longtext: 6rs4-pdb

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HEADER    HYDROLASE                               21-MAY-19   6RS4
TITLE     STRUCTURE OF TABERSONINE SYNTHASE - AN ALPHA-BETA HYDROLASE FROM
TITLE    2 CATHARANTHUS ROSEUS
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: TABERSONINE SYNTHASE;
COMPND   3 CHAIN: A, B;
COMPND   4 SYNONYM: HYDROLASE 2,CRHL2;
COMPND   5 EC: 4.-.-.-;
COMPND   6 ENGINEERED: YES;
COMPND   7 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: CATHARANTHUS ROSEUS;
SOURCE   3 ORGANISM_COMMON: MADAGASCAR PERIWINKLE;
SOURCE   4 ORGANISM_TAXID: 4058;
SOURCE   5 GENE: TS, HL2;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   8 EXPRESSION_SYSTEM_VARIANT: SOLUBL21;
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: POPINF
KEYWDS    ALKALOID, TABERSONINE, NATURAL PRODUCT, BIOSYNTHESIS, ALPHA/BETA
KEYWDS   2 HYDROLASE FOLD, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    L.CAPUTI,J.FRANKE,K.BUSSEY,S.C.FARROW,I.J.CURCINO VIEIRA,
AUTHOR   2 C.E.M.STEVENSON,D.M.LAWSON,S.E.O'CONNOR
REVDAT   1   04-MAR-20 6RS4    0
JRNL        AUTH   L.CAPUTI,J.FRANKE,K.BUSSEY,S.C.FARROW,I.J.C.VIEIRA,
JRNL        AUTH 2 C.E.M.STEVENSON,D.M.LAWSON,S.E.O'CONNOR
JRNL        TITL   STRUCTURAL BASIS OF CYCLOADDITION IN BIOSYNTHESIS OF IBOGA
JRNL        TITL 2 AND ASPIDOSPERMA ALKALOIDS.
JRNL        REF    NAT.CHEM.BIOL.                             2020
JRNL        REFN                   ESSN 1552-4469
JRNL        PMID   32066966
JRNL        DOI    10.1038/S41589-019-0460-X
REMARK   2
REMARK   2 RESOLUTION.    1.30 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0238
REMARK   3   AUTHORS     : MURSHUDOV,SKUBAK,LEBEDEV,PANNU,STEINER,
REMARK   3               : NICHOLLS,WINN,LONG,VAGIN
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.30
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 59.95
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.9
REMARK   3   NUMBER OF REFLECTIONS             : 168522
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.134
REMARK   3   R VALUE            (WORKING SET) : 0.133
REMARK   3   FREE R VALUE                     : 0.162
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 8863
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.30
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.33
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 12394
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 99.82
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2810
REMARK   3   BIN FREE R VALUE SET COUNT          : 689
REMARK   3   BIN FREE R VALUE                    : 0.3110
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4968
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 44
REMARK   3   SOLVENT ATOMS            : 699
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 10.60
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 16.17
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.63000
REMARK   3    B22 (A**2) : 1.80000
REMARK   3    B33 (A**2) : -1.59000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : -0.73000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.040
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.040
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.033
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 1.907
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.981
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.975
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  5463 ; 0.010 ; 0.013
REMARK   3   BOND LENGTHS OTHERS               (A):  4931 ; 0.001 ; 0.017
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  7475 ; 1.636 ; 1.639
REMARK   3   BOND ANGLES OTHERS          (DEGREES): 11481 ; 1.506 ; 1.570
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   710 ; 6.753 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   266 ;33.774 ;23.271
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   862 ;12.624 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    20 ;17.114 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   702 ; 0.098 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  6251 ; 0.009 ; 0.020
REMARK   3   GENERAL PLANES OTHERS             (A):  1169 ; 0.001 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2): 10394 ; 7.269 ; 3.000
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : NULL
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS U VALUES : REFINED INDIVIDUALLY
REMARK   4
REMARK   4 6RS4 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 21-MAY-19.
