANGL. DEV. =  13.4 DEGREES
REMARK 500    TRP C1016   CA  -  C   -  N   ANGL. DEV. =  13.4 DEGREES
REMARK 500    ARG B 152   NE  -  CZ  -  NH1 ANGL. DEV. =   3.2 DEGREES
REMARK 500    ARG B 241   NE  -  CZ  -  NH1 ANGL. DEV. =   4.2 DEGREES
REMARK 500    ARG B 241   NE  -  CZ  -  NH2 ANGL. DEV. =  -6.0 DEGREES
REMARK 500    ARG B 253   NE  -  CZ  -  NH1 ANGL. DEV. =   3.6 DEGREES
REMARK 500    ARG B 260   NE  -  CZ  -  NH1 ANGL. DEV. =   3.8 DEGREES
REMARK 500    ARG B 360   NE  -  CZ  -  NH1 ANGL. DEV. =   3.4 DEGREES
REMARK 500    ARG B 360   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.4 DEGREES
REMARK 500    ARG B 373   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.3 DEGREES
REMARK 500    ARG B 430   NE  -  CZ  -  NH1 ANGL. DEV. =   3.4 DEGREES
REMARK 500    ARG B 441   CG  -  CD  -  NE  ANGL. DEV. = -13.7 DEGREES
REMARK 500    ARG B 441   NE  -  CZ  -  NH2 ANGL. DEV. =  -4.3 DEGREES
REMARK 500    ARG B 445   CD  -  NE  -  CZ  ANGL. DEV. =  12.0 DEGREES
REMARK 500    ARG B 445   NE  -  CZ  -  NH1 ANGL. DEV. =   7.9 DEGREES
REMARK 500    ARG B 445   NE  -  CZ  -  NH2 ANGL. DEV. = -10.5 DEGREES
REMARK 500    ARG B 625   NE  -  CZ  -  NH1 ANGL. DEV. =   3.1 DEGREES
REMARK 500    ARG B 679   NE  -  CZ  -  NH1 ANGL. DEV. =   4.6 DEGREES
REMARK 500    ARG B 679   NE  -  CZ  -  NH2 ANGL. DEV. =  -4.8 DEGREES
REMARK 500    ARG B 685   NE  -  CZ  -  NH1 ANGL. DEV. =   3.1 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASP A 210     -155.19   -156.74
REMARK 500    ALA A 338     -168.92   -128.51
REMARK 500    ALA A 358     -111.99     49.08
REMARK 500    ARG A 445      -88.31   -110.03
REMARK 500    ALA A 527     -120.04     61.95
REMARK 500    SER A 559      -64.02   -103.44
REMARK 500    MET C1005     -102.33   -125.52
REMARK 500    SER C1010     -165.81     66.19
REMARK 500    ASP B 210     -155.21   -154.91
REMARK 500    ALA B 338     -167.06   -129.01
REMARK 500    ALA B 358     -113.91     55.14
REMARK 500    ARG B 445      -87.60   -114.89
REMARK 500    ALA B 527     -122.17     60.36
REMARK 500    SER B 559      -65.76   -101.67
REMARK 500    ARG B 685        2.53    -69.39
REMARK 500    MET D1005     -102.25   -126.56
REMARK 500    SER D1010     -164.04     66.77
REMARK 500    TRP D1016     -142.64   -112.53
REMARK 500    ASP D1017       56.36   -160.10
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 TRP C 1016     ASP C 1017                   64.21
REMARK 500
REMARK 500 REMARK: NULL
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 5TXC   RELATED DB: PDB
DBREF  5TXE A    1   695  UNP    E8RUP5   E8RUP5_ASTEC     1    695
DBREF  5TXE C 1001  1024  UNP    E8RUP8   E8RUP8_ASTEC    26     49
DBREF  5TXE B    1   695  UNP    E8RUP5   E8RUP5_ASTEC     1    695
