longtext: 6fr2-pdb

content
HEADER    HYDROLASE                               15-FEB-18   6FR2
TITLE     SOLUBLE EPOXIDE HYDROLASE IN COMPLEX WITH LK864
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: BIFUNCTIONAL EPOXIDE HYDROLASE 2;
COMPND   3 CHAIN: A;
COMPND   4 EC: 3.3.2.10,3.1.3.76;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: EPHX2;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 469008
KEYWDS    INHIBITOR, COMPLEX, SEH, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    J.S.KRAMER,D.POGORYELOV,M.KRASAVIN,E.PROSCHAK
REVDAT   1   12-SEP-18 6FR2    0
JRNL        AUTH   A.LUKIN,J.KRAMER,M.HARTMANN,L.WEIZEL,V.HERNANDEZ-OLMOS,
JRNL        AUTH 2 K.FALAHATI,I.BURGHARDT,N.KALINCHENKOVA,D.BAGNYUKOVA,
JRNL        AUTH 3 N.ZHURILO,J.RAUTIO,M.FORSBERG,J.IHALAINEN,S.AURIOLA,
JRNL        AUTH 4 J.LEPPANEN,I.KONSTANTINOV,D.POGORYELOV,E.PROSCHAK,D.DAR'IN,
JRNL        AUTH 5 M.KRASAVIN
JRNL        TITL   DISCOVERY OF POLAR SPIROCYCLIC ORALLY BIOAVAILABLE UREA
JRNL        TITL 2 INHIBITORS OF SOLUBLE EPOXIDE HYDROLASE.
JRNL        REF    BIOORG. CHEM.                 V.  80   655 2018
JRNL        REFN                   ISSN 1090-2120
JRNL        PMID   30059891
JRNL        DOI    10.1016/J.BIOORG.2018.07.014
REMARK   2
REMARK   2 RESOLUTION.    2.26 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.12_2829
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.26
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 46.09
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.340
REMARK   3   COMPLETENESS FOR RANGE        (%) : 98.6
REMARK   3   NUMBER OF REFLECTIONS             : 35259
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.170
REMARK   3   R VALUE            (WORKING SET) : 0.168
REMARK   3   FREE R VALUE                     : 0.210
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.040
REMARK   3   FREE R VALUE TEST SET COUNT      : 1776
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 46.0996 -  5.3152    0.99     2593   133  0.1642 0.2014
REMARK   3     2  5.3152 -  4.2197    0.99     2560   179  0.1296 0.1677
REMARK   3     3  4.2197 -  3.6866    0.98     2534   166  0.1365 0.1661
REMARK   3     4  3.6866 -  3.3496    0.99     2597   146  0.1459 0.2065
REMARK   3     5  3.3496 -  3.1096    1.00     2597   130  0.1678 0.2036
REMARK   3     6  3.1096 -  2.9263    1.00     2609   155  0.1757 0.3036
REMARK   3     7  2.9263 -  2.7798    1.00     2619    96  0.1883 0.2219
REMARK   3     8  2.7798 -  2.6588    0.99     2607   133  0.2020 0.2674
REMARK   3     9  2.6588 -  2.5564    1.00     2606   120  0.2060 0.2516
REMARK   3    10  2.5564 -  2.4682    1.00     2631   132  0.2296 0.2516
REMARK   3    11  2.4682 -  2.3911    1.00     2561   140  0.2432 0.3240
REMARK   3    12  2.3911 -  2.3227    1.00     2627   129  0.2653 0.2859
REMARK   3    13  2.3227 -  2.2616    0.90     2342   117  0.3071 0.3418
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : NULL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.250
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 24.190
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 46.28
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 54.68
