longtext: 6qi2-pdb

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HEADER    HYDROLASE                               17-JAN-19   6QI2
TITLE     TIME RESOLVED STRUCTURAL ANALYSIS OF THE FULL TURNOVER OF AN ENZYME -
TITLE    2 13536 MS
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: FLUOROACETATE DEHALOGENASE;
COMPND   3 CHAIN: A, B;
COMPND   4 EC: 3.8.1.3;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: RHODOPSEUDOMONAS PALUSTRIS;
SOURCE   3 ORGANISM_TAXID: 1076;
SOURCE   4 GENE: RPA1163;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 562
KEYWDS    TIME-RESOLVED, CATALYSIS, INTERMEDIATE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    E.C.SCHULZ,P.MEHRABI,E.F.PAI,D.MILLER
REVDAT   1   25-SEP-19 6QI2    0
JRNL        AUTH   P.MEHRABI,E.C.SCHULZ,R.DSOUZA,H.M.MULLER-WERKMEISTER,
JRNL        AUTH 2 F.TELLKAMP,R.J.D.MILLER,E.F.PAI
JRNL        TITL   TIME-RESOLVED CRYSTALLOGRAPHY REVEALS ALLOSTERIC
JRNL        TITL 2 COMMUNICATION ALIGNED WITH MOLECULAR BREATHING.
JRNL        REF    SCIENCE                       V. 365  1167 2019
JRNL        REFN                   ESSN 1095-9203
JRNL        PMID   31515393
JRNL        DOI    10.1126/SCIENCE.AAW9904
REMARK   2
REMARK   2 RESOLUTION.    1.73 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (1.13_2998: ???)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.73
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 20.39
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.350
REMARK   3   COMPLETENESS FOR RANGE        (%) : 97.8
REMARK   3   NUMBER OF REFLECTIONS             : 53237
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.197
REMARK   3   R VALUE            (WORKING SET) : 0.195
REMARK   3   FREE R VALUE                     : 0.234
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.930
REMARK   3   FREE R VALUE TEST SET COUNT      : 2625
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 20.3936 -  4.5984    1.00     2762   164  0.1926 0.1905
REMARK   3     2  4.5984 -  3.6567    1.00     2785   130  0.1708 0.1946
REMARK   3     3  3.6567 -  3.1965    1.00     2712   156  0.1827 0.2066
REMARK   3     4  3.1965 -  2.9051    1.00     2728   154  0.1934 0.2459
REMARK   3     5  2.9051 -  2.6974    1.00     2738   140  0.1982 0.2484
REMARK   3     6  2.6974 -  2.5387    1.00     2706   160  0.1930 0.2494
REMARK   3     7  2.5387 -  2.4118    1.00     2715   140  0.1936 0.2130
REMARK   3     8  2.4118 -  2.3069    1.00     2720   148  0.1845 0.2278
REMARK   3     9  2.3069 -  2.2182    1.00     2759   120  0.1950 0.2857
REMARK   3    10  2.2182 -  2.1418    1.00     2715   155  0.1938 0.2337
REMARK   3    11  2.1418 -  2.0749    1.00     2658   158  0.1949 0.2862
REMARK   3    12  2.0749 -  2.0156    1.00     2764   123  0.1965 0.2206
REMARK   3    13  2.0156 -  1.9626    1.00     2680   151  0.1951 0.2269
REMARK   3    14  1.9626 -  1.9147    1.00     2748   132  0.1965 0.2707
REMARK   3    15  1.9147 -  1.8712    1.00     2733   104  0.1999 0.2543
REMARK   3    16  1.8712 -  1.8315    1.00     2732   144  0.2296 0.3016
REMARK   3    17  1.8315 -  1.7948    1.00     2717   130  0.2472 0.3384
REMARK   3    18  1.7948 -  1.7610    1.00     2704   141  0.2790 0.3587
REMARK   3    19  1.7610 -  1.7296    0.57     1536    75  0.3332 0.3743
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : NULL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.220
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 24.680
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.014           4855
REMARK   3   ANGLE     :  1.437           6608
REMARK   3   CHIRALITY :  0.070            678
REMARK   3   PLANARITY :  0.010            866
REMARK   3   DIHEDRAL  : 18.629           2847
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 6QI2 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 17-JAN-19.
