longtext: 6o8v-pdb

content
HEADER    HYDROLASE                               12-MAR-19   6O8V
TITLE     THE STRUCTURE OF LIPASE FROM THERMOMYCES LANUGINOSA IN COMPLEX WITH 1,
TITLE    2 3 DIACYLGLYCEROL: RHOMBOHEDRAL CRYSTAL FORM
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LIPASE;
COMPND   3 CHAIN: A;
COMPND   4 SYNONYM: TRIACYLGLYCEROL LIPASE;
COMPND   5 EC: 3.1.1.3;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: THERMOMYCES LANUGINOSUS;
SOURCE   3 ORGANISM_TAXID: 5541;
SOURCE   4 GENE: LIP;
SOURCE   5 EXPRESSION_SYSTEM: THERMOACTINOMYCES VULGARIS;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 2026
KEYWDS    TRIMER, ESTERASE, LIPIDS, LIPASE, SUBSTRATE COMPLEX, CATALYSIS, ACYL
KEYWDS   2 INTERMEDIATE, LIPID BINDING PROTEIN, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    A.MCPHERSON
REVDAT   1   01-MAY-24 6O8V    0
JRNL        AUTH   A.MCPHERSON,B.S.LARSON,A.KALASKY
JRNL        TITL   THE CRYSTAL STRUCTURES OF THERMOMYCES (HUMICOLA) LANUGINOSA
JRNL        TITL 2 LIPASE IN COMPLEX WITH ENZYMATIC REACTANTS
JRNL        REF    CURR ENZYM INHIB              V.  16   199 2020
JRNL        REFN                   ESSN 1875-6662
JRNL        DOI    HTTP://DX.DOI.ORG/10.2174/1573408016999200511090910
REMARK   2
REMARK   2 RESOLUTION.    1.30 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.19RC7_4070
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : GEOSTD + MONOMER LIBRARY + CDL V1.2
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.30
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 80.52
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.370
REMARK   3   COMPLETENESS FOR RANGE        (%) : 87.9
REMARK   3   NUMBER OF REFLECTIONS             : 58352
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.111
REMARK   3   R VALUE            (WORKING SET) : 0.109
REMARK   3   FREE R VALUE                     : 0.141
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.910
REMARK   3   FREE R VALUE TEST SET COUNT      : 2865
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 80.5200 -  3.5400    1.00     3334   195  0.1442 0.1555
REMARK   3     2  3.5400 -  2.8100    1.00     3222   178  0.1171 0.1438
REMARK   3     3  2.8100 -  2.4600    1.00     3201   154  0.1133 0.1301
REMARK   3     4  2.4600 -  2.2300    1.00     3196   154  0.0940 0.1136
REMARK   3     5  2.2300 -  2.0700    1.00     3131   192  0.0897 0.1371
REMARK   3     6  2.0700 -  1.9500    1.00     3176   148  0.0948 0.1294
REMARK   3     7  1.9500 -  1.8500    1.00     3178   150  0.0897 0.1311
REMARK   3     8  1.8500 -  1.7700    1.00     3119   176  0.0895 0.1413
REMARK   3     9  1.7700 -  1.7000    1.00     3142   157  0.1004 0.1447
REMARK   3    10  1.7000 -  1.6400    1.00     3181   139  0.1182 0.1804
REMARK   3    11  1.6400 -  1.5900    1.00     3095   165  0.0973 0.1477
REMARK   3    12  1.5900 -  1.5500    1.00     3189   132  0.0877 0.1313
REMARK   3    13  1.5500 -  1.5100    0.99     3079   143  0.0861 0.1272
REMARK   3    14  1.5100 -  1.4700    0.96     3015   159  0.0786 0.1320
REMARK   3    15  1.4700 -  1.4400    0.92     2831   193  0.0770 0.1243
REMARK   3    16  1.4400 -  1.4100    0.86     2686   130  0.0822 0.1455
REMARK   3    17  1.4100 -  1.3800    0.74     2327   103  0.0949 0.1720
REMARK   3    18  1.3800 -  1.3500    0.57     1748   115  0.1201 0.1649
REMARK   3    19  1.3500 -  1.3300    0.38     1178    68  0.1546 0.1830
REMARK   3    20  1.3300 -  1.3000    0.14      459    14  0.2164 0.3480
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.093
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 13.226
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 13.90
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 20.61
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.010           2230
REMARK   3   ANGLE     :  1.155           3035
REMARK   3   CHIRALITY :  0.093            326
REMARK   3   PLANARITY :  0.018            390
REMARK   3   DIHEDRAL  : 11.383            826
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 6O8V COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 13-MAR-19.