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.011           2710
REMARK   3   ANGLE     :  0.955           3688
REMARK   3   CHIRALITY :  0.060            379
REMARK   3   PLANARITY :  0.007            474
REMARK   3   DIHEDRAL  : 25.429           1010
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 9
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 237 THROUGH 292 )
REMARK   3    ORIGIN FOR THE GROUP (A):  -6.6249  -7.1319 -34.1435
REMARK   3    T TENSOR
REMARK   3      T11:   0.2202 T22:   0.2227
REMARK   3      T33:   0.2385 T12:  -0.0026
REMARK   3      T13:   0.0255 T23:  -0.0017
REMARK   3    L TENSOR
REMARK   3      L11:   0.4883 L22:   0.7117
REMARK   3      L33:   0.3092 L12:   0.7476
REMARK   3      L13:  -0.0650 L23:  -0.1065
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0217 S12:  -0.0116 S13:   0.0528
REMARK   3      S21:  -0.0291 S22:   0.0155 S23:  -0.0579
REMARK   3      S31:   0.0330 S32:  -0.0023 S33:   0.0000
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 293 THROUGH 328 )
REMARK   3    ORIGIN FOR THE GROUP (A):   2.2884 -15.7093 -37.4347
REMARK   3    T TENSOR
REMARK   3      T11:   0.3306 T22:   0.3087
REMARK   3      T33:   0.3373 T12:  -0.0127
REMARK   3      T13:   0.0520 T23:  -0.0063
REMARK   3    L TENSOR
REMARK   3      L11:   0.3110 L22:   0.1616
REMARK   3      L33:   0.2562 L12:   0.2036
REMARK   3      L13:  -0.1012 L23:   0.1808
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1446 S12:  -0.0177 S13:  -0.0216
REMARK   3      S21:  -0.0278 S22:   0.0231 S23:  -0.1889
REMARK   3      S31:   0.0115 S32:   0.0685 S33:  -0.0000
REMARK   3   TLS GROUP : 3
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 329 THROUGH 370 )
REMARK   3    ORIGIN FOR THE GROUP (A):  -3.8212 -23.0067 -27.1340
REMARK   3    T TENSOR
REMARK   3      T11:   0.2401 T22:   0.2903
REMARK   3      T33:   0.3705 T12:   0.0034
REMARK   3      T13:  -0.0046 T23:   0.0249
REMARK   3    L TENSOR
REMARK   3      L11:   0.3201 L22:   0.0663
REMARK   3      L33:   0.4912 L12:   0.0507
REMARK   3      L13:  -0.0404 L23:   0.2002
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0614 S12:  -0.2259 S13:  -0.2133
REMARK   3      S21:   0.0215 S22:   0.0303 S23:  -0.2012
REMARK   3      S31:   0.0632 S32:  -0.0537 S33:  -0.0000
REMARK   3   TLS GROUP : 4
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 371 THROUGH 399 )
REMARK   3    ORIGIN FOR THE GROUP (A): -13.3053 -31.3425 -45.1712
REMARK   3    T TENSOR
REMARK   3      T11:   0.5637 T22:   0.3929
REMARK   3      T33:   0.4214 T12:  -0.0869
REMARK   3      T13:   0.0484 T23:  -0.0127
REMARK   3    L TENSOR
REMARK   3      L11:   0.2713 L22:   0.2135
REMARK   3      L33:   0.0449 L12:   0.0821
REMARK   3      L13:   0.0992 L23:   0.0262
REMARK   3    S TENSOR
REMARK   3      S11:  -0.2738 S12:   0.2211 S13:  -0.3432
REMARK   3      S21:  -0.3410 S22:   0.1576 S23:  -0.0663
REMARK   3      S31:   0.3942 S32:  -0.3891 S33:   0.0000
REMARK   3   TLS GROUP : 5
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 400 THROUGH 418 )
REMARK   3    ORIGIN FOR THE GROUP (A): -25.0625 -18.8325 -44.1364
REMARK   3    T TENSOR
REMARK   3      T11:   0.4387 T22:   0.4189
REMARK   3      T33:   0.4011 T12:  -0.0433
