longtext: 6syr-pdb

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HEADER    HYDROLASE                               01-OCT-19   6SYR
TITLE     THE WILD TYPE GLUCURONOYL ESTERASE OTCE15A FROM OPITUTUS TERRAE IN
TITLE    2 COMPLEX WITH D-GLUCURONATE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: GLUCURONOYL ESTERASE OTCE15A;
COMPND   3 CHAIN: A;
COMPND   4 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: OPITUTUS TERRAE PB90-1;
SOURCE   3 ORGANISM_TAXID: 452637;
SOURCE   4 GENE: OTER_0116;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21(DE3)
KEYWDS    ESTERASE, COMPLEX, BIOMASS, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    S.MAZURKEWICH,J.C.NAVARRO POULSEN,J.LARSBRINK,L.LO LEGGIO
REVDAT   1   27-NOV-19 6SYR    0
JRNL        AUTH   S.MAZURKEWICH,J.C.NAVARRO POULSEN,J.LARSBRINK,L.LO LEGGIO
JRNL        TITL   THE WILD TYPE GLUCURONOYL ESTERASE OTCE15A FROM OPITUTUS
JRNL        TITL 2 TERRAE IN COMPLEX WITH D-GLUCURONATE
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    1.49 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.15.2_3472
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.49
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 32.69
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.960
REMARK   3   COMPLETENESS FOR RANGE        (%) : 91.2
REMARK   3   NUMBER OF REFLECTIONS             : 47109
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.186
REMARK   3   R VALUE            (WORKING SET) : 0.183
REMARK   3   FREE R VALUE                     : 0.223
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 7.470
REMARK   3   FREE R VALUE TEST SET COUNT      : 3520
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 32.6900 -  4.3500    0.98     1882   151  0.1422 0.1864
REMARK   3     2  4.3500 -  3.4600    0.97     1866   150  0.1262 0.1604
REMARK   3     3  3.4600 -  3.0200    0.98     1868   148  0.1470 0.1907
REMARK   3     4  3.0200 -  2.7400    0.98     1865   152  0.1541 0.1910
REMARK   3     5  2.7400 -  2.5500    0.96     1826   145  0.1568 0.1775
REMARK   3     6  2.5500 -  2.4000    0.71     1373   116  0.1559 0.1894
REMARK   3     7  2.4000 -  2.2800    0.82     1556   128  0.1577 0.2038
REMARK   3     8  2.2800 -  2.1800    0.87     1676   135  0.1688 0.2280
REMARK   3     9  2.1800 -  2.0900    0.89     1693   129  0.1670 0.2476
REMARK   3    10  2.0900 -  2.0200    0.90     1716   138  0.1881 0.2557
REMARK   3    11  2.0200 -  1.9600    0.90     1730   140  0.1839 0.2094
REMARK   3    12  1.9600 -  1.9000    0.91     1765   132  0.1913 0.2478
REMARK   3    13  1.9000 -  1.8500    0.92     1730   147  0.1982 0.2816
REMARK   3    14  1.8500 -  1.8100    0.92     1746   142  0.2166 0.2626
REMARK   3    15  1.8100 -  1.7700    0.92     1771   142  0.2499 0.2825
REMARK   3    16  1.7700 -  1.7300    0.92     1746   137  0.2832 0.3166
REMARK   3    17  1.7300 -  1.6900    0.91     1755   146  0.2938 0.3424
REMARK   3    18  1.6900 -  1.6600    0.92     1764   151  0.3403 0.3842
REMARK   3    19  1.6600 -  1.6300    0.92     1738   139  0.3626 0.4019
REMARK   3    20  1.6300 -  1.6100    0.92     1786   146  0.3772 0.3597
REMARK   3    21  1.6100 -  1.5800    0.92     1741   135  0.4034 0.4996
REMARK   3    22  1.5800 -  1.5500    0.92     1770   147  0.4233 0.4493
REMARK   3    23  1.5500 -  1.5300    0.91     1749   132  0.4634 0.4636
REMARK   3    24  1.5300 -  1.5100    0.91     1762   150  0.5087 0.5013
REMARK   3    25  1.5100 -  1.4900    0.92     1715   142  0.5713 0.5672
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : NULL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.232
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 34.996
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 21.11
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 33.37
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.009           3252
REMARK   3   ANGLE     :  1.050           4445
REMARK   3   CHIRALITY :  0.076            469
REMARK   3   PLANARITY :  0.008            595
REMARK   3   DIHEDRAL  : 18.475           1177
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 6SYR COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 01-OCT-19.