REMARK 100 THE DEPOSITION ID IS D_1200013246.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 01-MAY-17
REMARK 200  TEMPERATURE           (KELVIN) : 293
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : PETRA III, DESY
REMARK 200  BEAMLINE                       : P11
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0089
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 53237
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.730
REMARK 200  RESOLUTION RANGE LOW       (A) : 81.480
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 97.7
REMARK 200  DATA REDUNDANCY                : 31.60
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 4.6000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.73
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.76
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 36.75
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 1.94
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 18-20 % (W/V)) PEG3350, 200 MM CACL2,
REMARK 280  AND 100 MM TRIS-HCL PH 8.5, BATCH MODE, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       39.30500
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 2520 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 19780 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -13.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLY A    -1
REMARK 465     HIS A     0
REMARK 465     MET A     1
REMARK 465     PRO A     2
REMARK 465     ASP A     3
REMARK 465     ALA A   301
REMARK 465     PRO A   302
REMARK 465     GLY A   303
REMARK 465     SER A   304
REMARK 465     GLY B    -1
REMARK 465     HIS B     0
REMARK 465     MET B     1
REMARK 465     PRO B     2
REMARK 465     ALA B   300
REMARK 465     ALA B   301
REMARK 465     PRO B   302
REMARK 465     GLY B   303
REMARK 465     SER B   304
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   NH2  ARG A    49     O    ALA A   193              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    PRO A  41       64.19   -105.67
REMARK 500    ASP A 110     -125.74     60.13
REMARK 500    SER A 123       57.59   -140.32
REMARK 500    ILE A 153       55.60   -113.22
REMARK 500    TYR A 224      -94.20   -119.66
REMARK 500    ALA A 258      174.15    178.03
REMARK 500    PRO B  41       69.48   -104.49
REMARK 500    GLN B  42     -168.28   -124.71
REMARK 500    ASP B 110     -126.30     59.04
REMARK 500    SER B 123       57.59   -147.44
REMARK 500    ASP B 173       72.11   -160.44
REMARK 500    TYR B 224      -93.97   -123.24
REMARK 500    ALA B 254      139.03    119.75
REMARK 500    GLN B 255        7.27    157.29
REMARK 500    SER B 256     -130.64    -62.53
REMARK 500    ALA B 257      -94.80    -55.42
REMARK 500    GLU B 277       88.18    -65.90
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH B 478        DISTANCE =  6.01 ANGSTROMS
