longtext: 6cof-pdb

content
HEADER    LYASE                                   12-MAR-18   6COF
TITLE     ATHNL ENANTIOSELECTIVITY MUTANT AT-A9-H7 APO, Y13C,Y121L,P126F,L128W,
TITLE    2 C131T,A209I
CAVEAT     6COF    RESIDUE A VAL 52 AND RESIDUE A GLU 53 THAT ARE NEXT TO EACH
CAVEAT   2 6COF    OTHER IN THE SAMPLE SEQUENCE ARE NOT PROPERLY LINKED:
CAVEAT   3 6COF    DISTANCE BETWEEN C AND N IS 1.20
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ALPHA-HYDROXYNITRILE LYASE;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: ATHNL;
COMPND   5 SYNONYM: ATHNL,(R)-HYDROXYNITRILE LYASE,(R)-OXYNITRILASE,
COMPND   6 METHYLESTERASE 5,ATMES5;
COMPND   7 EC: 4.1.2.10;
COMPND   8 ENGINEERED: YES;
COMPND   9 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: ARABIDOPSIS THALIANA;
SOURCE   3 ORGANISM_COMMON: MOUSE-EAR CRESS;
SOURCE   4 ORGANISM_TAXID: 3702;
SOURCE   5 GENE: HNL, MES5, AT5G10300, F18D22_70;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PET21A
KEYWDS    ALPHA BETA HYDROLASE GLOBULAR HYDROXYNITRILE LYASE, LYASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    B.J.JONES,R.J.KAZLAUSKAS,R.DESROULEAUX
REVDAT   1   20-MAR-19 6COF    0
JRNL        AUTH   B.J.JONES,R.DESROULEAUX,R.J.KAZLAUSKAS
JRNL        TITL   ATHNL ENANTIOSELECTIVITY MUTANTS
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    1.52 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0155
REMARK   3   AUTHORS     : MURSHUDOV,SKUBAK,LEBEDEV,PANNU,STEINER,
REMARK   3               : NICHOLLS,WINN,LONG,VAGIN
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.52
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 50.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   COMPLETENESS FOR RANGE        (%) : 98.9
REMARK   3   NUMBER OF REFLECTIONS             : 78115
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.160
REMARK   3   R VALUE            (WORKING SET) : 0.158
REMARK   3   FREE R VALUE                     : 0.191
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.100
REMARK   3   FREE R VALUE TEST SET COUNT      : 4180
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.52
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.56
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 5644
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 97.53
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2430
REMARK   3   BIN FREE R VALUE SET COUNT          : 279
REMARK   3   BIN FREE R VALUE                    : 0.2480
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4122
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 14
REMARK   3   SOLVENT ATOMS            : 359
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 24.39
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.34000
REMARK   3    B22 (A**2) : -0.32000
REMARK   3    B33 (A**2) : -0.03000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.068
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.072
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.050
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 2.751
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.974
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.960
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  4340 ; 0.034 ; 0.019
REMARK   3   BOND LENGTHS OTHERS               (A):  4129 ; 0.002 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  5874 ; 2.849 ; 1.961
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  9567 ; 1.292 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   540 ; 6.574 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   193 ;35.155 ;24.301
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   780 ;14.306 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    18 ;20.804 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   634 ; 0.179 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  4861 ; 0.015 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):   977 ; 0.002 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 2
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A     1        A   259
REMARK   3    ORIGIN FOR THE GROUP (A):  -7.8993  -2.6605   3.3449
REMARK   3    T TENSOR
REMARK   3      T11:   0.0119 T22:   0.0562
REMARK   3      T33:   0.0140 T12:  -0.0014
REMARK   3      T13:  -0.0076 T23:   0.0173
REMARK   3    L TENSOR
REMARK   3      L11:   0.5146 L22:   0.1740
REMARK   3      L33:   0.5934 L12:   0.0898
REMARK   3      L13:   0.0024 L23:   0.1062
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0534 S12:   0.1293 S13:   0.0659
REMARK   3      S21:  -0.0116 S22:   0.0219 S23:   0.0006
REMARK   3      S31:  -0.0455 S32:  -0.0640 S33:   0.0315
REMARK   3
REMARK   3   TLS GROUP : 2
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   B     1        B   259
REMARK   3    ORIGIN FOR THE GROUP (A):   9.4819  -7.4920  33.2045
REMARK   3    T TENSOR
REMARK   3      T11:   0.0028 T22:   0.0446
REMARK   3      T33:   0.0222 T12:  -0.0031
REMARK   3      T13:  -0.0041 T23:  -0.0185
REMARK   3    L TENSOR
REMARK   3      L11:   0.6818 L22:   0.2883
REMARK   3      L33:   0.1278 L12:   0.0544
REMARK   3      L13:   0.1973 L23:   0.0354
REMARK   3    S TENSOR
REMARK   3      S11:   0.0157 S12:  -0.1123 S13:   0.0061
REMARK   3      S21:   0.0100 S22:  -0.0024 S23:  -0.0197
REMARK   3      S31:   0.0024 S32:   0.0033 S33:  -0.0134
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : NULL
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS U VALUES : WITH TLS ADDED
REMARK   4
REMARK   4 6COF COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 13-MAR-18.