REMARK 100 THE DEPOSITION ID IS D_1000240208.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 15-FEB-18
REMARK 200  TEMPERATURE           (KELVIN) : 173
REMARK 200  PH                             : 6.0 - 7.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ALS
REMARK 200  BEAMLINE                       : 8.3.1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.03
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 300K
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : AIMLESS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 58358
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.300
REMARK 200  RESOLUTION RANGE LOW       (A) : 100.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 88.0
REMARK 200  DATA REDUNDANCY                : 31.00
REMARK 200  R MERGE                    (I) : 0.04900
REMARK 200  R SYM                      (I) : 0.04600
REMARK 200   FOR THE DATA SET  : 42.7000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.30
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.33
REMARK 200  COMPLETENESS FOR SHELL     (%) : 14.4
REMARK 200  DATA REDUNDANCY IN SHELL       : 4.20
REMARK 200  R MERGE FOR SHELL          (I) : 0.36000
REMARK 200  R SYM FOR SHELL            (I) : 0.34000
REMARK 200   FOR SHELL         : 2.600
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: 1TIB
REMARK 200
REMARK 200 REMARK: THIN NEEDLES OF INDETERMINATE CROSS SECTION
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 42.23
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.13
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: FROM A FILTERED, CRUDE NUTRIENT BROTH
REMARK 280  CONTAINING 30 - 50 MG/ML LIPASE PLUS 25% PEG 3350 PLUS 0.10 M
REMARK 280  MES BUFFER AT ROOM TEMPERATURE, PH 6.5, VAPOR DIFFUSION, SITTING
REMARK 280  DROP, TEMPERATURE 298K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: H 3 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z
REMARK 290       3555   -X+Y,-X,Z
REMARK 290       4555   Y,X,-Z
REMARK 290       5555   X-Y,-Y,-Z
REMARK 290       6555   -X,-X+Y,-Z
REMARK 290       7555   X+2/3,Y+1/3,Z+1/3
REMARK 290       8555   -Y+2/3,X-Y+1/3,Z+1/3
REMARK 290       9555   -X+Y+2/3,-X+1/3,Z+1/3
REMARK 290      10555   Y+2/3,X+1/3,-Z+1/3
REMARK 290      11555   X-Y+2/3,-Y+1/3,-Z+1/3
REMARK 290      12555   -X+2/3,-X+Y+1/3,-Z+1/3
REMARK 290      13555   X+1/3,Y+2/3,Z+2/3
REMARK 290      14555   -Y+1/3,X-Y+2/3,Z+2/3
REMARK 290      15555   -X+Y+1/3,-X+2/3,Z+2/3
REMARK 290      16555   Y+1/3,X+2/3,-Z+2/3
REMARK 290      17555   X-Y+1/3,-Y+2/3,-Z+2/3
REMARK 290      18555   -X+1/3,-X+Y+2/3,-Z+2/3
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   4 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   4  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   5  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   6 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   7  1.000000  0.000000  0.000000       38.18650
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       22.04699
REMARK 290   SMTRY3   7  0.000000  0.000000  1.000000       80.51700
REMARK 290   SMTRY1   8 -0.500000 -0.866025  0.000000       38.18650
REMARK 290   SMTRY2   8  0.866025 -0.500000  0.000000       22.04699
REMARK 290   SMTRY3   8  0.000000  0.000000  1.000000       80.51700