DBREF  5TXE D 1001  1024  UNP    E8RUP8   E8RUP8_ASTEC    26     49
SEQADV 5TXE ALA A  527  UNP  E8RUP5    SER   527 ENGINEERED MUTATION
SEQADV 5TXE GLY A  696  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE SER A  697  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE ARG A  698  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE SER A  699  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS A  700  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS A  701  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS A  702  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS A  703  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS A  704  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS A  705  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE ALA B  527  UNP  E8RUP5    SER   527 ENGINEERED MUTATION
SEQADV 5TXE GLY B  696  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE SER B  697  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE ARG B  698  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE SER B  699  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS B  700  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS B  701  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS B  702  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS B  703  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS B  704  UNP  E8RUP5              EXPRESSION TAG
SEQADV 5TXE HIS B  705  UNP  E8RUP5              EXPRESSION TAG
SEQRES   1 A  705  MET ARG SER SER LYS ILE ARG CYS PRO GLY ALA ILE ARG
SEQRES   2 A  705  VAL GLY THR LEU VAL THR ALA PHE GLY CYS LEU PRO HIS
SEQRES   3 A  705  VAL ALA PHE ALA ALA ALA ARG GLU ALA PRO PRO VAL THR
SEQRES   4 A  705  PRO GLU VAL LEU VAL ARG LEU ALA ASP ILE GLY THR MET
SEQRES   5 A  705  SER ALA SER GLU THR THR PRO LEU LEU SER LEU SER PRO
SEQRES   6 A  705  ASP GLY ARG TYR VAL ALA PHE GLN VAL ARG GLN ALA ASP
SEQRES   7 A  705  PRO VAL THR ASN LEU ASN VAL PHE ARG MET VAL VAL LYS
SEQRES   8 A  705  ALA THR ASP GLY ALA THR ASP ALA ILE ASP VAL ASP VAL
SEQRES   9 A  705  GLY GLY GLU TYR LEU PHE TRP THR ILE PRO SER TRP GLY
SEQRES  10 A  705  TYR ALA ARG ASN ALA PRO SER GLY ALA ASN LEU THR ILE
SEQRES  11 A  705  GLN PRO ARG TRP SER PRO SER GLY THR HIS LEU ALA TYR
SEQRES  12 A  705  LEU ARG GLN ASP GLN GLY ARG VAL ARG VAL TRP ARG ALA
SEQRES  13 A  705  SER VAL LYS GLY GLU GLY ALA SER PRO VAL ILE GLU ASP
SEQRES  14 A  705  ALA TYR ASP ILE GLU ASP VAL GLN TRP LEU ASP ASP ASN
SEQRES  15 A  705  THR LEU ILE TYR SER GLY ARG PRO GLY PHE VAL GLU ALA
SEQRES  16 A  705  GLU ALA GLU ILE GLU ARG GLU GLY ARG ARG GLY TRP VAL
SEQRES  17 A  705  TYR ASP GLU ARG PHE HIS PRO LEU THR GLY ALA ARG PRO
SEQRES  18 A  705  ARG VAL LEU GLU PRO ILE SER ILE VAL TYR GLN VAL LEU