REMARK 100 THE DEPOSITION ID IS D_1000231846.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 27-OCT-16
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.4
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : APS
REMARK 200  BEAMLINE                       : 19-ID
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97741
REMARK 200  MONOCHROMATOR                  : SI (111) ROSENBAUM-ROCK DOUBLE
REMARK 200                                   -CRYSTAL MONOCHROMATOR
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 82355
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.520
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.0
REMARK 200  DATA REDUNDANCY                : 5.300
REMARK 200  R MERGE                    (I) : 0.06900
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 6.4000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.52
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.55
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 5.20
REMARK 200  R MERGE FOR SHELL          (I) : 0.83700
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: REFMAC
REMARK 200 STARTING MODEL: PDBID 3DQZ
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 45.94
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.28
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1 M BIS-TRIS, 17% PEG 3350, PH 6.4,
REMARK 280  VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 292K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       24.92200
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       61.55950
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       43.54500
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       61.55950
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       24.92200
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       43.54500
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 2550 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 19580 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -41.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     LEU A   260
REMARK 465     LEU B   260
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    HOH A   563     O    HOH A   566              1.83
REMARK 500   NZ   LYS A   209     O    HOH A   401              2.00
REMARK 500   OG   SER A    68     O    HOH A   402              2.04
REMARK 500   OE2  GLU B    57     O    HOH B   401              2.07
REMARK 500   OG   SER B    68     O    HOH B   402              2.10
REMARK 500   O    HOH A   440     O    HOH A   501              2.13
REMARK 500   OE1  GLN B   244     O    HOH B   403              2.14
REMARK 500   O    HOH A   558     O    HOH B   574              2.15
REMARK 500   O    HOH A   535     O    HOH A   550              2.18
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),1X,F6.3)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION
REMARK 500    GLU A  38   CD    GLU A  38   OE2    -0.087
REMARK 500    ARG A  47   C     PRO A  48   N       0.146
REMARK 500    VAL A  52   C     GLU A  53   N      -0.139
REMARK 500    SER A  59   CB    SER A  59   OG     -0.085
REMARK 500    SER A 136   CB    SER A 136   OG     -0.122
REMARK 500    GLU A 168   CD    GLU A 168   OE2     0.115
REMARK 500    GLU A 193   CG    GLU A 193   CD      0.097
REMARK 500    ASP A 221   CB    ASP A 221   CG     -0.139
REMARK 500    LYS B  22   CD    LYS B  22   CE     -0.183
REMARK 500    GLU B  38   CD    GLU B  38   OE1    -0.082
REMARK 500    ARG B  47   CZ    ARG B  47   NH1     0.093