REMARK 100 THE DEPOSITION ID IS D_1292104569.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 09-DEC-18
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 7.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID30B
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.918374
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS EIGER X 4M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XDS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 47249
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.490
REMARK 200  RESOLUTION RANGE LOW       (A) : 32.690
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 91.2
REMARK 200  DATA REDUNDANCY                : 3.800
REMARK 200  R MERGE                    (I) : 0.06132
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 9.8800
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.49
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.54
REMARK 200  COMPLETENESS FOR SHELL     (%) : 91.4
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.90
REMARK 200  R MERGE FOR SHELL          (I) : 1.41600
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 0.820
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHENIX 1.15.2_3472
REMARK 200 STARTING MODEL: 6GS0
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): NULL
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): NULL
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: BASED ON THE MORPHEUS SCREEN FROM
REMARK 280  MOLECULAR DIMENSIONS: G12 - 0.1 M CARBOXYLIC ACIDS, 0.1 M BUFFER
REMARK 280  SYSTEM 3, PH 8.5, 50 % V/V PRECIPITANT MIX 4, PH 7.5, VAPOR
REMARK 280  DIFFUSION, SITTING DROP, TEMPERATURE 298K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 1230 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 15550 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -11.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     ARG A     2
REMARK 465     ASN A     3
REMARK 465     VAL A     4
REMARK 465     LEU A     5
REMARK 465     ALA A     6
REMARK 465     ALA A     7
REMARK 465     LEU A     8
REMARK 465     SER A     9
REMARK 465     LEU A    10
REMARK 465     LEU A    11
REMARK 465     PHE A    12
REMARK 465     THR A    13
REMARK 465     LEU A    14
REMARK 465     THR A    15
REMARK 465     SER A    16
REMARK 465     MET A    17
REMARK 465     GLN A    18
REMARK 465     ALA A    19
REMARK 465     THR A    20
REMARK 465     SER A    21
REMARK 465     ARG A    22
REMARK 465     PRO A    23
REMARK 465     ALA A    24
REMARK 465     ARG A    25
REMARK 465     LEU A    26
REMARK 465     ASP A    27
REMARK 465     ASP A    28
REMARK 465     THR A    29
REMARK 465     PRO A    30
REMARK 465     PRO A    31
REMARK 465     PRO A    32
REMARK 465     ALA A    33
REMARK 465     TYR A    34
REMARK 465     ALA A   432
REMARK 465     ARG A   433
REMARK 465     GLU A   434
REMARK 465     PRO A   435
REMARK 465     GLN A   436
REMARK 465     ARG A   437
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500  HH12  ARG A    51     OD2  ASP A   110     1654     1.56
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    VAL A  70      -71.11   -121.73
REMARK 500    ASP A 150      110.84    -35.78
REMARK 500    LEU A 220       31.57    -89.09
REMARK 500    ASP A 221     -118.54    147.34
REMARK 500    ALA A 222      -39.75     58.79
REMARK 500    ALA A 223     -148.83     54.40
REMARK 500    SER A 267     -127.73     57.59
REMARK 500    HIS A 328       37.49   -147.59
REMARK 500    ASP A 360       72.31     58.89
REMARK 500    LYS A 427      151.20    -44.92
REMARK 500
REMARK 500 REMARK: NULL
