longtext: 6isr-pdb

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HEADER    HYDROLASE                               18-NOV-18   6ISR
TITLE     STRUCTURE OF LIPASE MUTANT WITH CYS-HIS-ASP CATALYTIC TRIAD
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LIPASE B;
COMPND   3 CHAIN: A, B;
COMPND   4 SYNONYM: CALB;
COMPND   5 EC: 3.1.1.3;
COMPND   6 ENGINEERED: YES;
COMPND   7 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PSEUDOZYMA ANTARCTICA;
SOURCE   3 ORGANISM_COMMON: YEAST;
SOURCE   4 ORGANISM_TAXID: 84753;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008
KEYWDS    CALB, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    Y.X.CEN,J.H.ZHOU,Q.WU
REVDAT   1   24-JUL-19 6ISR    0
JRNL        AUTH   M.T.REETZ,J.H.ZHOU,T.S.MOODY,M.L.HUANG,Q.WU,Y.X.CEN,W.SINGH,
JRNL        AUTH 2 M.ARKIN
JRNL        TITL   ARTIFICIAL CYSTEINE-LIPASES WITH HIGH ACTIVITY AND ALTERED
JRNL        TITL 2 CATALYTIC MECHANISM CREATED BY LABORATORY EVOLUTION.
JRNL        REF    NAT COMMUN                                 2019
JRNL        REFN                   ESSN 2041-1723
JRNL        DOI    10.1038/S41467-019-11155-3
REMARK   2
REMARK   2 RESOLUTION.    2.60 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (1.10.1_2155: ???)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.60
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 42.25
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.360
REMARK   3   COMPLETENESS FOR RANGE        (%) : 91.4
REMARK   3   NUMBER OF REFLECTIONS             : 17958
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.181
REMARK   3   R VALUE            (WORKING SET) : 0.179
REMARK   3   FREE R VALUE                     : 0.228
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.880
REMARK   3   FREE R VALUE TEST SET COUNT      : 876
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 42.2545 -  4.6907    0.99     3190   154  0.1614 0.1838
REMARK   3     2  4.6907 -  3.7239    1.00     3078   187  0.1492 0.2059
REMARK   3     3  3.7239 -  3.2534    1.00     3095   151  0.1766 0.2497
REMARK   3     4  3.2534 -  2.9560    0.99     3079   150  0.2217 0.2708
REMARK   3     5  2.9560 -  2.7442    0.90     2752   157  0.2351 0.3017
REMARK   3     6  2.7442 -  2.5824    0.61     1888    77  0.2389 0.3290
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : NULL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.310
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 24.420
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.008           4790
REMARK   3   ANGLE     :  0.890           6561
REMARK   3   CHIRALITY :  0.055            757
REMARK   3   PLANARITY :  0.008            858
REMARK   3   DIHEDRAL  : 13.828           2891
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 1
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: ALL
REMARK   3    ORIGIN FOR THE GROUP (A): -12.6607  12.1282 -33.3530
REMARK   3    T TENSOR
REMARK   3      T11:   0.2020 T22:   0.2440
REMARK   3      T33:   0.2656 T12:  -0.0045
REMARK   3      T13:   0.0157 T23:   0.0112
REMARK   3    L TENSOR
REMARK   3      L11:  -0.0705 L22:   0.1138
REMARK   3      L33:   0.7371 L12:   0.0099
REMARK   3      L13:   0.0531 L23:   0.3759
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0095 S12:  -0.0073 S13:   0.0040
REMARK   3      S21:   0.0046 S22:   0.0117 S23:   0.0017
REMARK   3      S31:   0.0030 S32:  -0.0214 S33:   0.0028
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 6ISR COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 03-DEC-18.
