longtext: 5x6s-pdb

content
HEADER    HYDROLASE                               23-FEB-17   5X6S
TITLE     ACETYL XYLAN ESTERASE FROM ASPERGILLUS AWAMORI
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ACETYLXYLAN ESTERASE A;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: UNP RESIDUES 30-304;
COMPND   5 EC: 3.1.1.72;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: ASPERGILLUS AWAMORI;
SOURCE   3 ORGANISM_COMMON: BLACK KOJI MOLD;
SOURCE   4 ORGANISM_TAXID: 105351;
SOURCE   5 GENE: AXEA, ACEA;
SOURCE   6 EXPRESSION_SYSTEM: KOMAGATAELLA PASTORIS;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 4922;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: KM71H;
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PPICZALPHA
KEYWDS    ALPHA/BETA-HYDROLASE, ESTRASE_PHB, CAZY CE1, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    D.KOMIYA,T.KOSEKI,S.FUSHINOBU
REVDAT   1   23-AUG-17 5X6S    0
JRNL        AUTH   D.KOMIYA,A.HORI,T.ISHIDA,K.IGARASHI,M.SAMEJIMA,T.KOSEKI,
JRNL        AUTH 2 S.FUSHINOBU
JRNL        TITL   CRYSTAL STRUCTURE AND SUBSTRATE SPECIFICITY MODIFICATION OF
JRNL        TITL 2 ACETYL XYLAN ESTERASE FROM ASPERGILLUS LUCHUENSIS
JRNL        REF    APPL. ENVIRON. MICROBIOL.                  2017
JRNL        REFN                   ESSN 1098-5336
JRNL        PMID   28802264
JRNL        DOI    10.1128/AEM.01251-17
REMARK   1
REMARK   1 REFERENCE 1
REMARK   1  AUTH   T.KOSEKI,S.FURUSE,K.IWANO,H.SAKAI,H.MATSUZAWA
REMARK   1  TITL   AN ASPERGILLUS AWAMORI ACETYLESTERASE: PURIFICATION OF THE
REMARK   1  TITL 2 ENZYME, AND CLONING AND SEQUENCING OF THE GENE
REMARK   1  REF    BIOCHEM. J.                   V.T 2)   485 1997
REMARK   1  REFN                   ISSN 0264-6021
REMARK   1  PMID   9291122
REMARK   1  DOI    10.1042/BJ3260485
REMARK   1 REFERENCE 2
REMARK   1  AUTH   T.KOSEKI,Y.MIWA,S.FUSHINOBU,K.HASHIZUME
REMARK   1  TITL   BIOCHEMICAL CHARACTERIZATION OF RECOMBINANT ACETYL XYLAN
REMARK   1  TITL 2 ESTERASE FROM ASPERGILLUS AWAMORI EXPRESSED IN PICHIA
REMARK   1  TITL 3 PASTORIS: MUTATIONAL ANALYSIS OF CATALYTIC RESIDUES
REMARK   1  REF    BIOCHIM. BIOPHYS. ACTA        V.1749     7 2005
REMARK   1  REFN                   ISSN 0006-3002
REMARK   1  PMID   15848131
REMARK   1  DOI    10.1016/J.BBAPAP.2005.01.009
REMARK   2
REMARK   2 RESOLUTION.    1.90 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0155
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.90
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 87.08
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 98.1
REMARK   3   NUMBER OF REFLECTIONS             : 42045
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.153
REMARK   3   R VALUE            (WORKING SET) : 0.151
REMARK   3   FREE R VALUE                     : 0.185
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.100
REMARK   3   FREE R VALUE TEST SET COUNT      : 2238
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.90
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.95
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 3047
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 95.55
REMARK   3   BIN R VALUE           (WORKING SET) : 0.1950
REMARK   3   BIN FREE R VALUE SET COUNT          : 172
REMARK   3   BIN FREE R VALUE                    : 0.2410
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4148
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 96
REMARK   3   SOLVENT ATOMS            : 510
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 22.13
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.00000
REMARK   3    B22 (A**2) : 0.00000
REMARK   3    B33 (A**2) : 0.01000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.133
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.121
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): NULL
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): NULL
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.967
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.953
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  4376 ; 0.016 ; 0.020
REMARK   3   BOND LENGTHS OTHERS               (A):  3715 ; 0.000 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  5996 ; 1.522 ; 1.936
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  8591 ; 3.865 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   546 ; 6.129 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   188 ;36.061 ;24.894
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   564 ;12.223 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):     4 ; 7.467 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   640 ; 0.105 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  5078 ; 0.009 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  1042 ; 0.018 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  2190 ; 1.150 ; 2.114
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  2189 ; 1.139 ; 2.113
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  2734 ; 1.577 ; 3.164
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  2735 ; 1.582 ; 3.164
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  2186 ; 1.835 ; 2.244
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  2186 ; 1.835 ; 2.244
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  3263 ; 2.517 ; 3.323
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  5255 ; 3.898 ;26.248
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  5143 ; 3.689 ;25.703
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : NULL
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS
REMARK   4
REMARK   4 5X6S COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 23-FEB-17.