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA A 507  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 LEU A 250   O
REMARK 620 2 ASP A 253   O    93.8
REMARK 620 3 VAL A 256   O   114.6  79.9
REMARK 620 4 HOH A 750   O   109.0 156.3  84.5
REMARK 620 5 HOH A 778   O   133.5  86.6 111.2  82.5
REMARK 620 N                    1     2     3     4
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              MG A 502  MG
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HIS A 303   O
REMARK 620 2 GLU A 305   O    93.7
REMARK 620 3 HOH A 727   O   100.8  94.5
REMARK 620 4 HOH A 605   O    89.2  89.5 169.0
REMARK 620 5 HOH A 802   O   163.6  88.9  95.1  74.7
REMARK 620 6 HOH A 722   O    92.1 162.0 101.2  73.5  80.9
REMARK 620 N                    1     2     3     4     5
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GCU A 501
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue MG A 502
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 504
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 505
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 506
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NA A 507
DBREF  6SYR A    1   437  UNP    B1ZMF4   B1ZMF4_OPITP     1    437
SEQRES   1 A  437  MET ARG ASN VAL LEU ALA ALA LEU SER LEU LEU PHE THR
SEQRES   2 A  437  LEU THR SER MET GLN ALA THR SER ARG PRO ALA ARG LEU
SEQRES   3 A  437  ASP ASP THR PRO PRO PRO ALA TYR THR LEU PRO ASP PRO
SEQRES   4 A  437  LEU VAL GLY ALA ASP GLY THR ARG VAL HIS ASP ARG ALA
SEQRES   5 A  437  THR TRP GLN HIS ARG ARG ARG PRO GLU LEU LEU GLN LEU
SEQRES   6 A  437  PHE ALA ARG GLU VAL TYR GLY ARG THR PRO LEU GLY ARG
SEQRES   7 A  437  PRO GLU GLY MET VAL PHE LYS VAL THR THR MET GLU HIS
SEQRES   8 A  437  ALA ALA LEU GLY GLY ALA ALA THR ARG LYS GLU VAL THR
SEQRES   9 A  437  VAL ARG PHE GLY ARG ASP PRO ASN ALA PRO SER MET GLN
SEQRES  10 A  437  LEU LEU LEU TYR VAL PRO ASN ALA VAL ILE ALA ARG ALA
SEQRES  11 A  437  GLU ARG ALA PRO VAL PHE LEU GLY LEU ASN PHE TYR GLY
SEQRES  12 A  437  ASN HIS THR VAL HIS THR ASP PRO ALA ILE ALA LEU SER
SEQRES  13 A  437  ALA ARG TRP ILE PRO ALA GLU ALA PRO ASN GLY ALA ASN
SEQRES  14 A  437  HIS ARG ALA THR GLU ALA ALA ARG GLY SER ASP ALA GLN
SEQRES  15 A  437  LYS TRP PRO VAL GLU GLN ILE LEU ALA ARG GLY TYR ALA
SEQRES  16 A  437  VAL ALA THR VAL TYR CYS GLY ASP LEU CYS PRO ASP ARG
SEQRES  17 A  437  PRO ASP GLY LEU ASN ALA SER VAL ALA SER TRP LEU ASP
SEQRES  18 A  437  ALA ALA ALA GLY ASP GLN ARG ALA PRO ASP ALA TRP GLY
SEQRES  19 A  437  ALA ILE GLY VAL TRP ALA TRP GLY LEU SER ARG ALA LEU
SEQRES  20 A  437  ASP TYR LEU GLU THR ASP PRO LEU VAL ASP ALA SER ARG
SEQRES  21 A  437  VAL ALA VAL HIS GLY HIS SER ARG LEU GLY LYS ALA ALA
SEQRES  22 A  437  LEU TRP ALA GLY ALA GLN ASP ASP ARG PHE ALA LEU VAL
SEQRES  23 A  437  ILE SER ASN GLU SER GLY CYS GLY GLY ALA ALA LEU SER
SEQRES  24 A  437  LYS ARG ILE HIS GLY GLU THR VAL ALA ARG ILE ASN THR
SEQRES  25 A  437  VAL PHE PRO HIS TRP PHE ALA ARG ASN PHE ARG ARG TYR
SEQRES  26 A  437  ASP ASP HIS GLU GLU ALA LEU PRO VAL ASP GLN HIS GLU
SEQRES  27 A  437  LEU LEU ALA LEU VAL ALA PRO ARG PRO LEU TYR VAL ALA
SEQRES  28 A  437  SER ALA GLU ASP ASP ASP TRP ALA ASP PRO ARG GLY GLU
SEQRES  29 A  437  PHE LEU ALA VAL LYS ALA ALA GLU PRO VAL PHE ARG LEU
SEQRES  30 A  437  PHE GLY GLN THR GLY PRO SER GLY GLU ASP VAL PRO ARG
SEQRES  31 A  437  VAL ASN GLU PRO SER GLY GLY ALA LEU ARG TYR HIS ILE
SEQRES  32 A  437  ARG PRO GLY PRO HIS GLY MET THR ALA GLN ASP TRP ALA
SEQRES  33 A  437  PHE TYR LEU ALA PHE ALA ASP GLU TRP LEU LYS SER ALA
SEQRES  34 A  437  LEU PRO ALA ARG GLU PRO GLN ARG
HET    GCU  A 501      22
