longtext: 6grw-pdb

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HEADER    HYDROLASE                               12-JUN-18   6GRW
TITLE     GLUCURONOYL ESTERASE FROM OPITUTUS TERRAE (AU DERIVATIVE)
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: PUTATIVE ACETYL XYLAN ESTERASE;
COMPND   3 CHAIN: A;
COMPND   4 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: OPITUTUS TERRAE PB90-1;
SOURCE   3 ORGANISM_TAXID: 452637;
SOURCE   4 GENE: OTER_0116;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008
KEYWDS    CARBOHYDRATE ESTERASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    L.LO LEGGIO,J.LARSBRINK,R.MELAND KNUDSEN,S.MAZURKEWICH,J.C.NAVARRO
AUTHOR   2 POULSEN
REVDAT   1   15-AUG-18 6GRW    0
JRNL        AUTH   J.ARNLING BAATH,S.MAZURKEWICH,R.M.KNUDSEN,J.N.POULSEN,
JRNL        AUTH 2 L.OLSSON,L.LO LEGGIO,J.LARSBRINK
JRNL        TITL   BIOCHEMICAL AND STRUCTURAL FEATURES OF DIVERSE BACTERIAL
JRNL        TITL 2 GLUCURONOYL ESTERASES FACILITATING RECALCITRANT BIOMASS
JRNL        TITL 3 CONVERSION.
JRNL        REF    BIOTECHNOL BIOFUELS           V.  11   213 2018
JRNL        REFN                   ESSN 1754-6834
JRNL        PMID   30083226
JRNL        DOI    10.1186/S13068-018-1213-X
REMARK   2
REMARK   2 RESOLUTION.    1.50 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.13_2998
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.50
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 46.27
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.990
REMARK   3   COMPLETENESS FOR RANGE        (%) : 96.1
REMARK   3   NUMBER OF REFLECTIONS             : 50799
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.136
REMARK   3   R VALUE            (WORKING SET) : 0.135
REMARK   3   FREE R VALUE                     : 0.159
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.140
REMARK   3   FREE R VALUE TEST SET COUNT      : 2101
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 46.2900 -  3.7000    0.98     3324   144  0.1319 0.1405
REMARK   3     2  3.7000 -  2.9300    0.98     3309   142  0.1244 0.1503
REMARK   3     3  2.9300 -  2.5600    0.98     3311   143  0.1260 0.1439
REMARK   3     4  2.5600 -  2.3300    0.97     3291   142  0.1245 0.1570
REMARK   3     5  2.3300 -  2.1600    0.98     3321   143  0.1267 0.1410
REMARK   3     6  2.1600 -  2.0300    0.97     3258   141  0.1307 0.1590
REMARK   3     7  2.0300 -  1.9300    0.96     3266   141  0.1346 0.1777
REMARK   3     8  1.9300 -  1.8500    0.96     3246   140  0.1411 0.1838
REMARK   3     9  1.8500 -  1.7800    0.96     3242   140  0.1394 0.1660
REMARK   3    10  1.7800 -  1.7200    0.96     3223   139  0.1419 0.1657
REMARK   3    11  1.7200 -  1.6600    0.95     3228   139  0.1501 0.1786
REMARK   3    12  1.6600 -  1.6100    0.95     3217   139  0.1556 0.2002
REMARK   3    13  1.6100 -  1.5700    0.95     3205   138  0.1655 0.1797
REMARK   3    14  1.5700 -  1.5300    0.95     3162   136  0.1747 0.2246
REMARK   3    15  1.5300 -  1.5000    0.92     3095   134  0.1980 0.2402
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : NULL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.110
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 17.074
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 14.57
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 20.16
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.008           3293
REMARK   3   ANGLE     :  1.093           4504
REMARK   3   CHIRALITY :  0.076            471
REMARK   3   PLANARITY :  0.009            603
REMARK   3   DIHEDRAL  : 17.594           1208
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 6GRW COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 13-JUN-18.