REMARK 100 THE DEPOSITION ID IS D_1300003016.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 04-NOV-07
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : PHOTON FACTORY
REMARK 200  BEAMLINE                       : BL-6A
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.000
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 4R
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 44285
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.900
REMARK 200  RESOLUTION RANGE LOW       (A) : 87.080
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.1
REMARK 200  DATA REDUNDANCY                : 3.300
REMARK 200  R MERGE                    (I) : 0.07100
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 16.4000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.90
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.97
REMARK 200  COMPLETENESS FOR SHELL     (%) : 95.5
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.00
REMARK 200  R MERGE FOR SHELL          (I) : 0.35900
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.500
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: SAD
REMARK 200 SOFTWARE USED: SOLVE
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 50.91
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.51
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 21-22.5% PEG 8000, 0.1M MES-NAOH (PH
REMARK 280  5.0), 0.2M CALCIUM ACETATE, 5% MPD, VAPOR DIFFUSION, HANGING
REMARK 280  DROP, TEMPERATURE 298K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: H 3
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z
REMARK 290       3555   -X+Y,-X,Z
REMARK 290       4555   X+2/3,Y+1/3,Z+1/3
REMARK 290       5555   -Y+2/3,X-Y+1/3,Z+1/3
REMARK 290       6555   -X+Y+2/3,-X+1/3,Z+1/3
REMARK 290       7555   X+1/3,Y+2/3,Z+2/3
REMARK 290       8555   -Y+1/3,X-Y+2/3,Z+2/3
REMARK 290       9555   -X+Y+1/3,-X+2/3,Z+2/3
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       87.07550
REMARK 290   SMTRY2   4  0.000000  1.000000  0.000000       50.27306
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000       16.90267
REMARK 290   SMTRY1   5 -0.500000 -0.866025  0.000000       87.07550
REMARK 290   SMTRY2   5  0.866025 -0.500000  0.000000       50.27306
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000       16.90267
REMARK 290   SMTRY1   6 -0.500000  0.866025  0.000000       87.07550
REMARK 290   SMTRY2   6 -0.866025 -0.500000  0.000000       50.27306
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       16.90267
REMARK 290   SMTRY1   7  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000      100.54613
REMARK 290   SMTRY3   7  0.000000  0.000000  1.000000       33.80533
REMARK 290   SMTRY1   8 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   8  0.866025 -0.500000  0.000000      100.54613
REMARK 290   SMTRY3   8  0.000000  0.000000  1.000000       33.80533
REMARK 290   SMTRY1   9 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   9 -0.866025 -0.500000  0.000000      100.54613
REMARK 290   SMTRY3   9  0.000000  0.000000  1.000000       33.80533
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 4270 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 19350 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -23.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     SER A     1