REMARK 290   SMTRY1   9 -0.500000  0.866025  0.000000       38.18650
REMARK 290   SMTRY2   9 -0.866025 -0.500000  0.000000       22.04699
REMARK 290   SMTRY3   9  0.000000  0.000000  1.000000       80.51700
REMARK 290   SMTRY1  10 -0.500000  0.866025  0.000000       38.18650
REMARK 290   SMTRY2  10  0.866025  0.500000  0.000000       22.04699
REMARK 290   SMTRY3  10  0.000000  0.000000 -1.000000       80.51700
REMARK 290   SMTRY1  11  1.000000  0.000000  0.000000       38.18650
REMARK 290   SMTRY2  11  0.000000 -1.000000  0.000000       22.04699
REMARK 290   SMTRY3  11  0.000000  0.000000 -1.000000       80.51700
REMARK 290   SMTRY1  12 -0.500000 -0.866025  0.000000       38.18650
REMARK 290   SMTRY2  12 -0.866025  0.500000  0.000000       22.04699
REMARK 290   SMTRY3  12  0.000000  0.000000 -1.000000       80.51700
REMARK 290   SMTRY1  13  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  13  0.000000  1.000000  0.000000       44.09397
REMARK 290   SMTRY3  13  0.000000  0.000000  1.000000      161.03400
REMARK 290   SMTRY1  14 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2  14  0.866025 -0.500000  0.000000       44.09397
REMARK 290   SMTRY3  14  0.000000  0.000000  1.000000      161.03400
REMARK 290   SMTRY1  15 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2  15 -0.866025 -0.500000  0.000000       44.09397
REMARK 290   SMTRY3  15  0.000000  0.000000  1.000000      161.03400
REMARK 290   SMTRY1  16 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2  16  0.866025  0.500000  0.000000       44.09397
REMARK 290   SMTRY3  16  0.000000  0.000000 -1.000000      161.03400
REMARK 290   SMTRY1  17  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  17  0.000000 -1.000000  0.000000       44.09397
REMARK 290   SMTRY3  17  0.000000  0.000000 -1.000000      161.03400
REMARK 290   SMTRY1  18 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2  18 -0.866025  0.500000  0.000000       44.09397
REMARK 290   SMTRY3  18  0.000000  0.000000 -1.000000      161.03400
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: TRIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: TRIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 7810 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 30450 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -79.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 350   BIOMT2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 350   BIOMT3   2  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 350   BIOMT2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 350   BIOMT3   3  0.000000  0.000000  1.000000        0.00000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH A 691  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A 731  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A 780  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A 793  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A 796  LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A   -21
REMARK 465     ARG A   -20
REMARK 465     SER A   -19
REMARK 465     SER A   -18
REMARK 465     LEU A   -17
REMARK 465     VAL A   -16
REMARK 465     LEU A   -15
REMARK 465     PHE A   -14
REMARK 465     PHE A   -13
REMARK 465     VAL A   -12
REMARK 465     SER A   -11
