longtext: 6hxa-pdb

content
HEADER    LYASE                                   16-OCT-18   6HXA
TITLE     ANTI FROM P. LUMINESCENS CATALYSES TERMINAL POLYKETIDE SHORTENING IN
TITLE    2 THE BIOSYNTHESIS OF ANTHRAQUINONES
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ANTI;
COMPND   3 CHAIN: A;
COMPND   4 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PHOTORHABDUS LUMINESCENS;
SOURCE   3 ORGANISM_TAXID: 29488;
SOURCE   4 GENE: PLU4186;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   7 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   8 EXPRESSION_SYSTEM_PLASMID: PCOLA
KEYWDS    NATURAL PRODUCT BIOSYNTHESIS, TYPE II POLYKETIDE SYNTHASE SYSTEM,
KEYWDS   2 LYASE, ALPHA, BETA-HYDROLASE FOLD, CASCADE REACTION, RETRO CLAISEN
KEYWDS   3 REACTION, DIECKMANN CONDENSATION
EXPDTA    X-RAY DIFFRACTION
AUTHOR    Q.ZHOU,A.BRAEUER,G.GRAMMBITTER,M.SCHMALHOFER,P.SAURA,H.ADIHOU,
AUTHOR   2 V.R.I.KAILA,M.GROLL,H.BODE
REVDAT   1   07-AUG-19 6HXA    0
JRNL        AUTH   Q.ZHOU,A.BRAUER,H.ADIHOU,M.SCHMALHOFER,P.SAURA,
JRNL        AUTH 2 G.L.C.GRAMMBITTER,V.R.I.KAILA,M.GROLL,H.B.BODE
JRNL        TITL   MOLECULAR MECHANISM OF POLYKETIDE SHORTENING IN
JRNL        TITL 2 ANTHRAQUINONE BIOSYNTHESIS OFPHOTORHABDUS LUMINESCENS.
JRNL        REF    CHEM SCI                      V.  10  6341 2019
JRNL        REFN                   ISSN 2041-6520
JRNL        PMID   31341589
JRNL        DOI    10.1039/C9SC00749K
REMARK   2
REMARK   2 RESOLUTION.    1.85 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0230
REMARK   3   AUTHORS     : MURSHUDOV,SKUBAK,LEBEDEV,PANNU,STEINER,
REMARK   3               : NICHOLLS,WINN,LONG,VAGIN
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.85
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 30.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.7
REMARK   3   NUMBER OF REFLECTIONS             : 31340
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.170
REMARK   3   R VALUE            (WORKING SET) : 0.168
REMARK   3   FREE R VALUE                     : 0.204
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 1649
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.85
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.90
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 2295
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 100.0
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2890
REMARK   3   BIN FREE R VALUE SET COUNT          : 120
REMARK   3   BIN FREE R VALUE                    : 0.2960
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 3039
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 23
REMARK   3   SOLVENT ATOMS            : 182
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 37.11
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.36000
REMARK   3    B22 (A**2) : -4.11000
REMARK   3    B33 (A**2) : 3.75000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.390
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.126
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.093
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 7.206
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.971
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.960
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  3153 ; 0.003 ; 0.014
REMARK   3   BOND LENGTHS OTHERS               (A):  2808 ; 0.001 ; 0.017
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  4259 ; 0.789 ; 1.638
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  6568 ; 0.740 ; 1.647
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   376 ; 6.310 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   168 ;31.921 ;23.214
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   532 ;12.712 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    13 ;16.324 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   390 ; 0.039 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  3512 ; 0.003 ; 0.020
REMARK   3   GENERAL PLANES OTHERS             (A):   644 ; 0.001 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  1501 ; 1.039 ; 3.425
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  1500 ; 1.037 ; 3.424
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  1875 ; 1.495 ; 5.134
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  1876 ; 1.494 ; 5.136
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  1652 ; 1.043 ; 3.708
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  1652 ; 1.043 ; 3.708
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  2384 ; 1.384 ; 5.455
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  3655 ; 2.325 ;39.954
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  3623 ; 2.246 ;39.701
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  5956 ; 0.352 ; 3.000
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):    85 ;31.382 ; 5.000
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  5976 ;18.333 ; 5.000
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 1
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A     7        A   381
REMARK   3    ORIGIN FOR THE GROUP (A):  13.0990  17.1772  94.0862
REMARK   3    T TENSOR
REMARK   3      T11:   0.0005 T22:   0.0400
REMARK   3      T33:   0.0195 T12:   0.0004
REMARK   3      T13:   0.0016 T23:  -0.0003
REMARK   3    L TENSOR
REMARK   3      L11:   0.1063 L22:   0.0949
REMARK   3      L33:   0.0125 L12:   0.0423
REMARK   3      L13:   0.0011 L23:  -0.0118
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0013 S12:  -0.0016 S13:   0.0199
REMARK   3      S21:   0.0003 S22:   0.0017 S23:   0.0085
REMARK   3      S31:   0.0015 S32:   0.0058 S33:  -0.0004
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : NULL
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS
REMARK   4
REMARK   4 6HXA COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 16-OCT-18.