REMARK 465     SER B     1
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASN A  13       56.24   -152.79
REMARK 500    THR A  41       -4.94     71.48
REMARK 500    SER A 119     -119.92     54.45
REMARK 500    SER A 143       68.91     61.78
REMARK 500    TYR A 151      132.00    -34.46
REMARK 500    ASN B  13       58.81   -154.50
REMARK 500    ASN B  26       51.82    -90.41
REMARK 500    THR B  41       -4.58     66.36
REMARK 500    SER B 119     -123.50     56.11
REMARK 500    SER B 143       66.02     63.92
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 669        DISTANCE =  6.54 ANGSTROMS
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA A 304  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASN A  30   OD1
REMARK 620 2 HOH A 488   O    86.3
REMARK 620 3 HOH B 588   O   128.9  77.6
REMARK 620 4 HOH B 433   O    85.6 168.4 114.0
REMARK 620 5 HOH A 604   O    87.8  83.3 136.3  88.1
REMARK 620 N                    1     2     3     4
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA A 305  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 GLU A 134   O
REMARK 620 2 GLU B 134   O   167.6
REMARK 620 3 HOH B 551   O    88.7  90.1
REMARK 620 4 HOH A 607   O    80.8 101.3 168.3
REMARK 620 N                    1     2     3
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA B 306  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASN B  30   OD1
REMARK 620 2 HOH A 445   O    92.8
REMARK 620 3 HOH A 570   O    80.0  76.7
REMARK 620 4 HOH A 627   O   138.7 113.9  76.6
REMARK 620 5 HOH B 524   O    86.2 164.0 118.7  76.1
REMARK 620 6 HOH B 575   O    82.8  83.6 153.0 128.9  80.5
REMARK 620 N                    1     2     3     4     5
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA B 305  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 TYR B  48   O
REMARK 620 2 SER B  51   O    75.1
REMARK 620 3 SER B  51   OG   89.7  66.0
REMARK 620 4 TYR B  66   OH  114.9 169.8 110.7
REMARK 620 N                    1     2     3
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue MES A 303
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NA A 304
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NA A 305
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue MES B 303
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue MES B 304
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NA B 305
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NA B 306
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NAG A
REMARK 800  301 through NAG A 302 bound to ASN A 161
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NAG B
REMARK 800  301 through NAG B 302 bound to ASN B 161
DBREF  5X6S A    1   275  UNP    Q92194   AXE1_ASPAW      30    304
DBREF  5X6S B    1   275  UNP    Q92194   AXE1_ASPAW      30    304
SEQRES   1 A  275  SER GLY SER LEU GLN GLN VAL THR ASP PHE GLY ASP ASN
SEQRES   2 A  275  PRO THR ASN VAL GLY MET TYR ILE TYR VAL PRO ASN ASN
SEQRES   3 A  275  LEU ALA SER ASN PRO GLY ILE VAL VAL ALA ILE HIS TYR
SEQRES   4 A  275  CYS THR GLY THR GLY PRO GLY TYR TYR GLY ASP SER PRO
SEQRES   5 A  275  TYR ALA THR LEU SER GLU GLN TYR GLY PHE ILE VAL ILE
SEQRES   6 A  275  TYR PRO SER SER PRO TYR SER GLY GLY CYS TRP ASP VAL
SEQRES   7 A  275  SER SER GLN ALA THR LEU THR HIS ASN GLY GLY GLY ASN
