longtext: 6qgq-pdb

content
HEADER    HYDROLASE                               12-JAN-19   6QGQ
TITLE     CRYSTAL STRUCTURE OF APT1 C2S MUTANT BOUND TO PALMITIC ACID.
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ACYL-PROTEIN THIOESTERASE 1;
COMPND   3 CHAIN: A, C, D;
COMPND   4 SYNONYM: HAPT1,LYSOPHOSPHOLIPASE 1,LYSOPHOSPHOLIPASE I,LYSOPLA I;
COMPND   5 EC: 3.1.2.-;
COMPND   6 ENGINEERED: YES;
COMPND   7 MUTATION: YES;
COMPND   8 MOL_ID: 2;
COMPND   9 MOLECULE: ACYL-PROTEIN THIOESTERASE 1;
COMPND  10 CHAIN: B;
COMPND  11 SYNONYM: HAPT1,LYSOPHOSPHOLIPASE 1,LYSOPHOSPHOLIPASE I,LYSOPLA I;
COMPND  12 EC: 3.1.2.-;
COMPND  13 ENGINEERED: YES;
COMPND  14 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: LYPLA1, APT1, LPL1;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   8 MOL_ID: 2;
SOURCE   9 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE  10 ORGANISM_COMMON: HUMAN;
SOURCE  11 ORGANISM_TAXID: 9606;
SOURCE  12 GENE: LYPLA1, APT1, LPL1;
SOURCE  13 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE  14 EXPRESSION_SYSTEM_TAXID: 562
KEYWDS    ACYL-PROTEIN THIOESTERASE, PALMITIC ACID, DEPALMITOYLATION, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.AUDAGNOTTO,M.J.MARCAIDA,S.HO,F.POJER,G.VAN DER GOOT,M.DAL PERARO
REVDAT   3   31-MAR-21 6QGQ    1       JRNL
REVDAT   2   17-MAR-21 6QGQ    1       JRNL
REVDAT   1   05-FEB-20 6QGQ    0
JRNL        AUTH   L.ABRAMI,M.AUDAGNOTTO,S.HO,M.J.MARCAIDA,F.S.MESQUITA,
JRNL        AUTH 2 M.U.ANWAR,P.A.SANDOZ,G.FONTI,F.POJER,M.DAL PERARO,
JRNL        AUTH 3 F.G.VAN DER GOOT
JRNL        TITL   PALMITOYLATED ACYL PROTEIN THIOESTERASE APT2 DEFORMS
JRNL        TITL 2 MEMBRANES TO EXTRACT SUBSTRATE ACYL CHAINS.
JRNL        REF    NAT.CHEM.BIOL.                             2021
JRNL        REFN                   ESSN 1552-4469
JRNL        PMID   33707782
JRNL        DOI    10.1038/S41589-021-00753-2
REMARK   2
REMARK   2 RESOLUTION.    2.60 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (1.14_3260)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.60
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 46.62
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.340
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.9
REMARK   3   NUMBER OF REFLECTIONS             : 30461
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.193
REMARK   3   R VALUE            (WORKING SET) : 0.191
REMARK   3   FREE R VALUE                     : 0.238
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.860
REMARK   3   FREE R VALUE TEST SET COUNT      : 1481
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 46.6263 -  5.7822    1.00     2836   133  0.1884 0.2263
REMARK   3     2  5.7822 -  4.5908    1.00     2683   146  0.1719 0.2338
REMARK   3     3  4.5908 -  4.0109    1.00     2673   129  0.1551 0.1810
REMARK   3     4  4.0109 -  3.6443    1.00     2653   121  0.1762 0.2184
REMARK   3     5  3.6443 -  3.3832    1.00     2565   163  0.1853 0.2267
REMARK   3     6  3.3832 -  3.1838    1.00     2651   118  0.2057 0.2429
REMARK   3     7  3.1838 -  3.0244    1.00     2608   128  0.2205 0.2824
REMARK   3     8  3.0244 -  2.8927    1.00     2534   148  0.2313 0.2894
REMARK   3     9  2.8927 -  2.7814    1.00     2614   149  0.2295 0.2994
REMARK   3    10  2.7814 -  2.6854    1.00     2572   117  0.2303 0.3104
REMARK   3    11  2.6854 -  2.6015    0.99     2591   129  0.2258 0.2432
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : NULL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.260
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 21.470
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.003           6883
REMARK   3   ANGLE     :  0.541           9343
REMARK   3   CHIRALITY :  0.042           1065
REMARK   3   PLANARITY :  0.004           1205
REMARK   3   DIHEDRAL  :  6.818           4162
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 6QGQ COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 13-JAN-19.