SEQRES  19 A  705  ASP LEU LYS THR GLY THR ARG ARG ALA ALA THR PRO THR
SEQRES  20 A  705  GLU VAL ALA ARG LEU ARG GLU LYS PRO ASP PRO LEU ARG
SEQRES  21 A  705  ALA MET VAL GLY ARG THR THR PHE SER VAL SER ARG THR
SEQRES  22 A  705  ASP PRO GLN ASN ILE ASN ALA PRO THR THR LEU VAL ALA
SEQRES  23 A  705  ARG ARG GLY GLU GLY GLU PRO VAL ARG CYS ASP GLU GLU
SEQRES  24 A  705  ALA CYS GLN ASN ILE THR ARG MET TRP GLY ASP GLU THR
SEQRES  25 A  705  ALA ASN VAL LEU TYR PHE LEU ARG ARG GLU GLY TRP ALA
SEQRES  26 A  705  SER ASN GLU MET ALA LEU TYR ARG MET PRO ALA ASP ALA
SEQRES  27 A  705  LEU LYS PRO VAL ARG ILE TRP HIS ALA THR GLY LEU LEU
SEQRES  28 A  705  GLN GLY CYS GLU ARG GLN ALA LYS ARG LEU ILE CYS ALA
SEQRES  29 A  705  GLN GLU SER ALA LEU GLN PRO ARG ARG LEU VAL THR LEU
SEQRES  30 A  705  ASN LEU THR SER GLY GLN MET SER PRO LEU TYR ASP PRO
SEQRES  31 A  705  ASN PRO ASP LEU SER ARG TYR ARG LEU PRO LYS VAL GLU
SEQRES  32 A  705  ARG LEU THR LEU ARG ASN ARG ASN GLY ILE GLU VAL PHE
SEQRES  33 A  705  SER ASP LEU VAL LEU PRO PRO ASP TYR GLN LEU GLY THR
SEQRES  34 A  705  ARG LEU PRO LEU VAL ILE VAL GLN TYR SER SER ARG GLY
SEQRES  35 A  705  PHE LEU ARG GLY GLY THR GLY ASP GLU ASN PRO ILE LEU
SEQRES  36 A  705  PRO LEU ALA THR ALA GLY PHE ALA VAL LEU SER PHE HIS
SEQRES  37 A  705  SER PRO ARG SER GLU ALA SER TYR GLN ARG PHE THR SER
SEQRES  38 A  705  PRO ILE ALA GLN SER LYS ALA GLU TYR SER ASN TRP ARG
SEQRES  39 A  705  ASN ARG TRP ASN ILE LEU HIS THR LEU GLU ASP LEU ILE
SEQRES  40 A  705  ASP ASP LEU ASP ARG ARG GLY VAL ILE ASP PRO ALA ARG
SEQRES  41 A  705  VAL GLY LEU THR GLY LEU ALA ASP GLY ALA THR THR VAL
SEQRES  42 A  705  HIS PHE GLY LEU ILE ASN SER HIS ARG PHE ALA ALA ALA
SEQRES  43 A  705  VAL THR SER SER CYS CYS THR ASP SER PHE THR ALA SER
SEQRES  44 A  705  VAL MET ASN GLY PRO ARG ILE SER GLY ALA LEU LYS ALA
SEQRES  45 A  705  TYR GLY ILE GLU THR ASP GLN ALA ASP ASP GLY PRO PHE
SEQRES  46 A  705  TRP ALA ALA THR SER PHE VAL VAL ASN ALA SER ARG LEU
SEQRES  47 A  705  ASP THR PRO LEU LEU ILE GLN SER ALA ASP GLU GLU TYR
SEQRES  48 A  705  LEU GLY ALA LEU PRO GLY PHE THR ALA LEU GLN GLN ALA
SEQRES  49 A  705  ARG LYS PRO VAL GLU LEU ILE ILE TYR PRO ASN GLU HIS
SEQRES  50 A  705  HIS VAL LYS TRP GLN PRO ALA HIS ARG LEU ALA VAL TYR
SEQRES  51 A  705  ASN ARG THR ILE ASP TRP PHE ARG PHE TRP LEU MET ASP
SEQRES  52 A  705  GLN SER ASP PRO ALA PRO ASP LYS ALA ALA GLN TYR ASP
SEQRES  53 A  705  ARG TRP ARG ALA LEU ARG ALA LEU ARG GLN LYS SER PRO
SEQRES  54 A  705  SER PRO THR PRO ALA PRO GLY SER ARG SER HIS HIS HIS
SEQRES  55 A  705  HIS HIS HIS
SEQRES   1 C   24  GLY PRO THR PRO MET VAL GLY LEU ASP SER VAL SER GLY
SEQRES   2 C   24  GLN TYR TRP ASP GLN HIS ALA PRO LEU ALA ASP
SEQRES   1 B  705  MET ARG SER SER LYS ILE ARG CYS PRO GLY ALA ILE ARG