REMARK   3      T13:  -0.0736 T23:   0.0516
REMARK   3    L TENSOR
REMARK   3      L11:   0.1158 L22:   0.0579
REMARK   3      L33:   0.0048 L12:   0.0051
REMARK   3      L13:  -0.0561 L23:   0.0629
REMARK   3    S TENSOR
REMARK   3      S11:   0.1155 S12:  -0.1004 S13:  -0.1432
REMARK   3      S21:  -0.1951 S22:   0.0782 S23:   0.4641
REMARK   3      S31:   0.1648 S32:   0.1270 S33:  -0.0000
REMARK   3   TLS GROUP : 6
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 419 THROUGH 435 )
REMARK   3    ORIGIN FOR THE GROUP (A): -24.5846 -28.3296 -48.5732
REMARK   3    T TENSOR
REMARK   3      T11:   0.7719 T22:   0.7208
REMARK   3      T33:   0.7253 T12:  -0.1121
REMARK   3      T13:  -0.1203 T23:  -0.0747
REMARK   3    L TENSOR
REMARK   3      L11:   0.0612 L22:   0.0299
REMARK   3      L33:   0.0693 L12:   0.0340
REMARK   3      L13:  -0.0039 L23:   0.1146
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1248 S12:  -0.0381 S13:  -0.4049
REMARK   3      S21:  -0.3272 S22:   0.1865 S23:   0.0546
REMARK   3      S31:   0.0440 S32:  -0.4081 S33:  -0.0000
REMARK   3   TLS GROUP : 7
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 436 THROUGH 509 )
REMARK   3    ORIGIN FOR THE GROUP (A): -11.4401 -22.3832 -35.1620
REMARK   3    T TENSOR
REMARK   3      T11:   0.2993 T22:   0.2419
REMARK   3      T33:   0.2392 T12:  -0.0135
REMARK   3      T13:   0.0121 T23:   0.0258
REMARK   3    L TENSOR
REMARK   3      L11:   0.9913 L22:   1.1227
REMARK   3      L33:   0.6097 L12:   0.0661
REMARK   3      L13:   0.1301 L23:  -0.1306
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0459 S12:  -0.0544 S13:  -0.1275
REMARK   3      S21:  -0.1108 S22:   0.0134 S23:  -0.0435
REMARK   3      S31:   0.0675 S32:  -0.0676 S33:  -0.0000
REMARK   3   TLS GROUP : 8
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 510 THROUGH 530 )
REMARK   3    ORIGIN FOR THE GROUP (A): -17.9137 -19.8895 -24.4577
REMARK   3    T TENSOR
REMARK   3      T11:   0.3508 T22:   0.4221
REMARK   3      T33:   0.3879 T12:  -0.0270
REMARK   3      T13:   0.0333 T23:   0.0635
REMARK   3    L TENSOR
REMARK   3      L11:   0.3250 L22:   0.4341
REMARK   3      L33:   0.0740 L12:  -0.2097
REMARK   3      L13:  -0.2247 L23:   0.3601
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0034 S12:  -0.2008 S13:   0.0156
REMARK   3      S21:   0.0061 S22:   0.0412 S23:   0.1788
REMARK   3      S31:  -0.0501 S32:  -0.1574 S33:   0.0000
REMARK   3   TLS GROUP : 9
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 531 THROUGH 548 )
REMARK   3    ORIGIN FOR THE GROUP (A): -14.1672 -10.4860 -18.8647
REMARK   3    T TENSOR
REMARK   3      T11:   0.3921 T22:   0.4543
REMARK   3      T33:   0.3113 T12:   0.0210
REMARK   3      T13:   0.0013 T23:  -0.0195
REMARK   3    L TENSOR
REMARK   3      L11:   0.0966 L22:   0.3126
REMARK   3      L33:   0.3010 L12:  -0.0499
REMARK   3      L13:  -0.2172 L23:   0.1357
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1270 S12:  -0.4046 S13:   0.2106
REMARK   3      S21:   0.2733 S22:   0.1594 S23:   0.0719
REMARK   3      S31:  -0.3123 S32:  -0.1852 S33:  -0.0000
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 6FR2 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 16-FEB-18.