REMARK 100 THE DEPOSITION ID IS D_1200012442.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 18-JUN-16
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 7.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SLS
REMARK 200  BEAMLINE                       : X06SA
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS EIGER X 16M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 33019
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.850
REMARK 200  RESOLUTION RANGE LOW       (A) : 30.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.7
REMARK 200  DATA REDUNDANCY                : 6.700
REMARK 200  R MERGE                    (I) : 0.05700
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 18.9000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.85
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.95
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : 0.56000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 3.000
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: SHARP
REMARK 200 STARTING MODEL: SEMET DERIVATIVE; COORDINATES NOT DEPOSITED
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 41.00
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.08
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 100 MM HEPES, 0.1 M NAAC, 22% PEG
REMARK 280  3350, PH 7.5, VAPOR DIFFUSION, SITTING DROP, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 2 2 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z+1/2
REMARK 290       3555   -X,Y,-Z+1/2
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z
REMARK 290       6555   -X+1/2,-Y+1/2,Z+1/2
REMARK 290       7555   -X+1/2,Y+1/2,-Z+1/2
REMARK 290       8555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       45.62000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       45.62000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       27.05500
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       77.17000
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       27.05500
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       77.17000
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       45.62000
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       27.05500
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       77.17000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       45.62000
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       27.05500
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000       77.17000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 4510 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 28100 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -14.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000      182.48000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH A 617  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A 676  LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLY A   -14
REMARK 465     SER A   -13
REMARK 465     SER A   -12
REMARK 465     HIS A   -11
REMARK 465     HIS A   -10
REMARK 465     HIS A    -9
REMARK 465     HIS A    -8
REMARK 465     HIS A    -7
REMARK 465     HIS A    -6
REMARK 465     SER A    -5
REMARK 465     GLY A    -4
REMARK 465     ASP A    -3
REMARK 465     PRO A    -2
REMARK 465     ALA A    -1
REMARK 465     SER A     0
REMARK 465     MET A     1
REMARK 465     ASN A     2
REMARK 465     ASN A     3
REMARK 465     LYS A     4
REMARK 465     ASN A     5
REMARK 465     LYS A     6
REMARK 465     LYS A   382