SEQRES   2 B  705  VAL GLY THR LEU VAL THR ALA PHE GLY CYS LEU PRO HIS
SEQRES   3 B  705  VAL ALA PHE ALA ALA ALA ARG GLU ALA PRO PRO VAL THR
SEQRES   4 B  705  PRO GLU VAL LEU VAL ARG LEU ALA ASP ILE GLY THR MET
SEQRES   5 B  705  SER ALA SER GLU THR THR PRO LEU LEU SER LEU SER PRO
SEQRES   6 B  705  ASP GLY ARG TYR VAL ALA PHE GLN VAL ARG GLN ALA ASP
SEQRES   7 B  705  PRO VAL THR ASN LEU ASN VAL PHE ARG MET VAL VAL LYS
SEQRES   8 B  705  ALA THR ASP GLY ALA THR ASP ALA ILE ASP VAL ASP VAL
SEQRES   9 B  705  GLY GLY GLU TYR LEU PHE TRP THR ILE PRO SER TRP GLY
SEQRES  10 B  705  TYR ALA ARG ASN ALA PRO SER GLY ALA ASN LEU THR ILE
SEQRES  11 B  705  GLN PRO ARG TRP SER PRO SER GLY THR HIS LEU ALA TYR
SEQRES  12 B  705  LEU ARG GLN ASP GLN GLY ARG VAL ARG VAL TRP ARG ALA
SEQRES  13 B  705  SER VAL LYS GLY GLU GLY ALA SER PRO VAL ILE GLU ASP
SEQRES  14 B  705  ALA TYR ASP ILE GLU ASP VAL GLN TRP LEU ASP ASP ASN
SEQRES  15 B  705  THR LEU ILE TYR SER GLY ARG PRO GLY PHE VAL GLU ALA
SEQRES  16 B  705  GLU ALA GLU ILE GLU ARG GLU GLY ARG ARG GLY TRP VAL
SEQRES  17 B  705  TYR ASP GLU ARG PHE HIS PRO LEU THR GLY ALA ARG PRO
SEQRES  18 B  705  ARG VAL LEU GLU PRO ILE SER ILE VAL TYR GLN VAL LEU
SEQRES  19 B  705  ASP LEU LYS THR GLY THR ARG ARG ALA ALA THR PRO THR
SEQRES  20 B  705  GLU VAL ALA ARG LEU ARG GLU LYS PRO ASP PRO LEU ARG
SEQRES  21 B  705  ALA MET VAL GLY ARG THR THR PHE SER VAL SER ARG THR
SEQRES  22 B  705  ASP PRO GLN ASN ILE ASN ALA PRO THR THR LEU VAL ALA
SEQRES  23 B  705  ARG ARG GLY GLU GLY GLU PRO VAL ARG CYS ASP GLU GLU
SEQRES  24 B  705  ALA CYS GLN ASN ILE THR ARG MET TRP GLY ASP GLU THR
SEQRES  25 B  705  ALA ASN VAL LEU TYR PHE LEU ARG ARG GLU GLY TRP ALA
SEQRES  26 B  705  SER ASN GLU MET ALA LEU TYR ARG MET PRO ALA ASP ALA
SEQRES  27 B  705  LEU LYS PRO VAL ARG ILE TRP HIS ALA THR GLY LEU LEU
SEQRES  28 B  705  GLN GLY CYS GLU ARG GLN ALA LYS ARG LEU ILE CYS ALA
SEQRES  29 B  705  GLN GLU SER ALA LEU GLN PRO ARG ARG LEU VAL THR LEU
SEQRES  30 B  705  ASN LEU THR SER GLY GLN MET SER PRO LEU TYR ASP PRO
SEQRES  31 B  705  ASN PRO ASP LEU SER ARG TYR ARG LEU PRO LYS VAL GLU
SEQRES  32 B  705  ARG LEU THR LEU ARG ASN ARG ASN GLY ILE GLU VAL PHE
SEQRES  33 B  705  SER ASP LEU VAL LEU PRO PRO ASP TYR GLN LEU GLY THR
SEQRES  34 B  705  ARG LEU PRO LEU VAL ILE VAL GLN TYR SER SER ARG GLY
SEQRES  35 B  705  PHE LEU ARG GLY GLY THR GLY ASP GLU ASN PRO ILE LEU
SEQRES  36 B  705  PRO LEU ALA THR ALA GLY PHE ALA VAL LEU SER PHE HIS
SEQRES  37 B  705  SER PRO ARG SER GLU ALA SER TYR GLN ARG PHE THR SER
SEQRES  38 B  705  PRO ILE ALA GLN SER LYS ALA GLU TYR SER ASN TRP ARG
SEQRES  39 B  705  ASN ARG TRP ASN ILE LEU HIS THR LEU GLU ASP LEU ILE