SEQRES   8 A  275  SER ASN SER ILE ALA ASN MET VAL THR TRP THR ILE SER
SEQRES   9 A  275  LYS TYR GLY ALA ASP SER SER LYS VAL PHE VAL THR GLY
SEQRES  10 A  275  SER SER SER GLY ALA MET MET THR ASN VAL MET ALA ALA
SEQRES  11 A  275  THR TYR PRO GLU LEU PHE ALA ALA ALA THR VAL TYR SER
SEQRES  12 A  275  GLY VAL SER ALA GLY CYS PHE TYR SER ASN THR ASN GLN
SEQRES  13 A  275  VAL ASP GLY TRP ASN SER THR CYS ALA GLN GLY ASP VAL
SEQRES  14 A  275  ILE THR THR PRO GLU HIS TRP ALA SER ILE ALA GLU ALA
SEQRES  15 A  275  MET TYR SER GLY TYR SER GLY SER ARG PRO ARG MET GLN
SEQRES  16 A  275  ILE TYR HIS GLY SER ILE ASP THR THR LEU TYR PRO GLN
SEQRES  17 A  275  ASN TYR TYR GLU THR CYS LYS GLN TRP ALA GLY VAL PHE
SEQRES  18 A  275  GLY TYR ASP TYR SER ALA PRO GLU LYS THR GLU ALA ASN
SEQRES  19 A  275  THR PRO GLN THR ASN TYR GLU THR THR ILE TRP GLY ASP
SEQRES  20 A  275  SER LEU GLN GLY ILE PHE ALA THR GLY VAL GLY HIS THR
SEQRES  21 A  275  VAL PRO ILE HIS GLY ASP LYS ASP MET GLU TRP PHE GLY
SEQRES  22 A  275  PHE ALA
SEQRES   1 B  275  SER GLY SER LEU GLN GLN VAL THR ASP PHE GLY ASP ASN
SEQRES   2 B  275  PRO THR ASN VAL GLY MET TYR ILE TYR VAL PRO ASN ASN
SEQRES   3 B  275  LEU ALA SER ASN PRO GLY ILE VAL VAL ALA ILE HIS TYR
SEQRES   4 B  275  CYS THR GLY THR GLY PRO GLY TYR TYR GLY ASP SER PRO
SEQRES   5 B  275  TYR ALA THR LEU SER GLU GLN TYR GLY PHE ILE VAL ILE
SEQRES   6 B  275  TYR PRO SER SER PRO TYR SER GLY GLY CYS TRP ASP VAL
SEQRES   7 B  275  SER SER GLN ALA THR LEU THR HIS ASN GLY GLY GLY ASN
SEQRES   8 B  275  SER ASN SER ILE ALA ASN MET VAL THR TRP THR ILE SER
SEQRES   9 B  275  LYS TYR GLY ALA ASP SER SER LYS VAL PHE VAL THR GLY
SEQRES  10 B  275  SER SER SER GLY ALA MET MET THR ASN VAL MET ALA ALA
SEQRES  11 B  275  THR TYR PRO GLU LEU PHE ALA ALA ALA THR VAL TYR SER
SEQRES  12 B  275  GLY VAL SER ALA GLY CYS PHE TYR SER ASN THR ASN GLN
SEQRES  13 B  275  VAL ASP GLY TRP ASN SER THR CYS ALA GLN GLY ASP VAL
SEQRES  14 B  275  ILE THR THR PRO GLU HIS TRP ALA SER ILE ALA GLU ALA
SEQRES  15 B  275  MET TYR SER GLY TYR SER GLY SER ARG PRO ARG MET GLN
SEQRES  16 B  275  ILE TYR HIS GLY SER ILE ASP THR THR LEU TYR PRO GLN
SEQRES  17 B  275  ASN TYR TYR GLU THR CYS LYS GLN TRP ALA GLY VAL PHE
SEQRES  18 B  275  GLY TYR ASP TYR SER ALA PRO GLU LYS THR GLU ALA ASN
SEQRES  19 B  275  THR PRO GLN THR ASN TYR GLU THR THR ILE TRP GLY ASP
SEQRES  20 B  275  SER LEU GLN GLY ILE PHE ALA THR GLY VAL GLY HIS THR
SEQRES  21 B  275  VAL PRO ILE HIS GLY ASP LYS ASP MET GLU TRP PHE GLY
SEQRES  22 B  275  PHE ALA
MODRES 5X6S NAG A  301  NAG  -D
MODRES 5X6S NAG A  302  NAG  -D
MODRES 5X6S NAG B  301  NAG  -D
MODRES 5X6S NAG B  302  NAG  -D
HET    NAG  A 301      14
HET    NAG  A 302      14
HET    MES  A 303      12
HET     NA  A 304       1
HET     NA  A 305       1
HET    NAG  B 301      14
HET    NAG  B 302      14
HET    MES  B 303      12
HET    MES  B 304      12
HET     NA  B 305       1
HET     NA  B 306       1
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     MES 2-(N-MORPHOLINO)-ETHANESULFONIC ACID
HETNAM      NA SODIUM ION
FORMUL   3  NAG    4(C8 H15 N O6)
FORMUL   4  MES    3(C6 H13 N O4 S)
FORMUL   5   NA    4(NA 1+)
FORMUL  12  HOH   *510(H2 O)
HELIX    1 AA1 THR A   43  ASP A   50  1                                   8
HELIX    2 AA2 PRO A   52  GLY A   61  1                                  10