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOA A 401
DBREF  6QI2 A    1   302  UNP    Q6NAM1   DEHA_RHOPA       1    302
DBREF  6QI2 B    1   302  UNP    Q6NAM1   DEHA_RHOPA       1    302
SEQADV 6QI2 GLY A   -1  UNP  Q6NAM1              EXPRESSION TAG
SEQADV 6QI2 HIS A    0  UNP  Q6NAM1              EXPRESSION TAG
SEQADV 6QI2 GLY A  303  UNP  Q6NAM1              EXPRESSION TAG
SEQADV 6QI2 SER A  304  UNP  Q6NAM1              EXPRESSION TAG
SEQADV 6QI2 GLY B   -1  UNP  Q6NAM1              EXPRESSION TAG
SEQADV 6QI2 HIS B    0  UNP  Q6NAM1              EXPRESSION TAG
SEQADV 6QI2 GLY B  303  UNP  Q6NAM1              EXPRESSION TAG
SEQADV 6QI2 SER B  304  UNP  Q6NAM1              EXPRESSION TAG
SEQRES   1 A  306  GLY HIS MET PRO ASP LEU ALA ASP LEU PHE PRO GLY PHE
SEQRES   2 A  306  GLY SER GLU TRP ILE ASN THR SER SER GLY ARG ILE PHE
SEQRES   3 A  306  ALA ARG VAL GLY GLY ASP GLY PRO PRO LEU LEU LEU LEU
SEQRES   4 A  306  HIS GLY PHE PRO GLN THR HIS VAL MET TRP HIS ARG VAL
SEQRES   5 A  306  ALA PRO LYS LEU ALA GLU ARG PHE LYS VAL ILE VAL ALA
SEQRES   6 A  306  ASP LEU PRO GLY TYR GLY TRP SER ASP MET PRO GLU SER
SEQRES   7 A  306  ASP GLU GLN HIS THR PRO TYR THR LYS ARG ALA MET ALA
SEQRES   8 A  306  LYS GLN LEU ILE GLU ALA MET GLU GLN LEU GLY HIS VAL
SEQRES   9 A  306  HIS PHE ALA LEU ALA GLY HIS ASP ARG GLY ALA ARG VAL
SEQRES  10 A  306  SER TYR ARG LEU ALA LEU ASP SER PRO GLY ARG LEU SER
SEQRES  11 A  306  LYS LEU ALA VAL LEU ASP ILE LEU PRO THR TYR GLU TYR
SEQRES  12 A  306  TRP GLN ARG MET ASN ARG ALA TYR ALA LEU LYS ILE TYR
SEQRES  13 A  306  HIS TRP SER PHE LEU ALA GLN PRO ALA PRO LEU PRO GLU
SEQRES  14 A  306  ASN LEU LEU GLY GLY ASP PRO ASP PHE TYR VAL LYS ALA
SEQRES  15 A  306  LYS LEU ALA SER TRP THR ARG ALA GLY ASP LEU SER ALA
SEQRES  16 A  306  PHE ASP PRO ARG ALA VAL GLU HIS TYR ARG ILE ALA PHE
SEQRES  17 A  306  ALA ASP PRO MET ARG ARG HIS VAL MET CYS GLU ASP TYR
SEQRES  18 A  306  ARG ALA GLY ALA TYR ALA ASP PHE GLU HIS ASP LYS ILE
SEQRES  19 A  306  ASP VAL GLU ALA GLY ASN LYS ILE PRO VAL PRO MET LEU
SEQRES  20 A  306  ALA LEU TRP GLY ALA SER GLY ILE ALA GLN SER ALA ALA
SEQRES  21 A  306  THR PRO LEU ASP VAL TRP ARG LYS TRP ALA SER ASP VAL
SEQRES  22 A  306  GLN GLY ALA PRO ILE GLU SER GLY HIS PHE LEU PRO GLU
SEQRES  23 A  306  GLU ALA PRO ASP GLN THR ALA GLU ALA LEU VAL ARG PHE
SEQRES  24 A  306  PHE SER ALA ALA PRO GLY SER
SEQRES   1 B  306  GLY HIS MET PRO ASP LEU ALA ASP LEU PHE PRO GLY PHE
SEQRES   2 B  306  GLY SER GLU TRP ILE ASN THR SER SER GLY ARG ILE PHE
SEQRES   3 B  306  ALA ARG VAL GLY GLY ASP GLY PRO PRO LEU LEU LEU LEU
SEQRES   4 B  306  HIS GLY PHE PRO GLN THR HIS VAL MET TRP HIS ARG VAL
SEQRES   5 B  306  ALA PRO LYS LEU ALA GLU ARG PHE LYS VAL ILE VAL ALA
SEQRES   6 B  306  ASP LEU PRO GLY TYR GLY TRP SER ASP MET PRO GLU SER
SEQRES   7 B  306  ASP GLU GLN HIS THR PRO TYR THR LYS ARG ALA MET ALA