REMARK 465     ALA A   -10
REMARK 465     TRP A    -9
REMARK 465     THR A    -8
REMARK 465     ALA A    -7
REMARK 465     LEU A    -6
REMARK 465     ALA A    -5
REMARK 465     SER A    -4
REMARK 465     PRO A    -3
REMARK 465     ILE A    -2
REMARK 465     ARG A    -1
REMARK 465     ARG A     0
REMARK 465     GLY A   245
REMARK 465     GLY A   246
REMARK 465     ASN A   247
REMARK 465     ASN A   248
REMARK 465     GLN A   249
REMARK 465     PRO A   250
REMARK 465     ASN A   251
REMARK 465     ILE A   252
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     ASP A 242    CG   OD1  OD2
REMARK 480
REMARK 480 ZERO OCCUPANCY ATOM
REMARK 480 THE FOLLOWING RESIDUES HAVE ATOMS MODELED WITH ZERO
REMARK 480 OCCUPANCY. THE LOCATION AND PROPERTIES OF THESE ATOMS
REMARK 480 MAY NOT BE RELIABLE. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 480 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 480   M RES C SSEQI ATOMS
REMARK 480     ILE A  202   CB   CG1  CG2  CD1
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   HG   SER A   146     O    HOH A   404              1.32
REMARK 500  HD22  ASN A   162     O    HOH A   408              1.51
REMARK 500   OD1  ASN A    94     O    HOH A   401              1.55
REMARK 500   OG1  THR A   244     O    HOH A   402              1.79
REMARK 500   O    HOH A   402     O    HOH A   465              1.90
REMARK 500   OD1  ASP A   102     O    HOH A   403              1.92
REMARK 500   OG   SER A   146     O    HOH A   404              1.93
REMARK 500   O    HOH A   427     O    HOH A   566              1.99
REMARK 500   OD1  ASP A   102     O    HOH A   405              2.02
REMARK 500   O    HOH A   493     O    HOH A   705              2.05
REMARK 500   O    HOH A   485     O    HOH A   683              2.06
REMARK 500   O    HOH A   639     O    HOH A   740              2.09
REMARK 500   O    HOH A   427     O    HOH A   493              2.10
REMARK 500   O    HOH A   405     O    HOH A   664              2.11
REMARK 500   O    HOH A   422     O    HOH A   601              2.11
REMARK 500   O    HOH A   508     O    HOH A   721              2.12
REMARK 500   O    HOH A   428     O    HOH A   675              2.12
REMARK 500   O    ALA A    47     O    HOH A   406              2.18
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O    HOH A   740     O    HOH A   795     3555     2.03
REMARK 500   O    HOH A   736     O    HOH A   794     3555     2.10
REMARK 500   O    HOH A   749     O    HOH A   769    11445     2.13
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    CYS A  36     -155.57   -123.01
REMARK 500    CYS A  41       54.80   -149.78
REMARK 500    ARG A  84      -46.56   -137.63
REMARK 500    SER A 146     -129.19     60.53
REMARK 500    SER A 146     -129.22     60.83
REMARK 500    ASP A 242      -89.04   -179.40
REMARK 500    ALA A 243     -110.75    -98.90
REMARK 500    PHE A 262      -41.14     74.42
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: PLANAR GROUPS
REMARK 500
REMARK 500 PLANAR GROUPS IN THE FOLLOWING RESIDUES HAVE A TOTAL
REMARK 500 RMS DISTANCE OF ALL ATOMS FROM THE BEST-FIT PLANE
REMARK 500 BY MORE THAN AN EXPECTED VALUE OF 6*RMSD, WITH AN
REMARK 500 RMSD 0.02 ANGSTROMS, OR AT LEAST ONE ATOM HAS
REMARK 500 AN RMSD GREATER THAN THIS VALUE