HELIX    3 AA3 GLY A   90  GLY A  107  1                                  18
HELIX    4 AA4 SER A  119  TYR A  132  1                                  14
HELIX    5 AA5 PRO A  133  PHE A  136  5                                   4
HELIX    6 AA6 ASN A  161  GLN A  166  1                                   6
HELIX    7 AA7 THR A  172  MET A  183  1                                  12
HELIX    8 AA8 PRO A  207  GLY A  222  1                                  16
HELIX    9 AA9 HIS A  264  GLY A  273  1                                  10
HELIX   10 AB1 THR B   43  ASP B   50  1                                   8
HELIX   11 AB2 PRO B   52  GLY B   61  1                                  10
HELIX   12 AB3 GLY B   90  GLY B  107  1                                  18
HELIX   13 AB4 SER B  119  TYR B  132  1                                  14
HELIX   14 AB5 PRO B  133  PHE B  136  5                                   4
HELIX   15 AB6 ASN B  161  GLN B  166  1                                   6
HELIX   16 AB7 THR B  172  ALA B  182  1                                  11
HELIX   17 AB8 MET B  183  SER B  185  5                                   3
HELIX   18 AB9 PRO B  207  GLY B  222  1                                  16
HELIX   19 AC1 HIS B  264  GLY B  273  1                                  10
SHEET    1 AA110 SER A   3  VAL A   7  0
SHEET    2 AA110 GLY A  18  VAL A  23 -1  O  ILE A  21   N  GLN A   5
SHEET    3 AA110 ILE A  63  SER A  68 -1  O  TYR A  66   N  TYR A  20
SHEET    4 AA110 ILE A  33  ILE A  37  1  N  ALA A  36   O  ILE A  65
SHEET    5 AA110 VAL A 113  SER A 118  1  O  THR A 116   N  ILE A  37
SHEET    6 AA110 ALA A 138  TYR A 142  1  O  TYR A 142   N  GLY A 117
SHEET    7 AA110 ARG A 193  GLY A 199  1  O  TYR A 197   N  VAL A 141
SHEET    8 AA110 LEU A 249  ALA A 254  1  O  ILE A 252   N  ILE A 196
SHEET    9 AA110 TYR A 240  GLY A 246 -1  N  THR A 243   O  GLY A 251
SHEET   10 AA110 LYS A 230  GLU A 232 -1  N  GLU A 232   O  THR A 242
SHEET    1 AA210 SER B   3  VAL B   7  0
SHEET    2 AA210 GLY B  18  VAL B  23 -1  O  MET B  19   N  VAL B   7
SHEET    3 AA210 ILE B  63  SER B  68 -1  O  TYR B  66   N  TYR B  20
SHEET    4 AA210 ILE B  33  ILE B  37  1  N  VAL B  34   O  ILE B  65
SHEET    5 AA210 VAL B 113  SER B 118  1  O  PHE B 114   N  VAL B  35
SHEET    6 AA210 ALA B 138  TYR B 142  1  O  TYR B 142   N  GLY B 117
SHEET    7 AA210 ARG B 193  GLY B 199  1  O  GLN B 195   N  VAL B 141
SHEET    8 AA210 LEU B 249  ALA B 254  1  O  ILE B 252   N  ILE B 196
SHEET    9 AA210 TYR B 240  GLY B 246 -1  N  THR B 243   O  GLY B 251
SHEET   10 AA210 LYS B 230  GLU B 232 -1  N  GLU B 232   O  THR B 242
SSBOND   1 CYS A   40    CYS A   75                          1555   1555  2.01
SSBOND   2 CYS A  149    CYS A  164                          1555   1555  2.02
SSBOND   3 CYS B   40    CYS B   75                          1555   1555  2.04
SSBOND   4 CYS B  149    CYS B  164                          1555   1555  2.02
LINK         OD1 ASN A  30                NA    NA A 304     1555   1555  2.39
LINK         O   GLU A 134                NA    NA A 305     1555   1555  2.94
LINK         ND2 ASN A 161                 C1  NAG A 301     1555   1555  1.44
LINK         OD1 ASN B  30                NA    NA B 306     1555   1555  2.32