REMARK 100 THE DEPOSITION ID IS D_1200008485.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 16-SEP-17
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.0-6.55
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID29
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.072
REMARK 200  MONOCHROMATOR                  : 1.072
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XDS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 35335
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.260
REMARK 200  RESOLUTION RANGE LOW       (A) : 46.090
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.5
REMARK 200  DATA REDUNDANCY                : 3.400
REMARK 200  R MERGE                    (I) : 0.08100
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 9.4000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.26
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.30
REMARK 200  COMPLETENESS FOR SHELL     (%) : 82.7
REMARK 200  DATA REDUNDANCY IN SHELL       : 2.98
REMARK 200  R MERGE FOR SHELL          (I) : 0.78800
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 1.710
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: 4JNC
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 54.39
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.70
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 1 UL PROTEIN SOLUTION PROTEIN (5-10
REMARK 280  MG/ML , 50 MM NACL, 50 MM SODIUM PHOSPHATE, 10% GLYCEROL (98%),
REMARK 280  2 MM DTT AT PH 7.4) WAS MIXED IN DIFFERENT RATIOS (2/1, 1/1, 1/2)
REMARK 280  WITH PRECIPITANT SOLUTION (23 %-28 % (W/V) POLYETHYLENGLYCOL
REMARK 280  (PEG) 6000, 70 MM AMMONIUM ACETAT, 200 MM MAGNESIUM ACETAT, 100
REMARK 280  MM SODIUM CACODYLATE AT PH 6.1-6.5), VAPOR DIFFUSION, HANGING
REMARK 280  DROP, TEMPERATURE 277K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: I 2 2 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z
REMARK 290       3555   -X,Y,-Z
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z+1/2
REMARK 290       6555   -X+1/2,-Y+1/2,Z+1/2
REMARK 290       7555   -X+1/2,Y+1/2,-Z+1/2
REMARK 290       8555   X+1/2,-Y+1/2,-Z+1/2
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       40.04550
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       46.09000
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000       53.49900
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       40.04550
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       46.09000
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       53.49900
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       40.04550
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       46.09000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       53.49900
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       40.04550
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000       46.09000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000       53.49900
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 100 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 12810 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -8.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH A 786  LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A   219
REMARK 465     ALA A   220
REMARK 465     SER A   221
REMARK 465     LEU A   222
REMARK 465     ASN A   223
REMARK 465     THR A   224
REMARK 465     PRO A   225
REMARK 465     ALA A   226
REMARK 465     PRO A   227
REMARK 465     LEU A   228
REMARK 465     ASN A   548
REMARK 465     PRO A   549
REMARK 465     PRO A   550
REMARK 465     VAL A   551
REMARK 465     VAL A   552
REMARK 465     SER A   553
REMARK 465     LYS A   554
REMARK 465     MET A   555
REMARK 465     LEU A   556