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500  M RES CSSEQI        RMS     TYPE
REMARK 500    ARG A 205         0.14    SIDE CHAIN
REMARK 500    ARG A 232         0.08    SIDE CHAIN
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 785        DISTANCE =  5.81 ANGSTROMS
REMARK 525    HOH A 787        DISTANCE =  5.88 ANGSTROMS
REMARK 525    HOH A 788        DISTANCE =  6.01 ANGSTROMS
REMARK 525    HOH A 789        DISTANCE =  6.07 ANGSTROMS
REMARK 525    HOH A 790        DISTANCE =  6.14 ANGSTROMS
REMARK 525    HOH A 791        DISTANCE =  6.45 ANGSTROMS
REMARK 525    HOH A 792        DISTANCE =  6.55 ANGSTROMS
REMARK 525    HOH A 793        DISTANCE =  6.57 ANGSTROMS
REMARK 525    HOH A 794        DISTANCE =  6.89 ANGSTROMS
REMARK 525    HOH A 795        DISTANCE =  6.97 ANGSTROMS
REMARK 525    HOH A 796        DISTANCE =  7.19 ANGSTROMS
REMARK 525    HOH A 797        DISTANCE =  7.21 ANGSTROMS
REMARK 525    HOH A 798        DISTANCE =  7.78 ANGSTROMS
REMARK 525    HOH A 799        DISTANCE =  7.80 ANGSTROMS
REMARK 525    HOH A 800        DISTANCE =  7.88 ANGSTROMS
REMARK 525    HOH A 801        DISTANCE =  8.87 ANGSTROMS
REMARK 525    HOH A 802        DISTANCE =  9.20 ANGSTROMS
REMARK 525    HOH A 803        DISTANCE = 12.27 ANGSTROMS
REMARK 525    HOH A 804        DISTANCE = 13.95 ANGSTROMS
REMARK 525    HOH A 805        DISTANCE = 15.13 ANGSTROMS
REMARK 615
REMARK 615 ZERO OCCUPANCY ATOM
REMARK 615 THE FOLLOWING RESIDUES HAVE ATOMS MODELED WITH ZERO
REMARK 615 OCCUPANCY. THE LOCATION AND PROPERTIES OF THESE ATOMS
REMARK 615 MAY NOT BE RELIABLE. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 615 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 615   M RES C  SSEQI
REMARK 615     HOH A   459
DBREF  6O8V A  -21   269  UNP    O59952   LIP_THELA        1    291
SEQADV 6O8V LEU A   86  UNP  O59952    ILE   108 CONFLICT
SEQRES   1 A  291  MET ARG SER SER LEU VAL LEU PHE PHE VAL SER ALA TRP
SEQRES   2 A  291  THR ALA LEU ALA SER PRO ILE ARG ARG GLU VAL SER GLN
SEQRES   3 A  291  ASP LEU PHE ASN GLN PHE ASN LEU PHE ALA GLN TYR SER
SEQRES   4 A  291  ALA ALA ALA TYR CYS GLY LYS ASN ASN ASP ALA PRO ALA
SEQRES   5 A  291  GLY THR ASN ILE THR CYS THR GLY ASN ALA CYS PRO GLU
SEQRES   6 A  291  VAL GLU LYS ALA ASP ALA THR PHE LEU TYR SER PHE GLU
SEQRES   7 A  291  ASP SER GLY VAL GLY ASP VAL THR GLY PHE LEU ALA LEU
SEQRES   8 A  291  ASP ASN THR ASN LYS LEU ILE VAL LEU SER PHE ARG GLY
SEQRES   9 A  291  SER ARG SER LEU GLU ASN TRP ILE GLY ASN LEU ASN PHE
SEQRES  10 A  291  ASP LEU LYS GLU ILE ASN ASP ILE CYS SER GLY CYS ARG
SEQRES  11 A  291  GLY HIS ASP GLY PHE THR SER SER TRP ARG SER VAL ALA
SEQRES  12 A  291  ASP THR LEU ARG GLN LYS VAL GLU ASP ALA VAL ARG GLU
SEQRES  13 A  291  HIS PRO ASP TYR ARG VAL VAL PHE THR GLY HIS SER LEU
SEQRES  14 A  291  GLY GLY ALA LEU ALA THR VAL ALA GLY ALA ASP LEU ARG
SEQRES  15 A  291  GLY ASN GLY TYR ASP ILE ASP VAL PHE SER TYR GLY ALA
SEQRES  16 A  291  PRO ARG VAL GLY ASN ARG ALA PHE ALA GLU PHE LEU THR
SEQRES  17 A  291  VAL GLN THR GLY GLY THR LEU TYR ARG ILE THR HIS THR
SEQRES  18 A  291  ASN ASP ILE VAL PRO ARG LEU PRO PRO ARG GLU PHE GLY
SEQRES  19 A  291  TYR SER HIS SER SER PRO GLU TYR TRP ILE LYS SER GLY