REMARK 100 THE DEPOSITION ID IS D_1300009452.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 24-APR-18
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SSRF
REMARK 200  BEAMLINE                       : BL18U1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97930
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS3 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-3000
REMARK 200  DATA SCALING SOFTWARE          : HKL-3000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 19262
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.600
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.6
REMARK 200  DATA REDUNDANCY                : 4.500
REMARK 200  R MERGE                    (I) : 0.20000
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 8.8180
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.60
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.64
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : 0.82400
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 1.857
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHENIX
REMARK 200 STARTING MODEL: 1TCA
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 46.88
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.32
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1M SODIUM ACETATE PH 6.0 25% PEG4000
REMARK 280  8% ISOPROPANOL 0.5% N,N-DIMETHYLDODECYLAMINE OXIDE(DDAO), VAPOR
REMARK 280  DIFFUSION, SITTING DROP, TEMPERATURE 289K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       22.27950
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 140 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 12120 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -11.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 130 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 12160 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -11.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLY A    -3
REMARK 465     ALA A    -2
REMARK 465     MET A    -1
REMARK 465     ALA A     0
REMARK 465     LEU A     1
REMARK 465     PRO A     2
REMARK 465     SER A     3
REMARK 465     GLY B    -3
REMARK 465     ALA B    -2
REMARK 465     MET B    -1
REMARK 465     ALA B     0
REMARK 465     LEU B     1
REMARK 465     PRO B     2
REMARK 465     SER B     3
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OD2  ASP A   134     NE2  GLN A   157              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASN A  51      -92.49   -135.70
REMARK 500    ASN A  96       61.45     60.09
REMARK 500    CYS A 105     -131.14     56.49
REMARK 500    ASP A 134       66.40   -114.66
REMARK 500    ALA A 146       23.84   -143.97
REMARK 500    THR A 186       30.53    -89.52
REMARK 500    PRO A 198       -7.85    -58.18
REMARK 500    ASN A 206       -2.75     80.26
REMARK 500    THR A 310     -168.17   -128.77
REMARK 500    ASN B  51      -92.70   -136.72
REMARK 500    ASN B  96       60.73     60.55
REMARK 500    CYS B 105     -130.24     56.17
REMARK 500    ASP B 134       65.44   -113.32
REMARK 500    ALA B 146       24.07   -144.24
REMARK 500    THR B 186       30.61    -91.15
REMARK 500    PRO B 198       -7.30    -58.82
REMARK 500    ASN B 206       -1.64     78.04
REMARK 500    THR B 310     -168.69   -126.42
REMARK 500
REMARK 500 REMARK: NULL
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NI A 401  NI
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HIS A 224   NE2
REMARK 620 2 HOH A 525   O   116.8
REMARK 620 N                    1
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NI B 401  NI
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 CYS B 105   SG
REMARK 620 2 HIS B 224   NE2  97.1
REMARK 620 3 HOH B 525   O   132.0 111.6
REMARK 620 N                    1     2
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NI A 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue PEG A 402