REMARK 100 THE DEPOSITION ID IS D_1292102507.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 17-JUL-16
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : DIAMOND
REMARK 200  BEAMLINE                       : I03
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9795
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS3 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS 20160517
REMARK 200  DATA SCALING SOFTWARE          : AIMLESS 0.7.2
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 177427
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.300
REMARK 200  RESOLUTION RANGE LOW       (A) : 59.950
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 100.0
REMARK 200  DATA REDUNDANCY                : 6.700
REMARK 200  R MERGE                    (I) : 0.11100
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 10.8000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.30
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.32
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.9
REMARK 200  DATA REDUNDANCY IN SHELL       : 6.50
REMARK 200  R MERGE FOR SHELL          (I) : 1.68000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER 2.6.1
REMARK 200 STARTING MODEL: 2O7R
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 51.66
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.54
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL, PH 5, VAPOR DIFFUSION, SITTING
REMARK 280  DROP, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       54.48100
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLY A     0
REMARK 465     PRO A     1
REMARK 465     GLY A     2
REMARK 465     SER A     3
REMARK 465     SER A     4
REMARK 465     ASP A     5
REMARK 465     GLY B     0
REMARK 465     PRO B     1
REMARK 465     GLY B     2
REMARK 465     SER B     3
REMARK 465     SER B     4
REMARK 465     ASP B     5
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     GLU A   6    CG   CD   OE1  OE2
REMARK 470     LYS A  16    NZ
REMARK 470     LYS A  19    CG   CD   CE   NZ
REMARK 470     ARG A  22    CG   CD   NE   CZ   NH1  NH2
REMARK 470     GLU A  24    CD   OE1  OE2
REMARK 470     LEU A  26    CG   CD1  CD2
REMARK 470     HIS A  27    CG   ND1  CD2  CE1  NE2
REMARK 470     LYS A  63    NZ
REMARK 470     LYS A 104    CE   NZ
REMARK 470     LYS A 194    CD   CE   NZ
REMARK 470     LYS A 226    CE   NZ
REMARK 470     GLU A 304    CG   CD   OE1  OE2
REMARK 470     LYS A 309    CG   CD   CE   NZ
REMARK 470     ARG A 313    CZ   NH1  NH2
REMARK 470     LEU B  11    CD1  CD2
REMARK 470     LYS B  16    NZ
REMARK 470     LYS B  19    CG   CD   CE   NZ
REMARK 470     ARG B  22    CG   CD   NE   CZ   NH1  NH2
REMARK 470     GLU B  24    CG   CD   OE1  OE2
REMARK 470     HIS B  27    CG   ND1  CD2  CE1  NE2
REMARK 470     LYS B 104    CD   CE   NZ
REMARK 470     LYS B 146    CE   NZ
REMARK 470     LYS B 194    CD   CE   NZ
REMARK 470     LYS B 212    CD   CE   NZ
REMARK 470     ARG B 222    CZ   NH1  NH2
REMARK 470     LYS B 309    CD   CE   NZ
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    TYR A 254   CB  -  CG  -  CD1 ANGL. DEV. =   3.6 DEGREES
REMARK 500    TYR B 254   CB  -  CG  -  CD1 ANGL. DEV. =   4.4 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ALA A  80       48.56     82.13
REMARK 500    GLU A  85     -173.18     67.85
REMARK 500    SER A 170     -112.67     44.96
REMARK 500    GLN A 213     -157.56   -100.80
REMARK 500    ASN A 219      -89.38   -125.97
REMARK 500    ALA B  80       51.40     83.74
REMARK 500    PHE B  82       -1.43     69.29
REMARK 500    GLU B  85     -174.69     69.03