SEQRES   8 B  306  LYS GLN LEU ILE GLU ALA MET GLU GLN LEU GLY HIS VAL
SEQRES   9 B  306  HIS PHE ALA LEU ALA GLY HIS ASP ARG GLY ALA ARG VAL
SEQRES  10 B  306  SER TYR ARG LEU ALA LEU ASP SER PRO GLY ARG LEU SER
SEQRES  11 B  306  LYS LEU ALA VAL LEU ASP ILE LEU PRO THR TYR GLU TYR
SEQRES  12 B  306  TRP GLN ARG MET ASN ARG ALA TYR ALA LEU LYS ILE TYR
SEQRES  13 B  306  HIS TRP SER PHE LEU ALA GLN PRO ALA PRO LEU PRO GLU
SEQRES  14 B  306  ASN LEU LEU GLY GLY ASP PRO ASP PHE TYR VAL LYS ALA
SEQRES  15 B  306  LYS LEU ALA SER TRP THR ARG ALA GLY ASP LEU SER ALA
SEQRES  16 B  306  PHE ASP PRO ARG ALA VAL GLU HIS TYR ARG ILE ALA PHE
SEQRES  17 B  306  ALA ASP PRO MET ARG ARG HIS VAL MET CYS GLU ASP TYR
SEQRES  18 B  306  ARG ALA GLY ALA TYR ALA ASP PHE GLU HIS ASP LYS ILE
SEQRES  19 B  306  ASP VAL GLU ALA GLY ASN LYS ILE PRO VAL PRO MET LEU
SEQRES  20 B  306  ALA LEU TRP GLY ALA SER GLY ILE ALA GLN SER ALA ALA
SEQRES  21 B  306  THR PRO LEU ASP VAL TRP ARG LYS TRP ALA SER ASP VAL
SEQRES  22 B  306  GLN GLY ALA PRO ILE GLU SER GLY HIS PHE LEU PRO GLU
SEQRES  23 B  306  GLU ALA PRO ASP GLN THR ALA GLU ALA LEU VAL ARG PHE
SEQRES  24 B  306  PHE SER ALA ALA PRO GLY SER
HET    GOA  A 401       5
HETNAM     GOA GLYCOLIC ACID
HETSYN     GOA HYDROXYACETIC ACID; HYDROXYETHANOIC ACID
FORMUL   3  GOA    C2 H4 O3
FORMUL   4  HOH   *173(H2 O)
HELIX    1 AA1 THR A   43  HIS A   48  5                                   6
HELIX    2 AA2 VAL A   50  GLU A   56  1                                   7
HELIX    3 AA3 HIS A   80  TYR A   83  5                                   4
HELIX    4 AA4 THR A   84  LEU A   99  1                                  16
HELIX    5 AA5 ASP A  110  SER A  123  1                                  14
HELIX    6 AA6 PRO A  137  ARG A  144  1                                   8
HELIX    7 AA7 ASN A  146  ILE A  153  1                                   8
HELIX    8 AA8 ILE A  153  LEU A  159  1                                   7
HELIX    9 AA9 PRO A  164  GLY A  172  1                                   9
HELIX   10 AB1 ASP A  173  TRP A  185  1                                  13
HELIX   11 AB2 ASP A  195  ASP A  208  1                                  14
HELIX   12 AB3 ASP A  208  TYR A  224  1                                  17
HELIX   13 AB4 TYR A  224  GLY A  237  1                                  14
HELIX   14 AB5 GLY A  252  ALA A  258  1                                   7
HELIX   15 AB6 THR A  259  ALA A  268  1                                  10
HELIX   16 AB7 PHE A  281  ALA A  286  1                                   6
HELIX   17 AB8 ALA A  286  ALA A  300  1                                  15
HELIX   18 AB9 THR B   43  HIS B   48  5                                   6
HELIX   19 AC1 VAL B   50  ALA B   55  1                                   6
HELIX   20 AC2 HIS B   80  TYR B   83  5                                   4