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue PG4 A 403
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NI B 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue PEG B 402
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue PEG B 403
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue PG4 B 404
DBREF  6ISR A    1   317  UNP    P41365   LIPB_PSEA2      26    342
DBREF  6ISR B    1   317  UNP    P41365   LIPB_PSEA2      26    342
SEQADV 6ISR GLY A   -3  UNP  P41365              EXPRESSION TAG
SEQADV 6ISR ALA A   -2  UNP  P41365              EXPRESSION TAG
SEQADV 6ISR MET A   -1  UNP  P41365              EXPRESSION TAG
SEQADV 6ISR ALA A    0  UNP  P41365              EXPRESSION TAG
SEQADV 6ISR ALA A   57  UNP  P41365    THR    82 ENGINEERED MUTATION
SEQADV 6ISR THR A   89  UNP  P41365    ALA   114 ENGINEERED MUTATION
SEQADV 6ISR VAL A  104  UNP  P41365    TRP   129 ENGINEERED MUTATION
SEQADV 6ISR CYS A  105  UNP  P41365    SER   130 ENGINEERED MUTATION
SEQADV 6ISR GLY A  149  UNP  P41365    VAL   174 ENGINEERED MUTATION
SEQADV 6ISR TYR A  281  UNP  P41365    ALA   306 ENGINEERED MUTATION
SEQADV 6ISR TYR A  282  UNP  P41365    ALA   307 ENGINEERED MUTATION
SEQADV 6ISR GLY B   -3  UNP  P41365              EXPRESSION TAG
SEQADV 6ISR ALA B   -2  UNP  P41365              EXPRESSION TAG
SEQADV 6ISR MET B   -1  UNP  P41365              EXPRESSION TAG
SEQADV 6ISR ALA B    0  UNP  P41365              EXPRESSION TAG
SEQADV 6ISR ALA B   57  UNP  P41365    THR    82 ENGINEERED MUTATION
SEQADV 6ISR THR B   89  UNP  P41365    ALA   114 ENGINEERED MUTATION
SEQADV 6ISR VAL B  104  UNP  P41365    TRP   129 ENGINEERED MUTATION
SEQADV 6ISR CYS B  105  UNP  P41365    SER   130 ENGINEERED MUTATION
SEQADV 6ISR GLY B  149  UNP  P41365    VAL   174 ENGINEERED MUTATION
SEQADV 6ISR TYR B  281  UNP  P41365    ALA   306 ENGINEERED MUTATION
SEQADV 6ISR TYR B  282  UNP  P41365    ALA   307 ENGINEERED MUTATION
SEQRES   1 A  321  GLY ALA MET ALA LEU PRO SER GLY SER ASP PRO ALA PHE
SEQRES   2 A  321  SER GLN PRO LYS SER VAL LEU ASP ALA GLY LEU THR CYS
SEQRES   3 A  321  GLN GLY ALA SER PRO SER SER VAL SER LYS PRO ILE LEU
SEQRES   4 A  321  LEU VAL PRO GLY THR GLY THR THR GLY PRO GLN SER PHE
SEQRES   5 A  321  ASP SER ASN TRP ILE PRO LEU SER ALA GLN LEU GLY TYR
SEQRES   6 A  321  THR PRO CYS TRP ILE SER PRO PRO PRO PHE MET LEU ASN
SEQRES   7 A  321  ASP THR GLN VAL ASN THR GLU TYR MET VAL ASN ALA ILE
SEQRES   8 A  321  THR THR LEU TYR ALA GLY SER GLY ASN ASN LYS LEU PRO
SEQRES   9 A  321  VAL LEU THR VAL CYS GLN GLY GLY LEU VAL ALA GLN TRP
SEQRES  10 A  321  GLY LEU THR PHE PHE PRO SER ILE ARG SER LYS VAL ASP
SEQRES  11 A  321  ARG LEU MET ALA PHE ALA PRO ASP TYR LYS GLY THR VAL
SEQRES  12 A  321  LEU ALA GLY PRO LEU ASP ALA LEU ALA GLY SER ALA PRO
SEQRES  13 A  321  SER VAL TRP GLN GLN THR THR GLY SER ALA LEU THR THR
SEQRES  14 A  321  ALA LEU ARG ASN ALA GLY GLY LEU THR GLN ILE VAL PRO
SEQRES  15 A  321  THR THR ASN LEU TYR SER ALA THR ASP GLU ILE VAL GLN
SEQRES  16 A  321  PRO GLN VAL SER ASN SER PRO LEU ASP SER SER TYR LEU
SEQRES  17 A  321  PHE ASN GLY LYS ASN VAL GLN ALA GLN ALA VAL CYS GLY
SEQRES  18 A  321  PRO LEU PHE VAL ILE ASP HIS ALA GLY SER LEU THR SER
SEQRES  19 A  321  GLN PHE SER TYR VAL VAL GLY ARG SER ALA LEU ARG SER
SEQRES  20 A  321  THR THR GLY GLN ALA ARG SER ALA ASP TYR GLY ILE THR
SEQRES  21 A  321  ASP CYS ASN PRO LEU PRO ALA ASN ASP LEU THR PRO GLU
SEQRES  22 A  321  GLN LYS VAL ALA ALA ALA ALA LEU LEU ALA PRO TYR TYR
SEQRES  23 A  321  ALA ALA ILE VAL ALA GLY PRO LYS GLN ASN CYS GLU PRO
SEQRES  24 A  321  ASP LEU MET PRO TYR ALA ARG PRO PHE ALA VAL GLY LYS
SEQRES  25 A  321  ARG THR CYS SER GLY ILE VAL THR PRO
SEQRES   1 B  321  GLY ALA MET ALA LEU PRO SER GLY SER ASP PRO ALA PHE