REMARK 100 THE DEPOSITION ID IS D_1292100145.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 27-JUN-16
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SLS
REMARK 200  BEAMLINE                       : X06DA
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 2M-F
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 30525
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.600
REMARK 200  RESOLUTION RANGE LOW       (A) : 48.710
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.9
REMARK 200  DATA REDUNDANCY                : 7.900
REMARK 200  R MERGE                    (I) : 0.18000
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 9.6000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.60
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.74
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: 1FJ2
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 50.90
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.51
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 25% OF OF POLYETHYLENE GLYCOL (PEG)
REMARK 280  2000 MME, 0.3 M SODIUM ACETATE, 0.1 M SODIUM CACODYLATE PH 6.5;,
REMARK 280  VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 291K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z
REMARK 290       3555   -X+1/2,Y+1/2,-Z
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000       73.06900
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       80.36900
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       73.06900
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       80.36900
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2, 3, 4
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 3
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 4
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLY A    -2
REMARK 465     SER A    -1
REMARK 465     ARG A     0
REMARK 465     MET A     1
REMARK 465     SER A     2
REMARK 465     GLY A     3
REMARK 465     ASN A     4
REMARK 465     ASN A     5
REMARK 465     MET A     6
REMARK 465     SER A     7
REMARK 465     GLY B    -2
REMARK 465     SER B    -1
REMARK 465     ARG B     0
REMARK 465     MET B     1
REMARK 465     SER B     2
REMARK 465     GLY B     3
REMARK 465     ASN B     4
REMARK 465     ASN B     5
REMARK 465     MET B     6
REMARK 465     SER B     7
REMARK 465     THR B     8
REMARK 465     GLY C    -2
REMARK 465     SER C    -1
REMARK 465     ARG C     0
REMARK 465     MET C     1
REMARK 465     SER C     2
REMARK 465     GLY C     3
REMARK 465     ASN C     4
REMARK 465     ASN C     5
REMARK 465     MET C     6
REMARK 465     SER C     7
REMARK 465     THR C     8
REMARK 465     ASP C   230
REMARK 465     GLY D    -2
REMARK 465     SER D    -1
REMARK 465     ARG D     0
REMARK 465     MET D     1
REMARK 465     SER D     2
REMARK 465     GLY D     3
REMARK 465     ASN D     4
REMARK 465     ASN D     5
REMARK 465     MET D     6
REMARK 465     SER D     7
REMARK 465     THR D     8
REMARK 465     ASP D   230
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OD2  ASP C   174     O    HOH C   701              1.87
REMARK 500   OG   SER D   143     O    HOH D   401              1.93
REMARK 500   O    CYS A   169     O    HOH A   401              1.94