REMARK 100 THE DEPOSITION ID IS D_1200010018.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 26-AUG-17
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 8.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : PETRA III, DESY
REMARK 200  BEAMLINE                       : P11
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.979
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS BUILT=20180126
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 50802
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.500
REMARK 200  RESOLUTION RANGE LOW       (A) : 46.270
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 95.9
REMARK 200  DATA REDUNDANCY                : 3.550
REMARK 200  R MERGE                    (I) : 0.04200
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 16.6900
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.50
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.59
REMARK 200  COMPLETENESS FOR SHELL     (%) : 93.3
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.60
REMARK 200  R MERGE FOR SHELL          (I) : 0.31400
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 3.400
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHENIX 1.13_2998
REMARK 200 STARTING MODEL: STRUCTURE UNDER DEPOSITION
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 37.16
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 1.96
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: RESERVOIR COMPOSITION FOR CRYSTAL
REMARK 280  GROWTH: 0.03M MAGNESIUM CHLORIDE, 0.03 M CALCIUM CHLORIDE, 0.05M
REMARK 280  IMIDAZOLE, 0.05M MES, 10% V/V MPD, 10% W/V PEG 1000, 10% V/W PEG
REMARK 280  3500P. HEAVY ATOM DERIVATIZATION: 1.2UL CRYSTALLIZATION MOTHER
REMARK 280  LIQUOR CONTAINING 0.5MM KAU(CN)2 ADDED TO DROP CONTAINING
REMARK 280  CRYSTALS. DROP SIZE AND COMPOSITION: SITTING DROPS OF 0.4UL WERE
REMARK 280  MIXED IN A PROTEIN:RESERVOIR VOLUME RATIO OF 1:1 USING 45 MG/ML
REMARK 280  OF OTCE15A IN 20 MM TRIS PH 8.0, VAPOR DIFFUSION, SITTING DROP,
REMARK 280  TEMPERATURE 293.15K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 1630 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 15600 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -24.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLY A    -2
REMARK 465     HIS A    -1
REMARK 465     ASP A   221
REMARK 465     ALA A   222
REMARK 465     ALA A   223
REMARK 465     ALA A   432
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   HD1  HIS A   148     H    ASP A   150              1.31
REMARK 500   OE1  GLN A    64    HH12  ARG A    73              1.59
REMARK 500   O    PRO A   431     O    HOH A   601              1.65
REMARK 500   O    HOH A   927     O    HOH A   972              1.85
REMARK 500   O    HOH A   604     O    HOH A   927              2.09
REMARK 500   O    HOH A   935     O    HOH A   994              2.15
REMARK 500   O2   EDO A   508     O    HOH A   602              2.19
REMARK 500   O    PRO A   206     O    HOH A   603              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ALA A  33       62.04   -103.99
REMARK 500    VAL A  70      -73.43   -120.99
REMARK 500    ASP A 150      108.21    -25.42
REMARK 500    SER A 267     -128.53     56.78
REMARK 500    HIS A 328       41.70   -144.44
REMARK 500    ASP A 360       73.94     67.30
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 LEU A  430     PRO A  431                  122.42