SEQRES   2 B  321  SER GLN PRO LYS SER VAL LEU ASP ALA GLY LEU THR CYS
SEQRES   3 B  321  GLN GLY ALA SER PRO SER SER VAL SER LYS PRO ILE LEU
SEQRES   4 B  321  LEU VAL PRO GLY THR GLY THR THR GLY PRO GLN SER PHE
SEQRES   5 B  321  ASP SER ASN TRP ILE PRO LEU SER ALA GLN LEU GLY TYR
SEQRES   6 B  321  THR PRO CYS TRP ILE SER PRO PRO PRO PHE MET LEU ASN
SEQRES   7 B  321  ASP THR GLN VAL ASN THR GLU TYR MET VAL ASN ALA ILE
SEQRES   8 B  321  THR THR LEU TYR ALA GLY SER GLY ASN ASN LYS LEU PRO
SEQRES   9 B  321  VAL LEU THR VAL CYS GLN GLY GLY LEU VAL ALA GLN TRP
SEQRES  10 B  321  GLY LEU THR PHE PHE PRO SER ILE ARG SER LYS VAL ASP
SEQRES  11 B  321  ARG LEU MET ALA PHE ALA PRO ASP TYR LYS GLY THR VAL
SEQRES  12 B  321  LEU ALA GLY PRO LEU ASP ALA LEU ALA GLY SER ALA PRO
SEQRES  13 B  321  SER VAL TRP GLN GLN THR THR GLY SER ALA LEU THR THR
SEQRES  14 B  321  ALA LEU ARG ASN ALA GLY GLY LEU THR GLN ILE VAL PRO
SEQRES  15 B  321  THR THR ASN LEU TYR SER ALA THR ASP GLU ILE VAL GLN
SEQRES  16 B  321  PRO GLN VAL SER ASN SER PRO LEU ASP SER SER TYR LEU
SEQRES  17 B  321  PHE ASN GLY LYS ASN VAL GLN ALA GLN ALA VAL CYS GLY
SEQRES  18 B  321  PRO LEU PHE VAL ILE ASP HIS ALA GLY SER LEU THR SER
SEQRES  19 B  321  GLN PHE SER TYR VAL VAL GLY ARG SER ALA LEU ARG SER
SEQRES  20 B  321  THR THR GLY GLN ALA ARG SER ALA ASP TYR GLY ILE THR
SEQRES  21 B  321  ASP CYS ASN PRO LEU PRO ALA ASN ASP LEU THR PRO GLU
SEQRES  22 B  321  GLN LYS VAL ALA ALA ALA ALA LEU LEU ALA PRO TYR TYR
SEQRES  23 B  321  ALA ALA ILE VAL ALA GLY PRO LYS GLN ASN CYS GLU PRO
SEQRES  24 B  321  ASP LEU MET PRO TYR ALA ARG PRO PHE ALA VAL GLY LYS
SEQRES  25 B  321  ARG THR CYS SER GLY ILE VAL THR PRO
HET     NI  A 401       1
HET    PEG  A 402       7
HET    PG4  A 403      13
HET     NI  B 401       1
HET    PEG  B 402       7
HET    PEG  B 403       7
HET    PG4  B 404      13
HETNAM      NI NICKEL (II) ION
HETNAM     PEG DI(HYDROXYETHYL)ETHER
HETNAM     PG4 TETRAETHYLENE GLYCOL
FORMUL   3   NI    2(NI 2+)
FORMUL   4  PEG    3(C4 H10 O3)
FORMUL   5  PG4    2(C8 H18 O5)
FORMUL  10  HOH   *57(H2 O)
HELIX    1 AA1 PRO A   12  ALA A   18  1                                   7
HELIX    2 AA2 THR A   43  ASP A   49  1                                   7
HELIX    3 AA3 ASN A   51  LEU A   59  1                                   9
HELIX    4 AA4 ASP A   75  SER A   94  1                                  20
HELIX    5 AA5 CYS A  105  PHE A  118  1                                  14
HELIX    6 AA6 PRO A  119  ARG A  122  5                                   4
HELIX    7 AA7 THR A  138  GLY A  142  5                                   5
HELIX    8 AA8 PRO A  143  LEU A  147  5                                   5
HELIX    9 AA9 ALA A  151  GLN A  157  1                                   7
HELIX   10 AB1 SER A  161  ALA A  170  1                                  10
HELIX   11 AB2 ALA A  212  GLY A  217  1                                   6
HELIX   12 AB3 ALA A  225  SER A  230  1                                   6
HELIX   13 AB4 SER A  230  SER A  243  1                                  14
HELIX   14 AB5 GLY A  254  CYS A  258  5                                   5
HELIX   15 AB6 THR A  267  ALA A  276  1                                  10
HELIX   16 AB7 LEU A  278  ALA A  283  1                                   6
HELIX   17 AB8 ALA A  301  VAL A  306  5                                   6
HELIX   18 AB9 PRO B   12  ALA B   18  1                                   7
HELIX   19 AC1 THR B   43  ASP B   49  1                                   7
HELIX   20 AC2 ASN B   51  LEU B   59  1                                   9
HELIX   21 AC3 ASP B   75  SER B   94  1                                  20
HELIX   22 AC4 CYS B  105  PHE B  118  1                                  14
HELIX   23 AC5 PRO B  119  ARG B  122  5                                   4