REMARK 500   O    ASP B    74     O    HOH B   501              2.01
REMARK 500   NE2  GLN B   102     O    HOH B   502              2.01
REMARK 500   NZ   LYS B    50     O    HOH B   503              2.03
REMARK 500   CA   GLY B   159     O    HOH B   510              2.04
REMARK 500   O2   GOL D   301     O    HOH D   402              2.05
REMARK 500   O    HOH C   733     O    HOH C   737              2.09
REMARK 500   ND2  ASN A   106     O    HOH A   402              2.10
REMARK 500   OD1  ASN D   161     O    HOH D   403              2.12
REMARK 500   O    HOH B   531     O    HOH B   532              2.13
REMARK 500   O    LEU C   148     OG   SER C   151              2.15
REMARK 500   OE2  GLU C    84     O    HOH C   702              2.15
REMARK 500   OE1  GLU B   103     O    HOH B   502              2.18
REMARK 500   O    THR B    33     O    HOH B   504              2.18
REMARK 500   O    PRO D   228     O    HOH D   404              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O    HOH A   406     O    HOH C   703     4456     1.76
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ARG A  18      -75.47   -127.93
REMARK 500    ILE A  44       27.50   -144.14
REMARK 500    ASN A  64       58.30   -117.84
REMARK 500    SER A 119     -126.98     56.03
REMARK 500    ASN A 161       30.22    -99.95
REMARK 500    MET A 207     -151.50   -103.13
REMARK 500    ARG B  18     -109.08   -119.67
REMARK 500    SER B 119     -125.53     54.65
REMARK 500    CYS B 144     -161.35   -129.99
REMARK 500    SER B 151       -5.54    -57.37
REMARK 500    LYS B 190        1.35    -67.36
REMARK 500    MET B 207     -137.86    -93.68
REMARK 500    SER B 209     -147.56   -166.40
REMARK 500    ASP B 230       -5.64   -141.74
REMARK 500    ARG C  18      -99.10   -104.47
REMARK 500    ASN C  64       62.42   -114.85
REMARK 500    ASN C 111        0.86    -69.12
REMARK 500    SER C 119     -128.28     52.55
REMARK 500    ARG C 162      -57.55    -26.11
REMARK 500    PRO C 178      152.02    -48.95
REMARK 500    MET C 207     -149.99   -101.44
REMARK 500    ARG D  18     -156.44    -86.47
REMARK 500    LYS D 105        4.19    -68.65
REMARK 500    SER D 119     -123.21     55.91
REMARK 500    CYS D 144     -159.68   -129.28
REMARK 500    ASN D 194      109.14    -54.89
REMARK 500    MET D 207     -142.69    -92.91
REMARK 500    SER D 209     -155.40   -158.93
REMARK 500
REMARK 500 REMARK: NULL
REMARK 610
REMARK 610 MISSING HETEROATOM
REMARK 610 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 610 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 610 I=INSERTION CODE):
REMARK 610   M RES C SSEQI
REMARK 610     PLM D  302
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL A 301
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL A 302
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue PLM B 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL C 601
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL D 301
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue PLM D 302
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 6QGO   RELATED DB: PDB
REMARK 900 RELATED ID: 6QGN   RELATED DB: PDB
DBREF  6QGQ A    1   230  UNP    O75608   LYPA1_HUMAN      1    230