SEQRES  40 B  705  ASP ASP LEU ASP ARG ARG GLY VAL ILE ASP PRO ALA ARG
SEQRES  41 B  705  VAL GLY LEU THR GLY LEU ALA ASP GLY ALA THR THR VAL
SEQRES  42 B  705  HIS PHE GLY LEU ILE ASN SER HIS ARG PHE ALA ALA ALA
SEQRES  43 B  705  VAL THR SER SER CYS CYS THR ASP SER PHE THR ALA SER
SEQRES  44 B  705  VAL MET ASN GLY PRO ARG ILE SER GLY ALA LEU LYS ALA
SEQRES  45 B  705  TYR GLY ILE GLU THR ASP GLN ALA ASP ASP GLY PRO PHE
SEQRES  46 B  705  TRP ALA ALA THR SER PHE VAL VAL ASN ALA SER ARG LEU
SEQRES  47 B  705  ASP THR PRO LEU LEU ILE GLN SER ALA ASP GLU GLU TYR
SEQRES  48 B  705  LEU GLY ALA LEU PRO GLY PHE THR ALA LEU GLN GLN ALA
SEQRES  49 B  705  ARG LYS PRO VAL GLU LEU ILE ILE TYR PRO ASN GLU HIS
SEQRES  50 B  705  HIS VAL LYS TRP GLN PRO ALA HIS ARG LEU ALA VAL TYR
SEQRES  51 B  705  ASN ARG THR ILE ASP TRP PHE ARG PHE TRP LEU MET ASP
SEQRES  52 B  705  GLN SER ASP PRO ALA PRO ASP LYS ALA ALA GLN TYR ASP
SEQRES  53 B  705  ARG TRP ARG ALA LEU ARG ALA LEU ARG GLN LYS SER PRO
SEQRES  54 B  705  SER PRO THR PRO ALA PRO GLY SER ARG SER HIS HIS HIS
SEQRES  55 B  705  HIS HIS HIS
SEQRES   1 D   24  GLY PRO THR PRO MET VAL GLY LEU ASP SER VAL SER GLY
SEQRES   2 D   24  GLN TYR TRP ASP GLN HIS ALA PRO LEU ALA ASP
FORMUL   5  HOH   *910(H2 O)
HELIX    1 AA1 THR A   39  ARG A   45  1                                   7
HELIX    2 AA2 ARG A  189  GLY A  203  1                                  15
HELIX    3 AA3 ARG A  204  GLY A  206  5                                   3
HELIX    4 AA4 THR A  245  GLU A  254  1                                  10
HELIX    5 AA5 GLU A  298  GLN A  302  5                                   5
HELIX    6 AA6 GLY A  323  ASN A  327  5                                   5
HELIX    7 AA7 ASP A  393  TYR A  397  5                                   5
HELIX    8 AA8 PRO A  453  ALA A  460  1                                   8
HELIX    9 AA9 SER A  472  GLN A  477  5                                   6
HELIX   10 AB1 SER A  481  SER A  491  1                                  11
HELIX   11 AB2 TRP A  493  ARG A  513  1                                  21
HELIX   12 AB3 ALA A  527  SER A  540  1                                  14
HELIX   13 AB4 ASP A  554  SER A  559  1                                   6
HELIX   14 AB5 ASN A  562  LYS A  571  1                                  10
HELIX   15 AB6 ALA A  572  GLY A  574  5                                   3
HELIX   16 AB7 PHE A  585  THR A  589  5                                   5
HELIX   17 AB8 SER A  590  ALA A  595  1                                   6
HELIX   18 AB9 SER A  596  LEU A  598  5                                   3
HELIX   19 AC1 ASP A  608  LEU A  612  1                                   5
HELIX   20 AC2 ALA A  614  ALA A  624  1                                  11