SEQRES  20 A  291  THR LEU VAL PRO VAL THR ARG ASN ASP ILE VAL LYS ILE
SEQRES  21 A  291  GLU GLY ILE ASP ALA THR GLY GLY ASN ASN GLN PRO ASN
SEQRES  22 A  291  ILE PRO ASP ILE PRO ALA HIS LEU TRP TYR PHE GLY LEU
SEQRES  23 A  291  ILE GLY THR CYS LEU
HET    NAG  A 301      27
HET    PG4  A 302      27
HET    PO4  A 303       5
HET    PO4  A 304       5
HET    PO4  A 305       5
HET    PO4  A 306       5
HET    PO4  A 307       5
HET    LTV  A 308      51
HETNAM     NAG 2-ACETAMIDO-2-DEOXY-BETA-D-GLUCOPYRANOSE
HETNAM     PG4 TETRAETHYLENE GLYCOL
HETNAM     PO4 PHOSPHATE ION
HETNAM     LTV 2-HYDROXY-3-(OCTADECANOYLOXY)PROPYL PENTACOSANOATE
HETSYN     NAG N-ACETYL-BETA-D-GLUCOSAMINE; 2-ACETAMIDO-2-DEOXY-BETA-
HETSYN   2 NAG  D-GLUCOSE; 2-ACETAMIDO-2-DEOXY-D-GLUCOSE; 2-ACETAMIDO-
HETSYN   3 NAG  2-DEOXY-GLUCOSE; N-ACETYL-D-GLUCOSAMINE
FORMUL   2  NAG    C8 H15 N O6
FORMUL   3  PG4    C8 H18 O5
FORMUL   4  PO4    5(O4 P 3-)
FORMUL   9  LTV    C46 H90 O5
FORMUL  10  HOH   *405(H2 O)
HELIX    1 AA1 SER A    3  ALA A   20  1                                  18
HELIX    2 AA2 TYR A   21  ASN A   26  5                                   6
HELIX    3 AA3 CYS A   41  ALA A   47  1                                   7
HELIX    4 AA4 SER A   85  LEU A   93  1                                   9
HELIX    5 AA5 ASP A  111  HIS A  135  1                                  25
HELIX    6 AA6 SER A  146  ARG A  160  1                                  15
HELIX    7 AA7 ASN A  178  GLN A  188  1                                  11
HELIX    8 AA8 ILE A  202  LEU A  206  5                                   5
HELIX    9 AA9 PRO A  208  GLY A  212  5                                   5
HELIX   10 AB1 THR A  231  ASN A  233  5                                   3
HELIX   11 AB2 ASP A  254  TRP A  260  1                                   7
SHEET    1 AA1 8 ALA A  49  SER A  58  0
SHEET    2 AA1 8 VAL A  63  ASP A  70 -1  O  LEU A  67   N  TYR A  53
SHEET    3 AA1 8 LEU A  75  PHE A  80 -1  O  VAL A  77   N  ALA A  68
SHEET    4 AA1 8 ARG A 139  HIS A 145  1  O  THR A 143   N  LEU A  78
SHEET    5 AA1 8 ILE A 166  TYR A 171  1  O  ASP A 167   N  PHE A 142
SHEET    6 AA1 8 LEU A 193  HIS A 198  1  O  TYR A 194   N  VAL A 168
SHEET    7 AA1 8 GLU A 219  ILE A 222  1  O  ILE A 222   N  THR A 197
SHEET    8 AA1 8 ILE A 235  ILE A 238 -1  O  ILE A 238   N  GLU A 219
SHEET    1 AA2 2 LEU A  97  GLU A  99  0
SHEET    2 AA2 2 ARG A 108  HIS A 110 -1  O  GLY A 109   N  LYS A  98
SSBOND   1 CYS A   22    CYS A  268                          1555   1555  2.11
SSBOND   2 CYS A   36    CYS A   41                          1555   1555  2.04
SSBOND   3 CYS A  104    CYS A  107                          1555   1555  2.07
LINK         ND2 ASN A  33                 C1  NAG A 301     1555   1555  1.43
CISPEP   1 LEU A  206    PRO A  207          0       -17.05
CISPEP   2 SER A  217    PRO A  218          0        -0.60
CRYST1   76.373   76.373  241.551  90.00  90.00 120.00 H 3 2        18
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.013094  0.007560  0.000000        0.00000
SCALE2      0.000000  0.015119  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004140        0.00000
TER    4048      LEU A 269
MASTER      469    0    8   11   10    0    0    6 2520    1  137   23
END