LINK         O   TYR B  48                NA    NA B 305     1555   1555  2.79
LINK         O   SER B  51                NA    NA B 305     1555   1555  3.08
LINK         OG  SER B  51                NA    NA B 305     1555   1555  2.82
LINK         OH  TYR B  66                NA    NA B 305     1555   1555  2.72
LINK         O   GLU B 134                NA    NA A 305     1555   1555  2.59
LINK         ND2 ASN B 161                 C1  NAG B 301     1555   1555  1.35
LINK         O4  NAG A 301                 C1  NAG A 302     1555   1555  1.42
LINK        NA    NA A 304                 O   HOH A 488     1555   1555  2.55
LINK        NA    NA A 304                 O   HOH B 588     1555   1555  2.53
LINK        NA    NA A 304                 O   HOH B 433     1555   1555  2.24
LINK        NA    NA A 304                 O   HOH A 604     1555   1555  2.37
LINK        NA    NA A 305                 O   HOH B 551     1555   1555  2.40
LINK        NA    NA A 305                 O   HOH A 607     1555   1555  2.00
LINK         O4  NAG B 301                 C1  NAG B 302     1555   1555  1.41
LINK        NA    NA B 306                 O   HOH A 445     1555   1555  2.38
LINK        NA    NA B 306                 O   HOH A 570     1555   1555  2.60
LINK        NA    NA B 306                 O   HOH A 627     1555   1555  2.50
LINK        NA    NA B 306                 O   HOH B 524     1555   1555  2.31
LINK        NA    NA B 306                 O   HOH B 575     1555   1555  2.53
CISPEP   1 THR A  235    PRO A  236          0         5.40
CISPEP   2 HIS A  259    THR A  260          0        -3.13
CISPEP   3 THR B  235    PRO B  236          0         5.30
CISPEP   4 HIS B  259    THR B  260          0         3.91
SITE     1 AC1 10 TYR A  48  GLY A  49  SER A  51  THR A  55
SITE     2 AC1 10 HOH A 403  HOH A 427  HOH A 439  HOH A 532
SITE     3 AC1 10 GLN B   6  TYR B  20
SITE     1 AC2  6 ASN A  30  HOH A 488  HOH A 604  ASP B  12
SITE     2 AC2  6 HOH B 433  HOH B 588
SITE     1 AC3  4 GLU A 134  HOH A 607  GLU B 134  HOH B 551
SITE     1 AC4  8 GLN A   6  TYR A  20  TYR B  48  GLY B  49
SITE     2 AC4  8 SER B  51  THR B  55  HOH B 461  HOH B 539
SITE     1 AC5 11 GLU A 232  ALA A 233  THR A 235  HOH A 440
SITE     2 AC5 11 GLN B 166  GLY B 167  ASP B 168  TYR B 206
SITE     3 AC5 11 PRO B 207  GLN B 208  HOH B 414
SITE     1 AC6  5 TYR B  48  SER B  51  TYR B  53  ALA B  54
SITE     2 AC6  5 TYR B  66
SITE     1 AC7  6 HOH A 445  HOH A 570  HOH A 627  ASN B  30
SITE     2 AC7  6 HOH B 524  HOH B 575
SITE     1 AC8 10 CYS A 149  TYR A 151  SER A 152  ASN A 153
SITE     2 AC8 10 ASN A 161  VAL A 169  HIS A 175  ILE A 179
SITE     3 AC8 10 HOH A 519  HOH A 600
SITE     1 AC9 14 CYS B 149  TYR B 151  SER B 152  ASN B 153
SITE     2 AC9 14 ASN B 161  VAL B 169  HIS B 175  ILE B 179
SITE     3 AC9 14 HOH B 401  HOH B 426  HOH B 499  HOH B 531
SITE     4 AC9 14 HOH B 535  HOH B 572
CRYST1  174.151  174.151   50.708  90.00  90.00 120.00 H 3          18
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.005742  0.003315  0.000000        0.00000
SCALE2      0.000000  0.006630  0.000000        0.00000
SCALE3      0.000000  0.000000  0.019721        0.00000
TER    2075      ALA A 275
TER    4150      ALA B 275
MASTER      408    0   11   19   20    0   22    6 4754    2  127   44
END