HELIX   21 AC3 GLN A  642  ASP A  663  1                                  22
HELIX   22 AC4 ALA A  668  ASP A  670  5                                   3
HELIX   23 AC5 LYS A  671  ARG A  685  1                                  15
HELIX   24 AC6 THR B   39  ARG B   45  1                                   7
HELIX   25 AC7 ARG B  189  GLY B  203  1                                  15
HELIX   26 AC8 ARG B  204  GLY B  206  5                                   3
HELIX   27 AC9 THR B  245  GLU B  254  1                                  10
HELIX   28 AD1 GLU B  298  GLN B  302  5                                   5
HELIX   29 AD2 GLY B  323  ASN B  327  5                                   5
HELIX   30 AD3 ASP B  393  TYR B  397  5                                   5
HELIX   31 AD4 PRO B  453  ALA B  460  1                                   8
HELIX   32 AD5 GLU B  473  GLN B  477  5                                   5
HELIX   33 AD6 SER B  481  SER B  491  1                                  11
HELIX   34 AD7 TRP B  493  ARG B  513  1                                  21
HELIX   35 AD8 ALA B  527  SER B  540  1                                  14
HELIX   36 AD9 ASP B  554  SER B  559  1                                   6
HELIX   37 AE1 ASN B  562  LYS B  571  1                                  10
HELIX   38 AE2 ALA B  572  GLY B  574  5                                   3
HELIX   39 AE3 PHE B  585  THR B  589  5                                   5
HELIX   40 AE4 SER B  590  ALA B  595  1                                   6
HELIX   41 AE5 SER B  596  LEU B  598  5                                   3
HELIX   42 AE6 ASP B  608  LEU B  612  1                                   5
HELIX   43 AE7 ALA B  614  ALA B  624  1                                  11
HELIX   44 AE8 GLN B  642  ASP B  663  1                                  22
HELIX   45 AE9 ALA B  668  ASP B  670  5                                   3
HELIX   46 AF1 LYS B  671  ARG B  685  1                                  15
SHEET    1 AA1 3 ASP A  48  ILE A  49  0
SHEET    2 AA1 3 TYR A  69  ASP A  78 -1  O  ARG A  75   N  ASP A  48
SHEET    3 AA1 3 LEU A  61  LEU A  63 -1  N  SER A  62   O  ALA A  71
SHEET    1 AA2 4 ASP A  48  ILE A  49  0
SHEET    2 AA2 4 TYR A  69  ASP A  78 -1  O  ARG A  75   N  ASP A  48
SHEET    3 AA2 4 LEU A  83  ALA A  92 -1  O  VAL A  89   N  PHE A  72
SHEET    4 AA2 4 ILE A 100  VAL A 104 -1  O  VAL A 102   N  MET A  88
SHEET    1 AA3 4 ARG A 133  TRP A 134  0
SHEET    2 AA3 4 HIS A 140  ASP A 147 -1  O  ALA A 142   N  ARG A 133
SHEET    3 AA3 4 ARG A 150  SER A 157 -1  O  TRP A 154   N  TYR A 143
SHEET    4 AA3 4 ALA A 163  ILE A 167 -1  O  VAL A 166   N  VAL A 153
SHEET    1 AA4 4 ILE A 173  ASP A 180  0