REMARK 500
REMARK 500 REMARK: NULL
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              AU A 501  AU
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 CYS A 293   SG
REMARK 620 2 HOH A 902   O   100.1
REMARK 620 N                    1
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              AU A 502  AU
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 CYS A 293   SG
REMARK 620 2 GLY A 294   N    89.7
REMARK 620 N                    1
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              CA A 503  CA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HIS A 303   O
REMARK 620 2 GLU A 305   O    95.0
REMARK 620 3 HOH A 829   O   100.1  91.9
REMARK 620 4 HOH A 616   O    86.8  85.6 172.8
REMARK 620 5 HOH A 636   O    89.8 160.8 105.5  76.1
REMARK 620 6 HOH A 902   O   167.7  95.5  85.9  87.6  78.2
REMARK 620 N                    1     2     3     4     5
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              MG A 505  MG
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HOH A 953   O
REMARK 620 2 HOH A 614   O   164.3
REMARK 620 3 HOH A 676   O    73.1  96.2
REMARK 620 4 HOH A 717   O   102.5  85.1 165.9
REMARK 620 5 HOH A 916   O    82.0  87.2  92.8  73.2
REMARK 620 6 HOH A 964   O    94.2  98.6  97.4  96.3 167.6
REMARK 620 N                    1     2     3     4     5
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue AU A 501
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue AU A 502
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue CA A 503
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue MG A 504
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue MG A 505
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue MPD A 506
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 507
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 508
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 509
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 510
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 511
DBREF  6GRW A   33   432  UNP    B1ZMF4   B1ZMF4_OPITP    33    432
SEQADV 6GRW GLY A   -2  UNP  B1ZMF4              EXPRESSION TAG
SEQADV 6GRW HIS A   -1  UNP  B1ZMF4              EXPRESSION TAG
SEQADV 6GRW SER A    0  UNP  B1ZMF4              EXPRESSION TAG
SEQRES   1 A  403  GLY HIS SER ALA TYR THR LEU PRO ASP PRO LEU VAL GLY
SEQRES   2 A  403  ALA ASP GLY THR ARG VAL HIS ASP ARG ALA THR TRP GLN
SEQRES   3 A  403  HIS ARG ARG ARG PRO GLU LEU LEU GLN LEU PHE ALA ARG
SEQRES   4 A  403  GLU VAL TYR GLY ARG THR PRO LEU GLY ARG PRO GLU GLY
SEQRES   5 A  403  MET VAL PHE LYS VAL THR THR MET GLU HIS ALA ALA LEU
SEQRES   6 A  403  GLY GLY ALA ALA THR ARG LYS GLU VAL THR VAL ARG PHE
SEQRES   7 A  403  GLY ARG ASP PRO ASN ALA PRO SER MET GLN LEU LEU LEU
SEQRES   8 A  403  TYR VAL PRO ASN ALA VAL ILE ALA ARG ALA GLU ARG ALA
SEQRES   9 A  403  PRO VAL PHE LEU GLY LEU ASN PHE TYR GLY ASN HIS THR
SEQRES  10 A  403  VAL HIS THR ASP PRO ALA ILE ALA LEU SER ALA ARG TRP
SEQRES  11 A  403  ILE PRO ALA GLU ALA PRO ASN GLY ALA ASN HIS ARG ALA
SEQRES  12 A  403  THR GLU ALA ALA ARG GLY SER ASP ALA GLN LYS TRP PRO
SEQRES  13 A  403  VAL GLU GLN ILE LEU ALA ARG GLY TYR ALA VAL ALA THR
SEQRES  14 A  403  VAL TYR CYS GLY ASP LEU CYS PRO ASP ARG PRO ASP GLY
SEQRES  15 A  403  LEU ASN ALA SER VAL ALA SER TRP LEU ASP ALA ALA ALA