REMARK 465     LYS A   383
REMARK 465     GLY A   384
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O    HOH A   682     O    HOH A   682     3656     1.82
REMARK 500   OH   TYR A    20     CD   GLU A   160     1455     2.12
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    PHE A  27     -128.51     48.48
REMARK 500    ASP A 143     -106.36     65.31
REMARK 500    ILE A 180      -87.92   -100.87
REMARK 500    SER A 245     -122.86     56.84
REMARK 500    VAL A 356       -0.10     81.08
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: PLANAR GROUPS
REMARK 500
REMARK 500 PLANAR GROUPS IN THE FOLLOWING RESIDUES HAVE A TOTAL
REMARK 500 RMS DISTANCE OF ALL ATOMS FROM THE BEST-FIT PLANE
REMARK 500 BY MORE THAN AN EXPECTED VALUE OF 6*RMSD, WITH AN
REMARK 500 RMSD 0.02 ANGSTROMS, OR AT LEAST ONE ATOM HAS
REMARK 500 AN RMSD GREATER THAN THIS VALUE
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500  M RES CSSEQI        RMS     TYPE
REMARK 500    ARG A 117         0.09    SIDE CHAIN
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue PGE A 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue PG4 A 402
DBREF  6HXA A    1   384  UNP    Q7MZT8   Q7MZT8_PHOLL     1    384
SEQADV 6HXA GLY A  -14  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA SER A  -13  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA SER A  -12  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA HIS A  -11  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA HIS A  -10  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA HIS A   -9  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA HIS A   -8  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA HIS A   -7  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA HIS A   -6  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA SER A   -5  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA GLY A   -4  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA ASP A   -3  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA PRO A   -2  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA ALA A   -1  UNP  Q7MZT8              EXPRESSION TAG
SEQADV 6HXA SER A    0  UNP  Q7MZT8              EXPRESSION TAG
SEQRES   1 A  399  GLY SER SER HIS HIS HIS HIS HIS HIS SER GLY ASP PRO
SEQRES   2 A  399  ALA SER MET ASN ASN LYS ASN LYS PRO ASN ARG ILE SER
SEQRES   3 A  399  PRO GLU LEU LEU ALA THR CYS GLY TYR PHE MET PRO ARG
SEQRES   4 A  399  ILE PHE PHE LEU ASN SER GLN TYR ALA PRO GLN VAL HIS
SEQRES   5 A  399  TRP GLY ASP VAL VAL ALA ALA LEU SER HIS PHE PRO ALA
SEQRES   6 A  399  GLY ASN LEU ASP LEU SER SER GLU GLU PHE TRP TYR GLU
SEQRES   7 A  399  TRP MET ILE ASN TRP SER LYS VAL GLY ASP SER TYR ILE
SEQRES   8 A  399  ASN ILE ALA ASN SER ALA LYS SER GLU VAL SER HIS VAL
SEQRES   9 A  399  ARG ALA LEU ARG SER ALA ALA ALA CYS TYR HIS TRP ALA
SEQRES  10 A  399  GLU PHE MET TYR PHE SER ASP ARG SER ARG LYS ILE GLN
SEQRES  11 A  399  LEU ARG GLU TYR ILE ARG SER CYS PHE LEU SER SER ILE
SEQRES  12 A  399  LYS TYR SER ASP LEU LEU VAL ASP HIS GLN TYR ILE VAL
SEQRES  13 A  399  VAL ASP LYS PHE HIS MET PRO PHE PHE LEU ILE PHE PRO
SEQRES  14 A  399  LYS GLY TYR LYS GLU GLU GLU ASN HIS PRO LEU PRO CYS
SEQRES  15 A  399  VAL ILE LEU SER ASN GLY LEU ASP SER MET THR GLU ILE
SEQRES  16 A  399  GLU ILE LEU SER LEU ALA GLU PHE PHE LEU GLY LYS ASN
SEQRES  17 A  399  MET ALA VAL ALA ILE PHE ASP GLY PRO GLY GLN GLY ILE
SEQRES  18 A  399  ASN LEU GLY LYS SER PRO ILE ALA ILE ASP MET GLU LEU