REMARK 500    SER B 170     -114.20     46.68
REMARK 500    ASN B 219      -78.48   -112.32
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 848        DISTANCE =  6.11 ANGSTROMS
REMARK 525    HOH B 851        DISTANCE =  6.10 ANGSTROMS
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 402
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 403
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 404
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 405
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 406
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO B 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO B 402
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO B 403
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO B 404
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO B 405
DBREF1 6RS4 A    2   320  UNP                  TS_CATRO
DBREF2 6RS4 A     A0A2P1GIW3                          2         320
DBREF1 6RS4 B    2   320  UNP                  TS_CATRO
DBREF2 6RS4 B     A0A2P1GIW3                          2         320
SEQADV 6RS4 GLY A    0  UNP  A0A2P1GIW           EXPRESSION TAG
SEQADV 6RS4 PRO A    1  UNP  A0A2P1GIW           EXPRESSION TAG
SEQADV 6RS4 GLY B    0  UNP  A0A2P1GIW           EXPRESSION TAG
SEQADV 6RS4 PRO B    1  UNP  A0A2P1GIW           EXPRESSION TAG
SEQRES   1 A  321  GLY PRO GLY SER SER ASP GLU THR ILE PHE ASP LEU PRO
SEQRES   2 A  321  PRO TYR ILE LYS VAL PHE LYS ASP GLY ARG VAL GLU ARG
SEQRES   3 A  321  LEU HIS SER SER PRO TYR VAL PRO PRO SER LEU ASN ASP
SEQRES   4 A  321  PRO GLU THR GLY GLY VAL SER TRP LYS ASP VAL PRO ILE
SEQRES   5 A  321  SER SER VAL VAL SER ALA ARG ILE TYR LEU PRO LYS ILE
SEQRES   6 A  321  ASN ASN HIS ASP GLU LYS LEU PRO ILE ILE VAL TYR PHE
SEQRES   7 A  321  HIS GLY ALA GLY PHE CYS LEU GLU SER ALA PHE LYS SER
SEQRES   8 A  321  PHE PHE HIS THR TYR VAL LYS HIS PHE VAL ALA GLU ALA
SEQRES   9 A  321  LYS ALA ILE ALA VAL SER VAL GLU PHE ARG LEU ALA PRO
SEQRES  10 A  321  GLU ASN HIS LEU PRO ALA ALA TYR GLU ASP CYS TRP GLU
SEQRES  11 A  321  ALA LEU GLN TRP VAL ALA SER HIS VAL GLY LEU ASP ILE
SEQRES  12 A  321  SER SER LEU LYS THR CYS ILE ASP LYS ASP PRO TRP ILE
SEQRES  13 A  321  ILE ASN TYR ALA ASP PHE ASP ARG LEU TYR LEU TRP GLY
SEQRES  14 A  321  ASP SER THR GLY ALA ASN ILE VAL HIS ASN THR LEU ILE
SEQRES  15 A  321  ARG SER GLY LYS GLU LYS LEU ASN GLY GLY LYS VAL LYS
SEQRES  16 A  321  ILE LEU GLY ALA ILE LEU TYR TYR PRO TYR PHE LEU ILE
SEQRES  17 A  321  ARG THR SER SER LYS GLN SER ASP TYR MET GLU ASN GLU
SEQRES  18 A  321  TYR ARG SER TYR TRP LYS LEU ALA TYR PRO ASP ALA PRO
SEQRES  19 A  321  GLY GLY ASN ASP ASN PRO MET ILE ASN PRO THR ALA GLU
SEQRES  20 A  321  ASN ALA PRO ASP LEU ALA GLY TYR GLY CYS SER ARG LEU
SEQRES  21 A  321  LEU ILE SER MET VAL ALA ASP GLU ALA ARG ASP ILE THR
SEQRES  22 A  321  LEU LEU TYR ILE ASP ALA LEU GLU LYS SER GLY TRP LYS
SEQRES  23 A  321  GLY GLU LEU ASP VAL ALA ASP PHE ASP LYS GLN TYR PHE
SEQRES  24 A  321  GLU LEU PHE GLU MET GLU THR GLU VAL ALA LYS ASN MET
SEQRES  25 A  321  LEU ARG ARG LEU ALA SER PHE ILE LYS
SEQRES   1 B  321  GLY PRO GLY SER SER ASP GLU THR ILE PHE ASP LEU PRO
SEQRES   2 B  321  PRO TYR ILE LYS VAL PHE LYS ASP GLY ARG VAL GLU ARG
SEQRES   3 B  321  LEU HIS SER SER PRO TYR VAL PRO PRO SER LEU ASN ASP
SEQRES   4 B  321  PRO GLU THR GLY GLY VAL SER TRP LYS ASP VAL PRO ILE
SEQRES   5 B  321  SER SER VAL VAL SER ALA ARG ILE TYR LEU PRO LYS ILE
SEQRES   6 B  321  ASN ASN HIS ASP GLU LYS LEU PRO ILE ILE VAL TYR PHE
SEQRES   7 B  321  HIS GLY ALA GLY PHE CYS LEU GLU SER ALA PHE LYS SER