SEQRES  16 A  403  GLY ASP GLN ARG ALA PRO ASP ALA TRP GLY ALA ILE GLY
SEQRES  17 A  403  VAL TRP ALA TRP GLY LEU SER ARG ALA LEU ASP TYR LEU
SEQRES  18 A  403  GLU THR ASP PRO LEU VAL ASP ALA SER ARG VAL ALA VAL
SEQRES  19 A  403  HIS GLY HIS SER ARG LEU GLY LYS ALA ALA LEU TRP ALA
SEQRES  20 A  403  GLY ALA GLN ASP ASP ARG PHE ALA LEU VAL ILE SER ASN
SEQRES  21 A  403  GLU SER GLY CYS GLY GLY ALA ALA LEU SER LYS ARG ILE
SEQRES  22 A  403  HIS GLY GLU THR VAL ALA ARG ILE ASN THR VAL PHE PRO
SEQRES  23 A  403  HIS TRP PHE ALA ARG ASN PHE ARG ARG TYR ASP ASP HIS
SEQRES  24 A  403  GLU GLU ALA LEU PRO VAL ASP GLN HIS GLU LEU LEU ALA
SEQRES  25 A  403  LEU VAL ALA PRO ARG PRO LEU TYR VAL ALA SER ALA GLU
SEQRES  26 A  403  ASP ASP ASP TRP ALA ASP PRO ARG GLY GLU PHE LEU ALA
SEQRES  27 A  403  VAL LYS ALA ALA GLU PRO VAL PHE ARG LEU PHE GLY GLN
SEQRES  28 A  403  THR GLY PRO SER GLY GLU ASP VAL PRO ARG VAL ASN GLU
SEQRES  29 A  403  PRO SER GLY GLY ALA LEU ARG TYR HIS ILE ARG PRO GLY
SEQRES  30 A  403  PRO HIS GLY MET THR ALA GLN ASP TRP ALA PHE TYR LEU
SEQRES  31 A  403  ALA PHE ALA ASP GLU TRP LEU LYS SER ALA LEU PRO ALA
HET     AU  A 501       1
HET     AU  A 502       1
HET     CA  A 503       1
HET     MG  A 504       1
HET     MG  A 505       1
HET    MPD  A 506      22
HET    EDO  A 507      10
HET    EDO  A 508      10
HET    EDO  A 509      10
HET    EDO  A 510      10
HET    EDO  A 511      10
HETNAM      AU GOLD ION
HETNAM      CA CALCIUM ION
HETNAM      MG MAGNESIUM ION
HETNAM     MPD (4S)-2-METHYL-2,4-PENTANEDIOL
HETNAM     EDO 1,2-ETHANEDIOL
HETSYN     EDO ETHYLENE GLYCOL
FORMUL   2   AU    2(AU 1+)
FORMUL   4   CA    CA 2+
FORMUL   5   MG    2(MG 2+)
FORMUL   7  MPD    C6 H14 O2
FORMUL   8  EDO    5(C2 H6 O2)
FORMUL  13  HOH   *402(H2 O)
HELIX    1 AA1 ASP A   50  ARG A   57  1                                   8
HELIX    2 AA2 ARG A   57  VAL A   70  1                                  14
HELIX    3 AA3 LEU A   94  GLY A   96  5                                   3
HELIX    4 AA4 ASN A  124  ARG A  129  1                                   6
HELIX    5 AA5 GLY A  143  VAL A  147  5                                   5
HELIX    6 AA6 GLU A  174  ARG A  177  5                                   4
HELIX    7 AA7 ASP A  180  TRP A  184  5                                   5
HELIX    8 AA8 PRO A  185  ARG A  192  1                                   8
HELIX    9 AA9 GLY A  202  LEU A  204  5                                   3
HELIX   10 AB1 ASP A  210  ALA A  214  5                                   5
HELIX   11 AB2 SER A  215  LEU A  220  1                                   6
HELIX   12 AB3 GLY A  234  GLU A  251  1                                  18
HELIX   13 AB4 SER A  267  ASP A  280  1                                  14
HELIX   14 AB5 THR A  306  PHE A  314  1                                   9
HELIX   15 AB6 ALA A  319  ASP A  326  5                                   8
HELIX   16 AB7 HIS A  328  LEU A  332  5                                   5
HELIX   17 AB8 ASP A  335  LEU A  342  1                                   8
HELIX   18 AB9 ASP A  356  ALA A  359  5                                   4
HELIX   19 AC1 ASP A  360  PHE A  378  1                                  19
HELIX   20 AC2 THR A  411  LEU A  426  1                                  16
SHEET    1 AA1 9 VAL A  83  ALA A  93  0
SHEET    2 AA1 9 ALA A  98  ARG A 106 -1  O  THR A 104   N  LYS A  85