SEQRES  19 A  399  TYR VAL SER SER ILE VAL LYS LEU LEU GLU ASP ASP ALA
SEQRES  20 A  399  ARG ILE ASN SER ASN LEU LEU CYS PHE LEU GLY ILE SER
SEQRES  21 A  399  PHE GLY GLY TYR PHE ALA LEU ARG VAL ALA GLN ARG ILE
SEQRES  22 A  399  GLY ASP LYS PHE CYS CYS ILE VAL ASN LEU SER GLY GLY
SEQRES  23 A  399  PRO GLU ILE ALA GLU PHE ASP LYS LEU PRO ARG ARG LEU
SEQRES  24 A  399  LYS GLU ASP PHE GLN PHE ALA PHE MET GLN ASP ASN SER
SEQRES  25 A  399  HIS MET GLN SER ILE PHE ASP GLU ILE LYS LEU ASP ILE
SEQRES  26 A  399  SER LEU PRO CYS LYS THR LYS VAL PHE THR VAL HIS GLY
SEQRES  27 A  399  GLU LEU ASP ASP ILE PHE GLN ILE ASP LYS VAL LYS LYS
SEQRES  28 A  399  LEU ASP GLN LEU TRP GLY ASP ASN HIS GLN LEU LEU CYS
SEQRES  29 A  399  TYR GLU SER GLU ALA HIS VAL CYS LEU ASN LYS ILE ASN
SEQRES  30 A  399  GLU TYR MET ILE GLN VAL SER ASP TRP VAL SER GLU GLN
SEQRES  31 A  399  PHE TRP LEU ASN GLY TYR LYS LYS GLY
HET    PGE  A 401      10
HET    PG4  A 402      13
HETNAM     PGE TRIETHYLENE GLYCOL
HETNAM     PG4 TETRAETHYLENE GLYCOL
FORMUL   2  PGE    C6 H14 O4
FORMUL   3  PG4    C8 H18 O5
FORMUL   4  HOH   *182(H2 O)
HELIX    1 AA1 SER A   11  CYS A   18  1                                   8
HELIX    2 AA2 PHE A   21  PHE A   27  1                                   7
HELIX    3 AA3 HIS A   37  SER A   46  1                                  10
HELIX    4 AA4 SER A   57  ALA A   82  1                                  26
HELIX    5 AA5 SER A   84  PHE A  104  1                                  21
HELIX    6 AA6 ASP A  109  TYR A  130  1                                  22
HELIX    7 AA7 ILE A  180  GLY A  191  1                                  12
HELIX    8 AA8 GLN A  204  LEU A  208  5                                   5
HELIX    9 AA9 MET A  217  ASP A  231  1                                  15
HELIX   10 AB1 SER A  245  GLY A  259  1                                  15
HELIX   11 AB2 ASP A  260  PHE A  262  5                                   3
HELIX   12 AB3 GLU A  276  LEU A  280  5                                   5
HELIX   13 AB4 PRO A  281  MET A  293  1                                  13
HELIX   14 AB5 ASP A  295  SER A  297  5                                   3
HELIX   15 AB6 HIS A  298  LYS A  307  1                                  10
HELIX   16 AB7 GLN A  330  GLY A  342  1                                  13
HELIX   17 AB8 ALA A  354  ASN A  359  5                                   6
HELIX   18 AB9 LYS A  360  ASN A  379  1                                  20
SHEET    1 AA1 8 VAL A 135  VAL A 142  0
SHEET    2 AA1 8 PHE A 145  ILE A 152 -1  O  LEU A 151   N  ASP A 136
SHEET    3 AA1 8 ALA A 195  PHE A 199 -1  O  ILE A 198   N  PHE A 150
SHEET    4 AA1 8 LEU A 165  SER A 171  1  N  VAL A 168   O  ALA A 197
SHEET    5 AA1 8 ILE A 234  ILE A 244  1  O  ASN A 235   N  LEU A 165
SHEET    6 AA1 8 CYS A 264  LEU A 268  1  O  LEU A 268   N  GLY A 243
SHEET    7 AA1 8 LYS A 317  GLY A 323  1  O  PHE A 319   N  ASN A 267
SHEET    8 AA1 8 HIS A 345  TYR A 350  1  O  LEU A 348   N  THR A 320
SITE     1 AC1  3 TYR A  99  TYR A 119  HOH A 584
SITE     1 AC2  4 ALA A  82  SER A  84  TYR A 130  HOH A 614
CRYST1   54.110  154.340   91.240  90.00  90.00  90.00 C 2 2 21      8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.018481  0.000000  0.000000        0.00000
SCALE2      0.000000  0.006479  0.000000        0.00000
SCALE3      0.000000  0.000000  0.010960        0.00000
TER    3049      TYR A 381
MASTER      383    0    2   18    8    0    2    6 3244    1   23   31
END