SHEET    2 AA4 4 THR A 183  GLY A 188 -1  O  ILE A 185   N  GLN A 177
SHEET    3 AA4 4 VAL A 230  ASP A 235 -1  O  LEU A 234   N  LEU A 184
SHEET    4 AA4 4 ARG A 241  ALA A 243 -1  O  ARG A 242   N  VAL A 233
SHEET    1 AA5 9 TRP A 207  VAL A 208  0
SHEET    2 AA5 9 VAL B 628  TYR B 633 -1  O  ILE B 632   N  TRP A 207
SHEET    3 AA5 9 LEU B 602  ALA B 607  1  N  ILE B 604   O  ILE B 631
SHEET    4 AA5 9 ALA B 545  THR B 548  1  N  THR B 548   O  LEU B 603
SHEET    5 AA5 9 ILE B 516  GLY B 525  1  N  LEU B 523   O  VAL B 547
SHEET    6 AA5 9 LEU B 431  VAL B 436  1  N  LEU B 433   O  GLY B 522
SHEET    7 AA5 9 ALA B 463  PHE B 467  1  O  LEU B 465   N  VAL B 436
SHEET    8 AA5 9 GLU B 414  LEU B 421 -1  N  ASP B 418   O  SER B 466
SHEET    9 AA5 9 LYS B 401  ARG B 408 -1  N  LEU B 405   O  SER B 417
SHEET    1 AA6 4 ALA A 261  VAL A 263  0
SHEET    2 AA6 4 THR A 266  SER A 271 -1  O  PHE A 268   N  ALA A 261
SHEET    3 AA6 4 THR A 283  ARG A 288 -1  O  VAL A 285   N  SER A 269
SHEET    4 AA6 4 VAL A 294  ARG A 295 -1  O  VAL A 294   N  ALA A 286
SHEET    1 AA7 4 ILE A 304  ASP A 310  0
SHEET    2 AA7 4 VAL A 315  GLU A 322 -1  O  LEU A 319   N  ARG A 306
SHEET    3 AA7 4 GLU A 328  PRO A 335 -1  O  TYR A 332   N  PHE A 318
SHEET    4 AA7 4 VAL A 342  ALA A 347 -1  O  VAL A 342   N  ARG A 333
SHEET    1 AA8 4 LEU A 350  GLN A 357  0
SHEET    2 AA8 4 ARG A 360  SER A 367 -1  O  ILE A 362   N  GLU A 355
SHEET    3 AA8 4 GLN A 370  ASN A 378 -1  O  ARG A 373   N  GLN A 365
SHEET    4 AA8 4 MET A 384  TYR A 388 -1  O  TYR A 388   N  LEU A 374
SHEET    1 AA9 9 LYS A 401  ARG A 408  0
SHEET    2 AA9 9 GLU A 414  LEU A 421 -1  O  SER A 417   N  LEU A 405
SHEET    3 AA9 9 ALA A 463  PHE A 467 -1  O  SER A 466   N  ASP A 418
SHEET    4 AA9 9 LEU A 431  VAL A 436  1  N  VAL A 436   O  PHE A 467
SHEET    5 AA9 9 ILE A 516  GLY A 525  1  O  GLY A 522   N  LEU A 433
SHEET    6 AA9 9 ALA A 545  THR A 548  1  O  ALA A 545   N  LEU A 523
SHEET    7 AA9 9 LEU A 602  ALA A 607  1  O  GLN A 605   N  THR A 548
SHEET    8 AA9 9 VAL A 628  TYR A 633  1  O  ILE A 631   N  ILE A 604
SHEET    9 AA9 9 TRP B 207  VAL B 208 -1  O  TRP B 207   N  ILE A 632
SHEET    1 AB1 2 LEU C1008  ASP C1009  0
SHEET    2 AB1 2 GLN C1014  TYR C1015 -1  O  GLN C1014   N  ASP C1009
SHEET    1 AB2 3 ASP B  48  ILE B  49  0
SHEET    2 AB2 3 TYR B  69  ASP B  78 -1  O  ARG B  75   N  ASP B  48
SHEET    3 AB2 3 LEU B  61  LEU B  63 -1  N  SER B  62   O  ALA B  71
SHEET    1 AB3 4 ASP B  48  ILE B  49  0
SHEET    2 AB3 4 TYR B  69  ASP B  78 -1  O  ARG B  75   N  ASP B  48
SHEET    3 AB3 4 LEU B  83  ALA B  92 -1  O  VAL B  89   N  PHE B  72
SHEET    4 AB3 4 ILE B 100  VAL B 104 -1  O  VAL B 102   N  MET B  88
SHEET    1 AB4 4 ARG B 133  TRP B 134  0