DBREF  6QGQ B    1   230  UNP    O75608   LYPA1_HUMAN      1    230
DBREF  6QGQ C    1   230  UNP    O75608   LYPA1_HUMAN      1    230
DBREF  6QGQ D    1   230  UNP    O75608   LYPA1_HUMAN      1    230
SEQADV 6QGQ GLY A   -2  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ SER A   -1  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ ARG A    0  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ SER A    2  UNP  O75608    CYS     2 ENGINEERED MUTATION
SEQADV 6QGQ GLY B   -2  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ SER B   -1  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ ARG B    0  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ SER B    2  UNP  O75608    CYS     2 ENGINEERED MUTATION
SEQADV 6QGQ ALA B  231  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ GLY C   -2  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ SER C   -1  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ ARG C    0  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ SER C    2  UNP  O75608    CYS     2 ENGINEERED MUTATION
SEQADV 6QGQ GLY D   -2  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ SER D   -1  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ ARG D    0  UNP  O75608              EXPRESSION TAG
SEQADV 6QGQ SER D    2  UNP  O75608    CYS     2 ENGINEERED MUTATION
SEQRES   1 A  233  GLY SER ARG MET SER GLY ASN ASN MET SER THR PRO LEU
SEQRES   2 A  233  PRO ALA ILE VAL PRO ALA ALA ARG LYS ALA THR ALA ALA
SEQRES   3 A  233  VAL ILE PHE LEU HIS GLY LEU GLY ASP THR GLY HIS GLY
SEQRES   4 A  233  TRP ALA GLU ALA PHE ALA GLY ILE ARG SER SER HIS ILE
SEQRES   5 A  233  LYS TYR ILE CYS PRO HIS ALA PRO VAL ARG PRO VAL THR
SEQRES   6 A  233  LEU ASN MET ASN VAL ALA MET PRO SER TRP PHE ASP ILE
SEQRES   7 A  233  ILE GLY LEU SER PRO ASP SER GLN GLU ASP GLU SER GLY
SEQRES   8 A  233  ILE LYS GLN ALA ALA GLU ASN ILE LYS ALA LEU ILE ASP
SEQRES   9 A  233  GLN GLU VAL LYS ASN GLY ILE PRO SER ASN ARG ILE ILE
SEQRES  10 A  233  LEU GLY GLY PHE SER GLN GLY GLY ALA LEU SER LEU TYR
SEQRES  11 A  233  THR ALA LEU THR THR GLN GLN LYS LEU ALA GLY VAL THR
SEQRES  12 A  233  ALA LEU SER CYS TRP LEU PRO LEU ARG ALA SER PHE PRO
SEQRES  13 A  233  GLN GLY PRO ILE GLY GLY ALA ASN ARG ASP ILE SER ILE
SEQRES  14 A  233  LEU GLN CYS HIS GLY ASP CYS ASP PRO LEU VAL PRO LEU
SEQRES  15 A  233  MET PHE GLY SER LEU THR VAL GLU LYS LEU LYS THR LEU
SEQRES  16 A  233  VAL ASN PRO ALA ASN VAL THR PHE LYS THR TYR GLU GLY
SEQRES  17 A  233  MET MET HIS SER SER CYS GLN GLN GLU MET MET ASP VAL
SEQRES  18 A  233  LYS GLN PHE ILE ASP LYS LEU LEU PRO PRO ILE ASP
SEQRES   1 B  234  GLY SER ARG MET SER GLY ASN ASN MET SER THR PRO LEU
SEQRES   2 B  234  PRO ALA ILE VAL PRO ALA ALA ARG LYS ALA THR ALA ALA
SEQRES   3 B  234  VAL ILE PHE LEU HIS GLY LEU GLY ASP THR GLY HIS GLY
SEQRES   4 B  234  TRP ALA GLU ALA PHE ALA GLY ILE ARG SER SER HIS ILE
SEQRES   5 B  234  LYS TYR ILE CYS PRO HIS ALA PRO VAL ARG PRO VAL THR
SEQRES   6 B  234  LEU ASN MET ASN VAL ALA MET PRO SER TRP PHE ASP ILE
SEQRES   7 B  234  ILE GLY LEU SER PRO ASP SER GLN GLU ASP GLU SER GLY
SEQRES   8 B  234  ILE LYS GLN ALA ALA GLU ASN ILE LYS ALA LEU ILE ASP
SEQRES   9 B  234  GLN GLU VAL LYS ASN GLY ILE PRO SER ASN ARG ILE ILE