REMARK 500    GLU B  53   CD    GLU B  53   OE1     0.073
REMARK 500    GLU B  53   CD    GLU B  53   OE2    -0.100
REMARK 500    SER B 135   CB    SER B 135   OG     -0.080
REMARK 500    GLU B 138   CD    GLU B 138   OE1    -0.068
REMARK 500    GLU B 138   CD    GLU B 138   OE2     0.073
REMARK 500    TYR B 159   CZ    TYR B 159   CE2    -0.078
REMARK 500    GLU B 165   CD    GLU B 165   OE2     0.078
REMARK 500    GLU B 168   CD    GLU B 168   OE2     0.102
REMARK 500    GLU B 182   CG    GLU B 182   CD      0.101
REMARK 500    GLU B 182   CD    GLU B 182   OE1     0.102
REMARK 500    GLU B 207   CD    GLU B 207   OE2     0.090
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    GLU A  38   CG  -  CD  -  OE1 ANGL. DEV. = -12.1 DEGREES
REMARK 500    ASP A  45   CA  -  C   -  O   ANGL. DEV. =  13.8 DEGREES
REMARK 500    ARG A  47   NE  -  CZ  -  NH1 ANGL. DEV. =  -4.8 DEGREES
REMARK 500    ARG A  47   NE  -  CZ  -  NH1 ANGL. DEV. =   4.3 DEGREES
REMARK 500    ARG A  47   NE  -  CZ  -  NH2 ANGL. DEV. =  -5.3 DEGREES
REMARK 500    PRO A  48   C   -  N   -  CA  ANGL. DEV. =   9.5 DEGREES
REMARK 500    PHE A  82   CB  -  CG  -  CD2 ANGL. DEV. =   4.2 DEGREES
REMARK 500    ASP A  92   CB  -  CG  -  OD2 ANGL. DEV. =  -5.9 DEGREES
REMARK 500    ASP A 110   CB  -  CG  -  OD1 ANGL. DEV. =   5.5 DEGREES
REMARK 500    MET A 125   CG  -  SD  -  CE  ANGL. DEV. = -10.0 DEGREES
REMARK 500    SER A 136   CB  -  CA  -  C   ANGL. DEV. = -15.0 DEGREES
REMARK 500    ARG A 157   NE  -  CZ  -  NH1 ANGL. DEV. =  -3.0 DEGREES
REMARK 500    ARG A 175   NE  -  CZ  -  NH2 ANGL. DEV. =  -4.2 DEGREES
REMARK 500    PHE A 179   CB  -  CG  -  CD1 ANGL. DEV. =   4.4 DEGREES
REMARK 500    ASP A 183   CB  -  CG  -  OD2 ANGL. DEV. =   6.6 DEGREES
REMARK 500    ARG A 200   NE  -  CZ  -  NH1 ANGL. DEV. =   5.3 DEGREES
REMARK 500    ASP A 232   CB  -  CG  -  OD1 ANGL. DEV. =   5.6 DEGREES
REMARK 500    ARG B   3   NE  -  CZ  -  NH1 ANGL. DEV. =   3.4 DEGREES
REMARK 500    GLU B  38   OE1 -  CD  -  OE2 ANGL. DEV. =  13.0 DEGREES
REMARK 500    ASP B  56   CB  -  CG  -  OD2 ANGL. DEV. =  -6.0 DEGREES
REMARK 500    PHE B  80   CB  -  CG  -  CD2 ANGL. DEV. =   5.1 DEGREES
REMARK 500    MET B 123   CG  -  SD  -  CE  ANGL. DEV. =  10.3 DEGREES
REMARK 500    GLU B 138   OE1 -  CD  -  OE2 ANGL. DEV. =   7.4 DEGREES
REMARK 500    MET B 144   CG  -  SD  -  CE  ANGL. DEV. = -16.1 DEGREES
REMARK 500    ARG B 157   NE  -  CZ  -  NH1 ANGL. DEV. =   4.5 DEGREES
REMARK 500    ARG B 175   NE  -  CZ  -  NH1 ANGL. DEV. =   3.6 DEGREES
REMARK 500    ARG B 175   NE  -  CZ  -  NH2 ANGL. DEV. =  -5.5 DEGREES
REMARK 500    ARG B 200   NE  -  CZ  -  NH2 ANGL. DEV. =  -5.6 DEGREES
REMARK 500    CYS B 213   CB  -  CA  -  C   ANGL. DEV. =   7.8 DEGREES
REMARK 500    CYS B 213   CA  -  CB  -  SG  ANGL. DEV. =   7.5 DEGREES
REMARK 500    ASP B 232   CB  -  CG  -  OD1 ANGL. DEV. =   6.8 DEGREES
REMARK 500    ASP B 232   CB  -  CG  -  OD2 ANGL. DEV. =  -8.5 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    HIS A  15     -161.53   -126.55
REMARK 500    SER A  81     -117.21     60.57
REMARK 500    HIS B  15     -164.43   -128.21
REMARK 500    SER B  81     -119.74     59.50
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: MAIN CHAIN PLANARITY
REMARK 500
REMARK 500 THE FOLLOWING RESIDUES HAVE A PSEUDO PLANARITY
REMARK 500 TORSION ANGLE, C(I) - CA(I) - N(I+1) - O(I), GREATER
REMARK 500 10.0 DEGREES. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 500 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 500 I=INSERTION CODE).