SHEET    2 AB4 4 HIS B 140  ASP B 147 -1  O  ALA B 142   N  ARG B 133
SHEET    3 AB4 4 ARG B 150  SER B 157 -1  O  TRP B 154   N  TYR B 143
SHEET    4 AB4 4 ALA B 163  ILE B 167 -1  O  VAL B 166   N  VAL B 153
SHEET    1 AB5 4 ILE B 173  ASP B 180  0
SHEET    2 AB5 4 THR B 183  GLY B 188 -1  O  SER B 187   N  GLU B 174
SHEET    3 AB5 4 VAL B 230  ASP B 235 -1  O  LEU B 234   N  LEU B 184
SHEET    4 AB5 4 ARG B 241  ALA B 243 -1  O  ARG B 242   N  VAL B 233
SHEET    1 AB6 4 ALA B 261  VAL B 263  0
SHEET    2 AB6 4 THR B 266  SER B 271 -1  O  PHE B 268   N  ALA B 261
SHEET    3 AB6 4 THR B 283  ARG B 288 -1  O  VAL B 285   N  SER B 269
SHEET    4 AB6 4 VAL B 294  ARG B 295 -1  O  VAL B 294   N  ALA B 286
SHEET    1 AB7 4 ILE B 304  ASP B 310  0
SHEET    2 AB7 4 VAL B 315  GLU B 322 -1  O  TYR B 317   N  TRP B 308
SHEET    3 AB7 4 GLU B 328  PRO B 335 -1  O  TYR B 332   N  PHE B 318
SHEET    4 AB7 4 VAL B 342  ALA B 347 -1  O  ILE B 344   N  LEU B 331
SHEET    1 AB8 4 LEU B 350  GLN B 357  0
SHEET    2 AB8 4 ARG B 360  SER B 367 -1  O  ILE B 362   N  GLU B 355
SHEET    3 AB8 4 GLN B 370  ASN B 378 -1  O  LEU B 377   N  LEU B 361
SHEET    4 AB8 4 MET B 384  TYR B 388 -1  O  TYR B 388   N  LEU B 374
SHEET    1 AB9 2 LEU D1008  ASP D1009  0
SHEET    2 AB9 2 GLN D1014  TYR D1015 -1  O  GLN D1014   N  ASP D1009
SSBOND   1 CYS A  296    CYS A  301                          1555   1555  2.17
SSBOND   2 CYS A  354    CYS A  363                          1555   1555  2.17
SSBOND   3 CYS A  551    CYS A  552                          1555   1555  2.12
SSBOND   4 CYS B  296    CYS B  301                          1555   1555  2.21
SSBOND   5 CYS B  354    CYS B  363                          1555   1555  2.22
SSBOND   6 CYS B  551    CYS B  552                          1555   1555  2.17
LINK         N   GLY C1001                 CG  ASP C1009     1555   1555  1.25
LINK         N   GLY D1001                 CG  ASP D1009     1555   1555  1.27
CISPEP   1 GLU A  225    PRO A  226          0         8.46
CISPEP   2 GLN A  437    TYR A  438          0        -5.77
CISPEP   3 CYS A  551    CYS A  552          0         8.44
CISPEP   4 GLU B  225    PRO B  226          0        -0.33
CISPEP   5 GLN B  437    TYR B  438          0        -4.68
CISPEP   6 CYS B  551    CYS B  552          0         6.86
CRYST1  151.253  202.854  110.399  90.00  90.00  90.00 C 2 2 2      16
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.006611  0.000000  0.000000        0.00000
SCALE2      0.000000  0.004930  0.000000        0.00000
SCALE3      0.000000  0.000000  0.009058        0.00000
TER    5162      GLN A 686
TER    5288      ASP C1017
TER   10460      GLN B 686
TER   10595      GLN D1018
MASTER      561    0    0   46   76    0    0    611465    4   16  114
END