SEQRES  10 B  234  LEU GLY GLY PHE SER GLN GLY GLY ALA LEU SER LEU TYR
SEQRES  11 B  234  THR ALA LEU THR THR GLN GLN LYS LEU ALA GLY VAL THR
SEQRES  12 B  234  ALA LEU SER CYS TRP LEU PRO LEU ARG ALA SER PHE PRO
SEQRES  13 B  234  GLN GLY PRO ILE GLY GLY ALA ASN ARG ASP ILE SER ILE
SEQRES  14 B  234  LEU GLN CYS HIS GLY ASP CYS ASP PRO LEU VAL PRO LEU
SEQRES  15 B  234  MET PHE GLY SER LEU THR VAL GLU LYS LEU LYS THR LEU
SEQRES  16 B  234  VAL ASN PRO ALA ASN VAL THR PHE LYS THR TYR GLU GLY
SEQRES  17 B  234  MET MET HIS SER SER CYS GLN GLN GLU MET MET ASP VAL
SEQRES  18 B  234  LYS GLN PHE ILE ASP LYS LEU LEU PRO PRO ILE ASP ALA
SEQRES   1 C  233  GLY SER ARG MET SER GLY ASN ASN MET SER THR PRO LEU
SEQRES   2 C  233  PRO ALA ILE VAL PRO ALA ALA ARG LYS ALA THR ALA ALA
SEQRES   3 C  233  VAL ILE PHE LEU HIS GLY LEU GLY ASP THR GLY HIS GLY
SEQRES   4 C  233  TRP ALA GLU ALA PHE ALA GLY ILE ARG SER SER HIS ILE
SEQRES   5 C  233  LYS TYR ILE CYS PRO HIS ALA PRO VAL ARG PRO VAL THR
SEQRES   6 C  233  LEU ASN MET ASN VAL ALA MET PRO SER TRP PHE ASP ILE
SEQRES   7 C  233  ILE GLY LEU SER PRO ASP SER GLN GLU ASP GLU SER GLY
SEQRES   8 C  233  ILE LYS GLN ALA ALA GLU ASN ILE LYS ALA LEU ILE ASP
SEQRES   9 C  233  GLN GLU VAL LYS ASN GLY ILE PRO SER ASN ARG ILE ILE
SEQRES  10 C  233  LEU GLY GLY PHE SER GLN GLY GLY ALA LEU SER LEU TYR
SEQRES  11 C  233  THR ALA LEU THR THR GLN GLN LYS LEU ALA GLY VAL THR
SEQRES  12 C  233  ALA LEU SER CYS TRP LEU PRO LEU ARG ALA SER PHE PRO
SEQRES  13 C  233  GLN GLY PRO ILE GLY GLY ALA ASN ARG ASP ILE SER ILE
SEQRES  14 C  233  LEU GLN CYS HIS GLY ASP CYS ASP PRO LEU VAL PRO LEU
SEQRES  15 C  233  MET PHE GLY SER LEU THR VAL GLU LYS LEU LYS THR LEU
SEQRES  16 C  233  VAL ASN PRO ALA ASN VAL THR PHE LYS THR TYR GLU GLY
SEQRES  17 C  233  MET MET HIS SER SER CYS GLN GLN GLU MET MET ASP VAL
SEQRES  18 C  233  LYS GLN PHE ILE ASP LYS LEU LEU PRO PRO ILE ASP
SEQRES   1 D  233  GLY SER ARG MET SER GLY ASN ASN MET SER THR PRO LEU
SEQRES   2 D  233  PRO ALA ILE VAL PRO ALA ALA ARG LYS ALA THR ALA ALA
SEQRES   3 D  233  VAL ILE PHE LEU HIS GLY LEU GLY ASP THR GLY HIS GLY
SEQRES   4 D  233  TRP ALA GLU ALA PHE ALA GLY ILE ARG SER SER HIS ILE
SEQRES   5 D  233  LYS TYR ILE CYS PRO HIS ALA PRO VAL ARG PRO VAL THR
SEQRES   6 D  233  LEU ASN MET ASN VAL ALA MET PRO SER TRP PHE ASP ILE
SEQRES   7 D  233  ILE GLY LEU SER PRO ASP SER GLN GLU ASP GLU SER GLY
SEQRES   8 D  233  ILE LYS GLN ALA ALA GLU ASN ILE LYS ALA LEU ILE ASP
SEQRES   9 D  233  GLN GLU VAL LYS ASN GLY ILE PRO SER ASN ARG ILE ILE
SEQRES  10 D  233  LEU GLY GLY PHE SER GLN GLY GLY ALA LEU SER LEU TYR
SEQRES  11 D  233  THR ALA LEU THR THR GLN GLN LYS LEU ALA GLY VAL THR
SEQRES  12 D  233  ALA LEU SER CYS TRP LEU PRO LEU ARG ALA SER PHE PRO
SEQRES  13 D  233  GLN GLY PRO ILE GLY GLY ALA ASN ARG ASP ILE SER ILE
SEQRES  14 D  233  LEU GLN CYS HIS GLY ASP CYS ASP PRO LEU VAL PRO LEU
SEQRES  15 D  233  MET PHE GLY SER LEU THR VAL GLU LYS LEU LYS THR LEU
SEQRES  16 D  233  VAL ASN PRO ALA ASN VAL THR PHE LYS THR TYR GLU GLY
SEQRES  17 D  233  MET MET HIS SER SER CYS GLN GLN GLU MET MET ASP VAL
SEQRES  18 D  233  LYS GLN PHE ILE ASP LYS LEU LEU PRO PRO ILE ASP
HET    GOL  A 301      14
HET    GOL  A 302      14