REMARK 500
REMARK 500  M RES CSSEQI        ANGLE
REMARK 500    ILE A  44         10.57
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL A 301
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL A 302
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue CL A 303
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue CL B 301
DBREF  6COF A    1   258  UNP    Q9LFT6   HNL_ARATH        1    258
DBREF  6COF B    1   258  UNP    Q9LFT6   HNL_ARATH        1    258
SEQADV 6COF CYS A   14  UNP  Q9LFT6    TYR    14 ENGINEERED MUTATION
SEQADV 6COF LEU A  122  UNP  Q9LFT6    TYR   122 ENGINEERED MUTATION
SEQADV 6COF PHE A  126  UNP  Q9LFT6    PRO   126 ENGINEERED MUTATION
SEQADV 6COF TRP A  129  UNP  Q9LFT6    LEU   129 ENGINEERED MUTATION
SEQADV 6COF THR A  132  UNP  Q9LFT6    CYS   132 ENGINEERED MUTATION
SEQADV 6COF ILE A  210  UNP  Q9LFT6    ALA   210 ENGINEERED MUTATION
SEQADV 6COF GLY A  259  UNP  Q9LFT6              EXPRESSION TAG
SEQADV 6COF LEU A  260  UNP  Q9LFT6              EXPRESSION TAG
SEQADV 6COF CYS B   14  UNP  Q9LFT6    TYR    14 ENGINEERED MUTATION
SEQADV 6COF LEU B  122  UNP  Q9LFT6    TYR   122 ENGINEERED MUTATION
SEQADV 6COF PHE B  126  UNP  Q9LFT6    PRO   126 ENGINEERED MUTATION
SEQADV 6COF TRP B  129  UNP  Q9LFT6    LEU   129 ENGINEERED MUTATION
SEQADV 6COF THR B  132  UNP  Q9LFT6    CYS   132 ENGINEERED MUTATION
SEQADV 6COF ILE B  210  UNP  Q9LFT6    ALA   210 ENGINEERED MUTATION
SEQADV 6COF GLY B  259  UNP  Q9LFT6              EXPRESSION TAG
SEQADV 6COF LEU B  260  UNP  Q9LFT6              EXPRESSION TAG
SEQRES   1 A  260  MET GLU ARG LYS HIS HIS PHE VAL LEU VAL HIS ASN ALA
SEQRES   2 A  260  CYS HIS GLY ALA TRP ILE TRP TYR LYS LEU LYS PRO LEU
SEQRES   3 A  260  LEU GLU SER ALA GLY HIS ARG VAL THR ALA VAL GLU LEU
SEQRES   4 A  260  ALA ALA SER GLY ILE ASP PRO ARG PRO ILE GLN ALA VAL
SEQRES   5 A  260  GLU THR VAL ASP GLU TYR SER LYS PRO LEU ILE GLU THR
SEQRES   6 A  260  LEU LYS SER LEU PRO GLU ASN GLU GLU VAL ILE LEU VAL
SEQRES   7 A  260  GLY PHE SER PHE GLY GLY ILE ASN ILE ALA LEU ALA ALA
SEQRES   8 A  260  ASP ILE PHE PRO ALA LYS ILE LYS VAL LEU VAL PHE LEU
SEQRES   9 A  260  ASN ALA PHE LEU PRO ASP THR THR HIS VAL PRO SER HIS
SEQRES  10 A  260  VAL LEU ASP LYS LEU MET GLU MET PHE GLY GLY TRP GLY
SEQRES  11 A  260  ASP THR GLU PHE SER SER HIS GLU THR ARG ASN GLY THR
SEQRES  12 A  260  MET SER LEU LEU LYS MET GLY PRO LYS PHE MET LYS ALA
SEQRES  13 A  260  ARG LEU TYR GLN ASN CYS PRO ILE GLU ASP TYR GLU LEU
SEQRES  14 A  260  ALA LYS MET LEU HIS ARG GLN GLY SER PHE PHE THR GLU
SEQRES  15 A  260  ASP LEU SER LYS LYS GLU LYS PHE SER GLU GLU GLY TYR
SEQRES  16 A  260  GLY SER VAL GLN ARG VAL TYR VAL MET SER SER GLU ASP
SEQRES  17 A  260  LYS ILE ILE PRO CYS ASP PHE ILE ARG TRP MET ILE ASP
SEQRES  18 A  260  ASN PHE ASN VAL SER LYS VAL TYR GLU ILE ASP GLY GLY