REMARK 465     LEU A   557
REMARK 465     GLU A   558
REMARK 465     HIS A   559
REMARK 465     HIS A   560
REMARK 465     HIS A   561
REMARK 465     HIS A   562
REMARK 465     HIS A   563
REMARK 465     HIS A   564
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     LYS A 376    CG   CD   CE   NZ
REMARK 470     LYS A 421    CG   CD   CE   NZ
REMARK 470     GLU A 424    CG   CD   OE1  OE2
REMARK 470     TRP A 510    CG   CD1  CD2  NE1  CE2  CE3  CZ2
REMARK 470     TRP A 510    CZ3  CH2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    SER A 231     -163.54   -109.47
REMARK 500    GLU A 269     -140.87   -119.91
REMARK 500    ASP A 335     -129.92     64.74
REMARK 500    ASN A 359      -43.80     77.09
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 MET A  291     ASP A  292                  147.71
REMARK 500
REMARK 500 REMARK: NULL
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              MG A 602  MG
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASP A 509   OD1
REMARK 620 2 ASP A 509   OD2  57.2
REMARK 620 3 HOH A 802   O    95.7  95.5
REMARK 620 N                    1     2
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue E3N A 601
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue MG A 602
DBREF  6FR2 A  222   555  UNP    P34913   HYES_HUMAN     169    502
SEQADV 6FR2 MET A  219  UNP  P34913              INITIATING METHIONINE
SEQADV 6FR2 ALA A  220  UNP  P34913              EXPRESSION TAG
SEQADV 6FR2 SER A  221  UNP  P34913              EXPRESSION TAG
SEQADV 6FR2 LEU A  556  UNP  P34913              EXPRESSION TAG
SEQADV 6FR2 LEU A  557  UNP  P34913              EXPRESSION TAG
SEQADV 6FR2 GLU A  558  UNP  P34913              EXPRESSION TAG
SEQADV 6FR2 HIS A  559  UNP  P34913              EXPRESSION TAG
SEQADV 6FR2 HIS A  560  UNP  P34913              EXPRESSION TAG
SEQADV 6FR2 HIS A  561  UNP  P34913              EXPRESSION TAG
SEQADV 6FR2 HIS A  562  UNP  P34913              EXPRESSION TAG
SEQADV 6FR2 HIS A  563  UNP  P34913              EXPRESSION TAG
SEQADV 6FR2 HIS A  564  UNP  P34913              EXPRESSION TAG
SEQRES   1 A  346  MET ALA SER LEU ASN THR PRO ALA PRO LEU PRO THR SER
SEQRES   2 A  346  CYS ASN PRO SER ASP MET SER HIS GLY TYR VAL THR VAL
SEQRES   3 A  346  LYS PRO ARG VAL ARG LEU HIS PHE VAL GLU LEU GLY SER
SEQRES   4 A  346  GLY PRO ALA VAL CYS LEU CYS HIS GLY PHE PRO GLU SER
SEQRES   5 A  346  TRP TYR SER TRP ARG TYR GLN ILE PRO ALA LEU ALA GLN
SEQRES   6 A  346  ALA GLY TYR ARG VAL LEU ALA MET ASP MET LYS GLY TYR
SEQRES   7 A  346  GLY GLU SER SER ALA PRO PRO GLU ILE GLU GLU TYR CYS
SEQRES   8 A  346  MET GLU VAL LEU CYS LYS GLU MET VAL THR PHE LEU ASP
SEQRES   9 A  346  LYS LEU GLY LEU SER GLN ALA VAL PHE ILE GLY HIS ASP
SEQRES  10 A  346  TRP GLY GLY MET LEU VAL TRP TYR MET ALA LEU PHE TYR
SEQRES  11 A  346  PRO GLU ARG VAL ARG ALA VAL ALA SER LEU ASN THR PRO
SEQRES  12 A  346  PHE ILE PRO ALA ASN PRO ASN MET SER PRO LEU GLU SER
SEQRES  13 A  346  ILE LYS ALA ASN PRO VAL PHE ASP TYR GLN LEU TYR PHE
SEQRES  14 A  346  GLN GLU PRO GLY VAL ALA GLU ALA GLU LEU GLU GLN ASN
SEQRES  15 A  346  LEU SER ARG THR PHE LYS SER LEU PHE ARG ALA SER ASP
SEQRES  16 A  346  GLU SER VAL LEU SER MET HIS LYS VAL CYS GLU ALA GLY
SEQRES  17 A  346  GLY LEU PHE VAL ASN SER PRO GLU GLU PRO SER LEU SER
SEQRES  18 A  346  ARG MET VAL THR GLU GLU GLU ILE GLN PHE TYR VAL GLN
SEQRES  19 A  346  GLN PHE LYS LYS SER GLY PHE ARG GLY PRO LEU ASN TRP
SEQRES  20 A  346  TYR ARG ASN MET GLU ARG ASN TRP LYS TRP ALA CYS LYS
SEQRES  21 A  346  SER LEU GLY ARG LYS ILE LEU ILE PRO ALA LEU MET VAL
SEQRES  22 A  346  THR ALA GLU LYS ASP PHE VAL LEU VAL PRO GLN MET SER
SEQRES  23 A  346  GLN HIS MET GLU ASP TRP ILE PRO HIS LEU LYS ARG GLY
SEQRES  24 A  346  HIS ILE GLU ASP CYS GLY HIS TRP THR GLN MET ASP LYS