HET    PLM  B 401      49
HET    GOL  C 601      14
HET    GOL  D 301      14
HET    PLM  D 302      27
HETNAM     GOL GLYCEROL
HETNAM     PLM PALMITIC ACID
HETSYN     GOL GLYCERIN; PROPANE-1,2,3-TRIOL
FORMUL   5  GOL    4(C3 H8 O3)
FORMUL   7  PLM    2(C16 H32 O2)
FORMUL  11  HOH   *132(H2 O)
HELIX    1 AA1 GLY A   34  GLY A   43  1                                  10
HELIX    2 AA2 THR A   62  MET A   65  5                                   4
HELIX    3 AA3 ASP A   85  ASN A  106  1                                  22
HELIX    4 AA4 PRO A  109  ASN A  111  5                                   3
HELIX    5 AA5 SER A  119  LEU A  130  1                                  12
HELIX    6 AA6 LEU A  148  PHE A  152  5                                   5
HELIX    7 AA7 GLY A  159  ILE A  164  5                                   6
HELIX    8 AA8 PRO A  178  VAL A  193  1                                  16
HELIX    9 AA9 ASN A  194  ALA A  196  5                                   3
HELIX   10 AB1 CYS A  211  LEU A  226  1                                  16
HELIX   11 AB2 GLY B   34  GLY B   43  1                                  10
HELIX   12 AB3 THR B   62  MET B   65  5                                   4
HELIX   13 AB4 ASP B   85  ASN B  106  1                                  22
HELIX   14 AB5 PRO B  109  ASN B  111  5                                   3
HELIX   15 AB6 SER B  119  THR B  131  1                                  13
HELIX   16 AB7 LEU B  148  PHE B  152  5                                   5
HELIX   17 AB8 PRO B  178  VAL B  193  1                                  16
HELIX   18 AB9 ASN B  194  ALA B  196  5                                   3
HELIX   19 AC1 CYS B  211  LEU B  226  1                                  16
HELIX   20 AC2 GLY C   34  GLY C   43  1                                  10
HELIX   21 AC3 THR C   62  MET C   65  5                                   4
HELIX   22 AC4 ASP C   85  ASN C  106  1                                  22
HELIX   23 AC5 PRO C  109  ASN C  111  5                                   3
HELIX   24 AC6 SER C  119  THR C  132  1                                  14
HELIX   25 AC7 LEU C  148  PHE C  152  5                                   5
HELIX   26 AC8 PRO C  178  VAL C  193  1                                  16
HELIX   27 AC9 ASN C  194  ALA C  196  5                                   3
HELIX   28 AD1 CYS C  211  LEU C  226  1                                  16
HELIX   29 AD2 GLY D   34  GLY D   43  1                                  10
HELIX   30 AD3 THR D   62  MET D   65  5                                   4
HELIX   31 AD4 ASP D   85  LYS D  105  1                                  21
HELIX   32 AD5 PRO D  109  ASN D  111  5                                   3
HELIX   33 AD6 SER D  119  THR D  132  1                                  14
HELIX   34 AD7 LEU D  148  PHE D  152  5                                   5
HELIX   35 AD8 PRO D  178  VAL D  193  1                                  16
HELIX   36 AD9 ASN D  194  ALA D  196  5                                   3
HELIX   37 AE1 CYS D  211  LEU D  226  1                                  16
SHEET    1 AA1 7 ALA A  12  VAL A  14  0
SHEET    2 AA1 7 ILE A  49  CYS A  53 -1  O  CYS A  53   N  ALA A  12