SEQRES  19 A  260  ASP HIS MET VAL MET LEU SER LYS PRO GLN LYS LEU PHE
SEQRES  20 A  260  ASP SER LEU SER ALA ILE ALA THR ASP TYR MET GLY LEU
SEQRES   1 B  260  MET GLU ARG LYS HIS HIS PHE VAL LEU VAL HIS ASN ALA
SEQRES   2 B  260  CYS HIS GLY ALA TRP ILE TRP TYR LYS LEU LYS PRO LEU
SEQRES   3 B  260  LEU GLU SER ALA GLY HIS ARG VAL THR ALA VAL GLU LEU
SEQRES   4 B  260  ALA ALA SER GLY ILE ASP PRO ARG PRO ILE GLN ALA VAL
SEQRES   5 B  260  GLU THR VAL ASP GLU TYR SER LYS PRO LEU ILE GLU THR
SEQRES   6 B  260  LEU LYS SER LEU PRO GLU ASN GLU GLU VAL ILE LEU VAL
SEQRES   7 B  260  GLY PHE SER PHE GLY GLY ILE ASN ILE ALA LEU ALA ALA
SEQRES   8 B  260  ASP ILE PHE PRO ALA LYS ILE LYS VAL LEU VAL PHE LEU
SEQRES   9 B  260  ASN ALA PHE LEU PRO ASP THR THR HIS VAL PRO SER HIS
SEQRES  10 B  260  VAL LEU ASP LYS LEU MET GLU MET PHE GLY GLY TRP GLY
SEQRES  11 B  260  ASP THR GLU PHE SER SER HIS GLU THR ARG ASN GLY THR
SEQRES  12 B  260  MET SER LEU LEU LYS MET GLY PRO LYS PHE MET LYS ALA
SEQRES  13 B  260  ARG LEU TYR GLN ASN CYS PRO ILE GLU ASP TYR GLU LEU
SEQRES  14 B  260  ALA LYS MET LEU HIS ARG GLN GLY SER PHE PHE THR GLU
SEQRES  15 B  260  ASP LEU SER LYS LYS GLU LYS PHE SER GLU GLU GLY TYR
SEQRES  16 B  260  GLY SER VAL GLN ARG VAL TYR VAL MET SER SER GLU ASP
SEQRES  17 B  260  LYS ILE ILE PRO CYS ASP PHE ILE ARG TRP MET ILE ASP
SEQRES  18 B  260  ASN PHE ASN VAL SER LYS VAL TYR GLU ILE ASP GLY GLY
SEQRES  19 B  260  ASP HIS MET VAL MET LEU SER LYS PRO GLN LYS LEU PHE
SEQRES  20 B  260  ASP SER LEU SER ALA ILE ALA THR ASP TYR MET GLY LEU
HET    GOL  A 301       6
HET    GOL  A 302       6
HET     CL  A 303       1
HET     CL  B 301       1
HETNAM     GOL GLYCEROL
HETNAM      CL CHLORIDE ION
HETSYN     GOL GLYCERIN; PROPANE-1,2,3-TRIOL
FORMUL   3  GOL    2(C3 H8 O3)
FORMUL   5   CL    2(CL 1-)
FORMUL   7  HOH   *359(H2 O)
HELIX    1 AA1 GLY A   16  TYR A   21  5                                   6
HELIX    2 AA2 LYS A   22  ALA A   30  1                                   9
HELIX    3 AA3 PRO A   48  VAL A   52  5                                   5
HELIX    4 AA4 THR A   54  SER A   68  1                                  15
HELIX    5 AA5 PHE A   82  PHE A   94  1                                  13
HELIX    6 AA6 SER A  116  PHE A  126  1                                  11
HELIX    7 AA7 GLY A  150  LEU A  158  1                                   9
HELIX    8 AA8 PRO A  163  HIS A  174  1                                  12
HELIX    9 AA9 PHE A  180  LYS A  187  1                                   8
HELIX   10 AB1 GLY A  194  VAL A  198  5                                   5
HELIX   11 AB2 PRO A  212  PHE A  223  1                                  12
HELIX   12 AB3 MET A  237  LYS A  242  1                                   6
HELIX   13 AB4 LYS A  242  TYR A  257  1                                  16