SEQRES   8 B  321  PHE PHE HIS THR TYR VAL LYS HIS PHE VAL ALA GLU ALA
SEQRES   9 B  321  LYS ALA ILE ALA VAL SER VAL GLU PHE ARG LEU ALA PRO
SEQRES  10 B  321  GLU ASN HIS LEU PRO ALA ALA TYR GLU ASP CYS TRP GLU
SEQRES  11 B  321  ALA LEU GLN TRP VAL ALA SER HIS VAL GLY LEU ASP ILE
SEQRES  12 B  321  SER SER LEU LYS THR CYS ILE ASP LYS ASP PRO TRP ILE
SEQRES  13 B  321  ILE ASN TYR ALA ASP PHE ASP ARG LEU TYR LEU TRP GLY
SEQRES  14 B  321  ASP SER THR GLY ALA ASN ILE VAL HIS ASN THR LEU ILE
SEQRES  15 B  321  ARG SER GLY LYS GLU LYS LEU ASN GLY GLY LYS VAL LYS
SEQRES  16 B  321  ILE LEU GLY ALA ILE LEU TYR TYR PRO TYR PHE LEU ILE
SEQRES  17 B  321  ARG THR SER SER LYS GLN SER ASP TYR MET GLU ASN GLU
SEQRES  18 B  321  TYR ARG SER TYR TRP LYS LEU ALA TYR PRO ASP ALA PRO
SEQRES  19 B  321  GLY GLY ASN ASP ASN PRO MET ILE ASN PRO THR ALA GLU
SEQRES  20 B  321  ASN ALA PRO ASP LEU ALA GLY TYR GLY CYS SER ARG LEU
SEQRES  21 B  321  LEU ILE SER MET VAL ALA ASP GLU ALA ARG ASP ILE THR
SEQRES  22 B  321  LEU LEU TYR ILE ASP ALA LEU GLU LYS SER GLY TRP LYS
SEQRES  23 B  321  GLY GLU LEU ASP VAL ALA ASP PHE ASP LYS GLN TYR PHE
SEQRES  24 B  321  GLU LEU PHE GLU MET GLU THR GLU VAL ALA LYS ASN MET
SEQRES  25 B  321  LEU ARG ARG LEU ALA SER PHE ILE LYS
HET    EDO  A 401       4
HET    EDO  A 402       4
HET    EDO  A 403       4
HET    EDO  A 404       4
HET    EDO  A 405       4
HET    EDO  A 406       8
HET    EDO  B 401       8
HET    EDO  B 402       4
HET    EDO  B 403       4
HET    EDO  B 404       4
HET    EDO  B 405       8
HETNAM     EDO 1,2-ETHANEDIOL
HETSYN     EDO ETHYLENE GLYCOL
FORMUL   3  EDO    11(C2 H6 O2)
FORMUL  14  HOH   *699(H2 O)
HELIX    1 AA1 LYS A   89  LYS A  104  1                                  16
HELIX    2 AA2 PRO A  121  SER A  136  1                                  16
HELIX    3 AA3 ASP A  141  THR A  147  5                                   7
HELIX    4 AA4 ASP A  152  TYR A  158  1                                   7
HELIX    5 AA5 SER A  170  GLY A  184  1                                  15
HELIX    6 AA6 LYS A  187  LYS A  192  5                                   6
HELIX    7 AA7 SER A  214  ASN A  219  1                                   6
HELIX    8 AA8 ASN A  219  TYR A  229  1                                  11
HELIX    9 AA9 GLY A  234  ASN A  238  5                                   5
HELIX   10 AB1 ASP A  250  TYR A  254  5                                   5
HELIX   11 AB2 GLU A  267  GLY A  283  1                                  17
HELIX   12 AB3 GLN A  296  GLU A  302  5                                   7
HELIX   13 AB4 THR A  305  SER A  317  1                                  13
HELIX   14 AB5 LYS B   89  LYS B  104  1                                  16
HELIX   15 AB6 PRO B  121  SER B  136  1                                  16
HELIX   16 AB7 ASP B  141  THR B  147  5                                   7
HELIX   17 AB8 ASP B  152  TYR B  158  1                                   7
HELIX   18 AB9 SER B  170  GLU B  186  1                                  17
HELIX   19 AC1 LYS B  187  LYS B  192  5                                   6
HELIX   20 AC2 SER B  214  ASN B  219  1                                   6
HELIX   21 AC3 ASN B  219  TYR B  229  1                                  11
HELIX   22 AC4 GLY B  234  ASN B  238  5                                   5
HELIX   23 AC5 ASP B  250  TYR B  254  5                                   5
HELIX   24 AC6 ALA B  268  SER B  282  1                                  15
HELIX   25 AC7 GLN B  296  GLU B  302  5                                   7