SHEET    3 AA1 7 ALA A  22  LEU A  27  1  N  VAL A  24   O  LYS A  50
SHEET    4 AA1 7 ILE A 113  PHE A 118  1  O  ILE A 114   N  ILE A  25
SHEET    5 AA1 7 GLY A 138  LEU A 142  1  O  GLY A 138   N  LEU A 115
SHEET    6 AA1 7 ILE A 166  GLY A 171  1  O  LEU A 167   N  VAL A 139
SHEET    7 AA1 7 VAL A 198  TYR A 203  1  O  LYS A 201   N  GLN A 168
SHEET    1 AA2 2 VAL A  58  PRO A  60  0
SHEET    2 AA2 2 ALA A  68  PRO A  70 -1  O  MET A  69   N  ARG A  59
SHEET    1 AA3 7 ALA B  12  VAL B  14  0
SHEET    2 AA3 7 ILE B  49  CYS B  53 -1  O  TYR B  51   N  VAL B  14
SHEET    3 AA3 7 ALA B  22  LEU B  27  1  N  VAL B  24   O  ILE B  52
SHEET    4 AA3 7 ILE B 113  PHE B 118  1  O  PHE B 118   N  LEU B  27
SHEET    5 AA3 7 GLY B 138  LEU B 142  1  O  LEU B 142   N  GLY B 117
SHEET    6 AA3 7 SER B 165  GLY B 171  1  O  LEU B 167   N  VAL B 139
SHEET    7 AA3 7 VAL B 198  TYR B 203  1  O  THR B 199   N  GLN B 168
SHEET    1 AA4 2 VAL B  58  PRO B  60  0
SHEET    2 AA4 2 ALA B  68  PRO B  70 -1  O  MET B  69   N  ARG B  59
SHEET    1 AA5 7 ALA C  12  VAL C  14  0
SHEET    2 AA5 7 ILE C  49  CYS C  53 -1  O  CYS C  53   N  ALA C  12
SHEET    3 AA5 7 ALA C  22  LEU C  27  1  N  VAL C  24   O  ILE C  52
SHEET    4 AA5 7 ILE C 113  PHE C 118  1  O  ILE C 114   N  ALA C  23
SHEET    5 AA5 7 GLY C 138  LEU C 142  1  O  LEU C 142   N  GLY C 117
SHEET    6 AA5 7 SER C 165  GLY C 171  1  O  LEU C 167   N  VAL C 139
SHEET    7 AA5 7 VAL C 198  TYR C 203  1  O  TYR C 203   N  HIS C 170
SHEET    1 AA6 2 VAL C  58  PRO C  60  0
SHEET    2 AA6 2 ALA C  68  PRO C  70 -1  O  MET C  69   N  ARG C  59
SHEET    1 AA7 7 ALA D  12  VAL D  14  0
SHEET    2 AA7 7 ILE D  49  CYS D  53 -1  O  CYS D  53   N  ALA D  12
SHEET    3 AA7 7 ALA D  22  LEU D  27  1  N  VAL D  24   O  ILE D  52
SHEET    4 AA7 7 ILE D 113  PHE D 118  1  O  ILE D 114   N  ILE D  25
SHEET    5 AA7 7 GLY D 138  LEU D 142  1  O  LEU D 142   N  GLY D 117
SHEET    6 AA7 7 ILE D 166  GLY D 171  1  O  LEU D 167   N  VAL D 139
SHEET    7 AA7 7 VAL D 198  TYR D 203  1  O  THR D 199   N  GLN D 168
SHEET    1 AA8 2 VAL D  58  PRO D  60  0
SHEET    2 AA8 2 ALA D  68  PRO D  70 -1  O  MET D  69   N  ARG D  59
SITE     1 AC1  2 TRP A  37  MET B  65
SITE     1 AC2  3 MET A  65  HOH A 403  GLY B  36
SITE     1 AC3  9 LEU B  30  LEU B  78  SER B 119  GLN B 120
SITE     2 AC3  9 TRP B 145  ARG B 149  PHE B 181  THR B 185
SITE     3 AC3  9 HOH B 505
SITE     1 AC4  5 ASP C  32  GLY C  36  TRP C  37  SER C 210
SITE     2 AC4  5 HOH C 722
SITE     1 AC5  6 MET C  65  ASP D  32  GLY D  36  TRP D  37
SITE     2 AC5  6 SER D 210  HOH D 402
SITE     1 AC6 11 LEU D  30  ILE D  75  GLY D  77  LEU D  78
SITE     2 AC6 11 SER D  79  SER D 119  GLN D 120  TRP D 145
SITE     3 AC6 11 VAL D 177  PHE D 181  HOH D 407
CRYST1  146.138  160.738   40.679  90.00  90.00  90.00 P 21 21 2    12
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.006843  0.000000  0.000000        0.00000
SCALE2      0.000000  0.006221  0.000000        0.00000
SCALE3      0.000000  0.000000  0.024583        0.00000
TER    1679      ASP A 230
TER    3356      ALA B 231
TER    5020      ILE C 229
TER    6684      ILE D 229
MASTER      397    0    6   37   36    0   12    6 6865    4  132   72
END