HELIX   14 AB5 GLY B   16  TYR B   21  5                                   6
HELIX   15 AB6 LYS B   22  ALA B   30  1                                   9
HELIX   16 AB7 PRO B   48  VAL B   52  5                                   5
HELIX   17 AB8 THR B   54  SER B   68  1                                  15
HELIX   18 AB9 PHE B   82  PHE B   94  1                                  13
HELIX   19 AC1 SER B  116  MET B  125  1                                  10
HELIX   20 AC2 GLY B  150  LEU B  158  1                                   9
HELIX   21 AC3 PRO B  163  HIS B  174  1                                  12
HELIX   22 AC4 PHE B  180  LYS B  186  1                                   7
HELIX   23 AC5 GLY B  194  VAL B  198  5                                   5
HELIX   24 AC6 PRO B  212  PHE B  223  1                                  12
HELIX   25 AC7 MET B  237  LYS B  242  1                                   6
HELIX   26 AC8 LYS B  242  TYR B  257  1                                  16
SHEET    1 AA1 6 ARG A  33  VAL A  37  0
SHEET    2 AA1 6 HIS A   6  VAL A  10  1  N  LEU A   9   O  THR A  35
SHEET    3 AA1 6 VAL A  75  PHE A  80  1  O  VAL A  78   N  VAL A   8
SHEET    4 AA1 6 ILE A  98  LEU A 104  1  O  VAL A 102   N  LEU A  77
SHEET    5 AA1 6 ARG A 200  SER A 205  1  O  VAL A 203   N  PHE A 103
SHEET    6 AA1 6 VAL A 228  ILE A 231  1  O  TYR A 229   N  MET A 204
SHEET    1 AA2 2 GLU A 133  THR A 139  0
SHEET    2 AA2 2 GLY A 142  LYS A 148 -1  O  LYS A 148   N  GLU A 133
SHEET    1 AA3 6 ARG B  33  ALA B  36  0
SHEET    2 AA3 6 HIS B   6  VAL B  10  1  N  LEU B   9   O  THR B  35
SHEET    3 AA3 6 VAL B  75  PHE B  80  1  O  VAL B  78   N  VAL B   8
SHEET    4 AA3 6 ILE B  98  LEU B 104  1  O  VAL B 102   N  LEU B  77
SHEET    5 AA3 6 ARG B 200  SER B 205  1  O  VAL B 203   N  PHE B 103
SHEET    6 AA3 6 VAL B 228  ILE B 231  1  O  ILE B 231   N  MET B 204
SHEET    1 AA4 2 GLU B 133  THR B 139  0
SHEET    2 AA4 2 GLY B 142  LYS B 148 -1  O  LEU B 146   N  SER B 135
SITE     1 AC1  8 GLU A  53  ASN A 141  GLY A 142  MET A 144
SITE     2 AC1  8 ASP A 183  LYS A 186  HOH A 455  HOH A 476
SITE     1 AC2  6 GLY A 128  GLY A 130  LYS A 152  PHE A 153
SITE     2 AC2  6 ARG A 157  HOH A 464
SITE     1 AC3  5 ASN A  12  ALA A  13  SER A  81  PHE A  82
SITE     2 AC3  5 PHE A 179
SITE     1 AC4  6 ASN B  12  ALA B  13  SER B  81  PHE B  82
SITE     2 AC4  6 PHE B 179  HOH B 531
CRYST1   49.844   87.090  123.119  90.00  90.00  90.00 P 21 21 21    8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.020063  0.000000  0.000000        0.00000
SCALE2      0.000000  0.011482  0.000000        0.00000
SCALE3      0.000000  0.000000  0.008122        0.00000
TER    2104      GLY A 259
TER    4216      GLY B 259
MASTER      445    0    4   26   16    0    8    6 4495    2   12   40
END