HELIX   21 AC3 THR B   84  LEU B   99  1                                  16
HELIX   22 AC4 ASP B  110  SER B  123  1                                  14
HELIX   23 AC5 PRO B  137  ARG B  144  1                                   8
HELIX   24 AC6 ASN B  146  ILE B  153  1                                   8
HELIX   25 AC7 ILE B  153  LEU B  159  1                                   7
HELIX   26 AC8 PRO B  164  ASP B  173  1                                  10
HELIX   27 AC9 ASP B  173  TRP B  185  1                                  13
HELIX   28 AD1 ASP B  195  ASP B  208  1                                  14
HELIX   29 AD2 ASP B  208  TYR B  224  1                                  17
HELIX   30 AD3 TYR B  224  GLY B  237  1                                  14
HELIX   31 AD4 THR B  259  ALA B  268  1                                  10
HELIX   32 AD5 PHE B  281  ALA B  286  1                                   6
HELIX   33 AD6 ALA B  286  SER B  299  1                                  14
SHEET    1 AA1 8 GLY A  12  ILE A  16  0
SHEET    2 AA1 8 ILE A  23  GLY A  29 -1  O  ILE A  23   N  ILE A  16
SHEET    3 AA1 8 LYS A  59  ALA A  63 -1  O  VAL A  60   N  GLY A  28
SHEET    4 AA1 8 PRO A  33  LEU A  37  1  N  LEU A  36   O  ILE A  61
SHEET    5 AA1 8 PHE A 104  HIS A 109  1  O  ALA A 105   N  LEU A  35
SHEET    6 AA1 8 LEU A 127  LEU A 133  1  O  ALA A 131   N  LEU A 106
SHEET    7 AA1 8 MET A 244  GLY A 249  1  O  LEU A 247   N  VAL A 132
SHEET    8 AA1 8 VAL A 271  ILE A 276  1  O  GLN A 272   N  ALA A 246
SHEET    1 AA2 8 GLY B  12  ILE B  16  0
SHEET    2 AA2 8 ILE B  23  GLY B  29 -1  O  VAL B  27   N  GLY B  12
SHEET    3 AA2 8 LYS B  59  ALA B  63 -1  O  VAL B  62   N  ARG B  26
SHEET    4 AA2 8 PRO B  33  LEU B  37  1  N  LEU B  34   O  ILE B  61
SHEET    5 AA2 8 PHE B 104  HIS B 109  1  O  ALA B 105   N  LEU B  35
SHEET    6 AA2 8 LEU B 127  LEU B 133  1  O  ALA B 131   N  LEU B 106
SHEET    7 AA2 8 MET B 244  GLY B 249  1  O  LEU B 245   N  VAL B 132
SHEET    8 AA2 8 VAL B 271  ILE B 276  1  O  GLN B 272   N  ALA B 246
CISPEP   1 PHE A   40    PRO A   41          0         2.14
CISPEP   2 ALA A  163    PRO A  164          0         4.45
CISPEP   3 PHE B   40    PRO B   41          0        -1.33
CISPEP   4 ALA B  163    PRO B  164          0         2.77
SITE     1 AC1  9 ASP A 110  ARG A 111  ARG A 114  ILE A 135
SITE     2 AC1  9 HIS A 155  TRP A 156  TYR A 219  ILE A 253
SITE     3 AC1  9 HOH A 501
CRYST1   41.380   78.610   83.530  90.00 102.73  90.00 P 1 21 1      4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.024166  0.000000  0.005459        0.00000
SCALE2      0.000000  0.012721  0.000000        0.00000
SCALE3      0.000000  0.000000  0.012273        0.00000
TER    2349      ALA A 300
TER    4701      SER B 299
MASTER      289    0    1   33   16    0    3    6 4877    2    5   48
END