SEQRES  25 A  346  PRO THR GLU VAL ASN GLN ILE LEU ILE LYS TRP LEU ASP
SEQRES  26 A  346  SER ASP ALA ARG ASN PRO PRO VAL VAL SER LYS MET LEU
SEQRES  27 A  346  LEU GLU HIS HIS HIS HIS HIS HIS
HET    E3N  A 601      33
HET     MG  A 602       1
HETNAM     E3N 1-[(4~{S})-9-PROPAN-2-YLSULFONYL-1-OXA-9-
HETNAM   2 E3N  AZASPIRO[5.5]UNDECAN-4-YL]-3-[[4-(TRIFLUOROMETHYLOXY)
HETNAM   3 E3N  PHENYL]METHYL]UREA
HETNAM      MG MAGNESIUM ION
FORMUL   2  E3N    C21 H30 F3 N3 O5 S
FORMUL   3   MG    MG 2+
FORMUL   4  HOH   *107(H2 O)
HELIX    1 AA1 ASN A  233  MET A  237  5                                   5
HELIX    2 AA2 SER A  270  ARG A  275  5                                   6
HELIX    3 AA3 GLN A  277  ALA A  284  1                                   8
HELIX    4 AA4 GLU A  304  TYR A  308  5                                   5
HELIX    5 AA5 CYS A  309  LEU A  324  1                                  16
HELIX    6 AA6 ASP A  335  TYR A  348  1                                  14
HELIX    7 AA7 SER A  370  ALA A  377  1                                   8
HELIX    8 AA8 ASN A  378  PHE A  381  5                                   4
HELIX    9 AA9 ASP A  382  PHE A  387  1                                   6
HELIX   10 AB1 GLY A  391  ASN A  400  1                                  10
HELIX   11 AB2 ASN A  400  PHE A  409  1                                  10
HELIX   12 AB3 ALA A  411  SER A  415  5                                   5
HELIX   13 AB4 SER A  418  HIS A  420  5                                   3
HELIX   14 AB5 LYS A  421  GLY A  426  1                                   6
HELIX   15 AB6 THR A  443  LYS A  455  1                                  13
HELIX   16 AB7 PHE A  459  ASN A  464  1                                   6
HELIX   17 AB8 TRP A  465  ARG A  467  5                                   3
HELIX   18 AB9 ASN A  468  LYS A  478  1                                  11
HELIX   19 AC1 VAL A  500  GLN A  505  5                                   6
HELIX   20 AC2 HIS A  506  TRP A  510  5                                   5
HELIX   21 AC3 TRP A  525  LYS A  530  1                                   6
HELIX   22 AC4 LYS A  530  ALA A  546  1                                  17
SHEET    1 AA1 8 SER A 238  LYS A 245  0
SHEET    2 AA1 8 VAL A 248  LEU A 255 -1  O  PHE A 252   N  GLY A 240
SHEET    3 AA1 8 ARG A 287  MET A 291 -1  O  VAL A 288   N  LEU A 255
SHEET    4 AA1 8 ALA A 260  CYS A 264  1  N  VAL A 261   O  ARG A 287
SHEET    5 AA1 8 ALA A 329  HIS A 334  1  O  VAL A 330   N  CYS A 262
SHEET    6 AA1 8 VAL A 352  LEU A 358  1  O  LEU A 358   N  GLY A 333
SHEET    7 AA1 8 ALA A 488  ALA A 493  1  O  VAL A 491   N  SER A 357
SHEET    8 AA1 8 LEU A 514  ILE A 519  1  O  LYS A 515   N  ALA A 488
LINK         OD1 ASP A 509                MG    MG A 602     1555   1555  2.36
LINK         OD2 ASP A 509                MG    MG A 602     1555   1555  2.24
LINK        MG    MG A 602                 O   HOH A 802     1555   1555  2.65
CISPEP   1 PHE A  267    PRO A  268          0       -10.38
SITE     1 AC1 16 ASP A 335  TRP A 336  MET A 339  ILE A 375
SITE     2 AC1 16 PHE A 381  TYR A 383  GLN A 384  SER A 415
SITE     3 AC1 16 MET A 419  TYR A 466  MET A 469  VAL A 498
SITE     4 AC1 16 HIS A 524  TRP A 525  HOH A 717  HOH A 805
SITE     1 AC2  3 HIS A 506  ASP A 509  HOH A 802
CRYST1   80.091   92.180  106.998  90.00  90.00  90.00 I 2 2 2       8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.012486  0.000000  0.000000        0.00000
SCALE2      0.000000  0.010848  0.000000        0.00000
SCALE3      0.000000  0.000000  0.009346        0.00000
TER    2595      ARG A 547
MASTER      467    0    2   22    8    0    5    6 2689    1   37   27
END