HELIX   24 AC6 PRO B  143  LEU B  147  5                                   5
HELIX   25 AC7 ALA B  151  GLN B  157  1                                   7
HELIX   26 AC8 SER B  161  ALA B  170  1                                  10
HELIX   27 AC9 ALA B  212  GLY B  217  1                                   6
HELIX   28 AD1 ALA B  225  SER B  230  1                                   6
HELIX   29 AD2 SER B  230  SER B  243  1                                  14
HELIX   30 AD3 GLY B  254  CYS B  258  5                                   5
HELIX   31 AD4 THR B  267  ALA B  276  1                                  10
HELIX   32 AD5 LEU B  278  ILE B  285  5                                   8
HELIX   33 AD6 ALA B  301  VAL B  306  5                                   6
SHEET    1 AA1 7 LEU A  20  CYS A  22  0
SHEET    2 AA1 7 THR A  62  ILE A  66 -1  O  TRP A  65   N  THR A  21
SHEET    3 AA1 7 PRO A  33  VAL A  37  1  N  ILE A  34   O  CYS A  64
SHEET    4 AA1 7 LEU A  99  VAL A 104  1  O  LEU A 102   N  LEU A  35
SHEET    5 AA1 7 VAL A 125  PHE A 131  1  O  MET A 129   N  VAL A 101
SHEET    6 AA1 7 THR A 179  TYR A 183  1  O  THR A 180   N  ALA A 130
SHEET    7 AA1 7 LYS A 208  GLN A 211  1  O  VAL A 210   N  ASN A 181
SHEET    1 AA2 2 ARG A 309  THR A 310  0
SHEET    2 AA2 2 GLY A 313  ILE A 314 -1  O  GLY A 313   N  THR A 310
SHEET    1 AA3 7 LEU B  20  CYS B  22  0
SHEET    2 AA3 7 THR B  62  ILE B  66 -1  O  TRP B  65   N  THR B  21
SHEET    3 AA3 7 PRO B  33  VAL B  37  1  N  ILE B  34   O  CYS B  64
SHEET    4 AA3 7 LEU B  99  VAL B 104  1  O  LEU B 102   N  LEU B  35
SHEET    5 AA3 7 VAL B 125  PHE B 131  1  O  MET B 129   N  VAL B 101
SHEET    6 AA3 7 THR B 179  TYR B 183  1  O  THR B 180   N  ALA B 130
SHEET    7 AA3 7 LYS B 208  GLN B 211  1  O  VAL B 210   N  ASN B 181
SHEET    1 AA4 2 ARG B 309  THR B 310  0
SHEET    2 AA4 2 GLY B 313  ILE B 314 -1  O  GLY B 313   N  THR B 310
SSBOND   1 CYS A   22    CYS A   64                          1555   1555  2.05
SSBOND   2 CYS A  216    CYS A  258                          1555   1555  2.05
SSBOND   3 CYS A  293    CYS A  311                          1555   1555  2.05
SSBOND   4 CYS B   22    CYS B   64                          1555   1555  2.05
SSBOND   5 CYS B  216    CYS B  258                          1555   1555  2.04
SSBOND   6 CYS B  293    CYS B  311                          1555   1555  2.07
LINK         NE2 HIS A 224                NI    NI A 401     1555   1555  2.35
LINK         SG  CYS B 105                NI    NI B 401     1555   1555  2.58
LINK         NE2 HIS B 224                NI    NI B 401     1555   1555  2.34
LINK        NI    NI A 401                 O   HOH A 525     1555   1555  2.66
LINK        NI    NI B 401                 O   HOH B 525     1555   1555  2.77
CISPEP   1 PRO A   69    PRO A   70          0       -11.07
CISPEP   2 GLN A  191    PRO A  192          0         3.33
CISPEP   3 PRO B   69    PRO B   70          0        -9.25
CISPEP   4 GLN B  191    PRO B  192          0         3.01
SITE     1 AC1  3 CYS A 105  HIS A 224  HOH A 525
SITE     1 AC2  4 ASP A  17  LEU A  20  PRO A  27  THR A  89
SITE     1 AC3  3 THR A 116  ARG A 122  ALA A 170
SITE     1 AC4  3 CYS B 105  HIS B 224  HOH B 525
SITE     1 AC5  5 LYS B  32  ARG B 127  ARG B 242  SER B 243
SITE     2 AC5  5 THR B 244
SITE     1 AC6  2 VAL B 194  GLN B 211
SITE     1 AC7  2 ARG B 122  ALA B 170
CRYST1   52.288   44.559  132.947  90.00  89.41  90.00 P 1 21 1      4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.019125  0.000000 -0.000197        0.00000
SCALE2      0.000000  0.022442  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007522        0.00000
TER    2319      PRO A 317
TER    4627      PRO B 317
MASTER      331    0    7   33   18    0    8    6 4720    2   66   50
END