HELIX   26 AC8 THR B  305  SER B  317  1                                  13
SHEET    1 AA1 3 THR A   7  LEU A  11  0
SHEET    2 AA1 3 ILE A  15  PHE A  18 -1  O  ILE A  15   N  LEU A  11
SHEET    3 AA1 3 VAL A  23  ARG A  25 -1  O  GLU A  24   N  LYS A  16
SHEET    1 AA2 8 SER A  45  PRO A  50  0
SHEET    2 AA2 8 SER A  56  LEU A  61 -1  O  ILE A  59   N  LYS A  47
SHEET    3 AA2 8 ILE A 106  GLU A 111 -1  O  GLU A 111   N  SER A  56
SHEET    4 AA2 8 LEU A  71  PHE A  77  1  N  ILE A  74   O  ILE A 106
SHEET    5 AA2 8 ALA A 159  ASP A 169  1  O  TYR A 165   N  ILE A  73
SHEET    6 AA2 8 GLY A 197  TYR A 201  1  O  TYR A 201   N  GLY A 168
SHEET    7 AA2 8 ARG A 258  MET A 263  1  O  LEU A 260   N  LEU A 200
SHEET    8 AA2 8 GLU A 287  ASP A 292  1  O  ASP A 289   N  LEU A 259
SHEET    1 AA3 3 THR B   7  LEU B  11  0
SHEET    2 AA3 3 ILE B  15  PHE B  18 -1  O  VAL B  17   N  PHE B   9
SHEET    3 AA3 3 VAL B  23  GLU B  24 -1  O  GLU B  24   N  LYS B  16
SHEET    1 AA4 8 SER B  45  PRO B  50  0
SHEET    2 AA4 8 SER B  56  LEU B  61 -1  O  ILE B  59   N  LYS B  47
SHEET    3 AA4 8 ILE B 106  GLU B 111 -1  O  GLU B 111   N  SER B  56
SHEET    4 AA4 8 LEU B  71  PHE B  77  1  N  ILE B  74   O  ILE B 106
SHEET    5 AA4 8 ALA B 159  ASP B 169  1  O  TYR B 165   N  ILE B  73
SHEET    6 AA4 8 GLY B 197  TYR B 201  1  O  TYR B 201   N  GLY B 168
SHEET    7 AA4 8 ARG B 258  MET B 263  1  O  LEU B 260   N  LEU B 200
SHEET    8 AA4 8 GLU B 287  ASP B 292  1  O  ASP B 289   N  LEU B 259
CISPEP   1 PRO A   12    PRO A   13          0        -2.79
CISPEP   2 ALA A  115    PRO A  116          0        -1.24
CISPEP   3 LEU A  120    PRO A  121          0        10.53
CISPEP   4 PRO B   12    PRO B   13          0        -3.40
CISPEP   5 ALA B  115    PRO B  116          0        -4.05
CISPEP   6 LEU B  120    PRO B  121          0         9.39
SITE     1 AC1  7 GLY A  81  SER A 170  THR A 171  TYR A 204
SITE     2 AC1  7 HOH A 505  HOH A 697  HOH A 733
SITE     1 AC2  5 SER A  28  GLU A  85  LYS A  89  EDO A 403
SITE     2 AC2  5 HOH A 587
SITE     1 AC3  7 TYR A  31  SER A  86  LYS A  89  EDO A 402
SITE     2 AC3  7 HOH A 720  HOH A 729  TYR B  31
SITE     1 AC4  8 HIS A 137  VAL A 138  GLY A 139  LEU A 140
SITE     2 AC4  8 ILE A 156  PHE A 161  HOH A 510  HOH A 702
SITE     1 AC5  7 LEU A 206  ILE A 207  ARG A 208  ARG A 222
SITE     2 AC5  7 ASN A 236  ASP A 237  HOH A 633
SITE     1 AC6 10 GLU A 129  GLN A 132  TRP A 133  SER A 136
SITE     2 AC6 10 LYS A 151  HOH A 507  HOH A 521  HOH A 522
SITE     3 AC6 10 HOH A 537  HOH A 547
SITE     1 AC7 11 GLY B  81  SER B 170  THR B 171  TYR B 202
SITE     2 AC7 11 TYR B 204  TYR B 221  TYR B 224  HOH B 509
SITE     3 AC7 11 HOH B 556  HOH B 565  HOH B 632
SITE     1 AC8  5 PRO A  30  TYR A  31  HOH A 729  LYS B  89
SITE     2 AC8  5 HOH B 569
SITE     1 AC9  7 HIS B 137  VAL B 138  GLY B 139  LEU B 140
SITE     2 AC9  7 PHE B 161  HOH B 547  HOH B 715
SITE     1 AD1  5 ARG B  25  SER B  86  LYS B  89  HOH B 520
SITE     2 AD1  5 HOH B 607
SITE     1 AD2 10 GLU B 129  GLN B 132  TRP B 133  SER B 136
SITE     2 AD2 10 LYS B 151  HOH B 506  HOH B 511  HOH B 527
SITE     3 AD2 10 HOH B 528  HOH B 553
CRYST1   56.485  108.962   63.235  90.00 108.54  90.00 P 1 21 1      4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.017704  0.000000  0.005937        0.00000
SCALE2      0.000000  0.009178  0.000000        0.00000
SCALE3      0.000000  0.000000  0.016680        0.00000
TER    2611      LYS A 320
TER    5236      LYS B 320
MASTER      383    0   11   26   22    0   25    6 5711    2   56   50
END