SHEET    3 AA1 9 SER A 115  PRO A 123 -1  O  LEU A 118   N  VAL A 103
SHEET    4 AA1 9 ALA A 195  TYR A 200 -1  O  THR A 198   N  LEU A 119
SHEET    5 AA1 9 ALA A 133  ASN A 140  1  N  PRO A 134   O  ALA A 195
SHEET    6 AA1 9 VAL A 256  HIS A 266  1  O  ALA A 262   N  VAL A 135
SHEET    7 AA1 9 LEU A 285  ASN A 289  1  O  ILE A 287   N  VAL A 263
SHEET    8 AA1 9 LEU A 348  ALA A 353  1  O  TYR A 349   N  SER A 288
SHEET    9 AA1 9 LEU A 399  ARG A 404  1  O  ARG A 400   N  VAL A 350
SHEET    1 AA2 2 GLY A 167  ALA A 168  0
SHEET    2 AA2 2 ARG A 171  ALA A 172 -1  O  ARG A 171   N  ALA A 168
LINK         SG  CYS A 293                AU    AU A 501     1555   1555  2.25
LINK         SG  CYS A 293                AU    AU A 502     1555   1555  2.44
LINK         N   GLY A 294                AU    AU A 502     1555   1555  2.72
LINK         O   HIS A 303                CA    CA A 503     1555   1555  2.26
LINK         O   GLU A 305                CA    CA A 503     1555   1555  2.37
LINK        AU    AU A 501                 O   HOH A 902     1555   1555  2.48
LINK        CA    CA A 503                 O   HOH A 829     1555   1555  2.40
LINK        CA    CA A 503                 O   HOH A 616     1555   1555  2.32
LINK        CA    CA A 503                 O   HOH A 636     1555   1555  2.34
LINK        CA    CA A 503                 O   HOH A 902     1555   1555  2.38
LINK        MG    MG A 504                 O   HOH A 852     1555   1555  2.64
LINK        MG    MG A 505                 O   HOH A 953     1555   1555  2.66
LINK        MG    MG A 505                 O   HOH A 614     1555   1655  2.02
LINK        MG    MG A 505                 O   HOH A 676     1555   1655  2.11
LINK        MG    MG A 505                 O   HOH A 717     1555   1655  2.08
LINK        MG    MG A 505                 O   HOH A 916     1555   1655  2.30
LINK        MG    MG A 505                 O   HOH A 964     1555   1655  2.07
CISPEP   1 ALA A  344    PRO A  345          0         5.06
SITE     1 AC1  8 CYS A 293  GLU A 305  THR A 306  VAL A 307
SITE     2 AC1  8  AU A 502   CA A 503  HOH A 756  HOH A 902
SITE     1 AC2  6 CYS A 293  GLY A 294  SER A 299  PHE A 322
SITE     2 AC2  6  AU A 501  HOH A 655
SITE     1 AC3  7 HIS A 303  GLU A 305   AU A 501  HOH A 616
SITE     2 AC3  7 HOH A 636  HOH A 829  HOH A 902
SITE     1 AC4  3 LEU A  94  GLY A  95  HOH A 852
SITE     1 AC5  6 HOH A 614  HOH A 676  HOH A 717  HOH A 916
SITE     2 AC5  6 HOH A 953  HOH A 964
SITE     1 AC6  5 VAL A  41  GLY A  42  ALA A  43  GLN A 413
SITE     2 AC6  5 HOH A 719
SITE     1 AC7  6 GLY A  81  ARG A 109  ARG A 362  ASP A 387
SITE     2 AC7  6 VAL A 388  HOH A 723
SITE     1 AC8  5 ARG A 158  TRP A 159  PRO A 206  HOH A 602
SITE     2 AC8  5 HOH A 832
SITE     1 AC9  5 ALA A 284  LYS A 427  EDO A 510  HOH A 621
SITE     2 AC9  5 HOH A 662
SITE     1 AD1  6 SER A 259  ARG A 282  LYS A 427  EDO A 509
SITE     2 AD1  6 HOH A 662  HOH A 797
SITE     1 AD2  4 ALA A 130  ASP A 226  GLN A 227  HOH A 619
CRYST1   43.693   44.608   50.558  76.57  67.17  70.65 P 1           1
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.022887 -0.008038 -0.008738        0.00000
SCALE2      0.000000  0.023760 -0.002852        0.00000
SCALE3      0.000000  0.000000  0.021614        0.00000
TER    6250      PRO A 431
MASTER      351    0   11   20   11    0   20    6 3508    1   88   31
END