HET     MG  A 502       1
HET    EDO  A 503      10
HET    EDO  A 504      10
HET    EDO  A 505      10
HET    EDO  A 506      10
HET     NA  A 507       1
HETNAM     GCU D-GLUCURONIC ACID
HETNAM      MG MAGNESIUM ION
HETNAM     EDO 1,2-ETHANEDIOL
HETNAM      NA SODIUM ION
HETSYN     EDO ETHYLENE GLYCOL
FORMUL   2  GCU    C6 H10 O7
FORMUL   3   MG    MG 2+
FORMUL   4  EDO    4(C2 H6 O2)
FORMUL   8   NA    NA 1+
FORMUL   9  HOH   *227(H2 O)
HELIX    1 AA1 ASP A   50  ARG A   57  1                                   8
HELIX    2 AA2 ARG A   57  VAL A   70  1                                  14
HELIX    3 AA3 LEU A   94  GLY A   96  5                                   3
HELIX    4 AA4 ASN A  124  ALA A  128  1                                   5
HELIX    5 AA5 GLY A  143  VAL A  147  5                                   5
HELIX    6 AA6 THR A  173  ARG A  177  5                                   5
HELIX    7 AA7 ASP A  180  TRP A  184  5                                   5
HELIX    8 AA8 PRO A  185  ARG A  192  1                                   8
HELIX    9 AA9 GLY A  202  LEU A  204  5                                   3
HELIX   10 AB1 ASP A  210  ALA A  214  5                                   5
HELIX   11 AB2 SER A  215  LEU A  220  1                                   6
HELIX   12 AB3 GLY A  234  ASP A  253  1                                  20
HELIX   13 AB4 SER A  267  ASP A  280  1                                  14
HELIX   14 AB5 THR A  306  PHE A  314  1                                   9
HELIX   15 AB6 ALA A  319  ASP A  326  5                                   8
HELIX   16 AB7 HIS A  328  LEU A  332  5                                   5
HELIX   17 AB8 ASP A  335  LEU A  342  1                                   8
HELIX   18 AB9 ASP A  356  ALA A  359  5                                   4
HELIX   19 AC1 ASP A  360  PHE A  378  1                                  19
HELIX   20 AC2 THR A  411  LEU A  426  1                                  16
SHEET    1 AA1 9 VAL A  83  ALA A  93  0
SHEET    2 AA1 9 ALA A  98  ARG A 106 -1  O  ARG A 106   N  VAL A  83
SHEET    3 AA1 9 SER A 115  PRO A 123 -1  O  LEU A 118   N  VAL A 103
SHEET    4 AA1 9 ALA A 195  TYR A 200 -1  O  VAL A 196   N  TYR A 121
SHEET    5 AA1 9 ALA A 133  ASN A 140  1  N  PHE A 136   O  ALA A 197
SHEET    6 AA1 9 VAL A 256  HIS A 266  1  O  ALA A 262   N  LEU A 137
SHEET    7 AA1 9 LEU A 285  ASN A 289  1  O  LEU A 285   N  VAL A 263
SHEET    8 AA1 9 LEU A 348  ALA A 353  1  O  TYR A 349   N  SER A 288
SHEET    9 AA1 9 LEU A 399  ARG A 404  1  O  ARG A 400   N  VAL A 350
LINK         O   LEU A 250                NA    NA A 507     1555   1555  2.46
LINK         O   ASP A 253                NA    NA A 507     1555   1555  2.53
LINK         O   VAL A 256                NA    NA A 507     1555   1555  2.65
LINK         O   HIS A 303                MG    MG A 502     1555   1555  2.18
LINK         O   GLU A 305                MG    MG A 502     1555   1555  2.25
LINK        MG    MG A 502                 O   HOH A 727     1555   1555  2.42
LINK        MG    MG A 502                 O   HOH A 605     1555   1555  2.63
LINK        MG    MG A 502                 O   HOH A 802     1555   1555  2.52
LINK        MG    MG A 502                 O   HOH A 722     1555   1555  2.18
LINK        NA    NA A 507                 O   HOH A 750     1555   1555  2.26
LINK        NA    NA A 507                 O   HOH A 778     1555   1555  2.49
CISPEP   1 ALA A  344    PRO A  345          0         0.68
SITE     1 AC1 12 SER A 267  ARG A 268  LYS A 271  GLU A 305
SITE     2 AC1 12 ILE A 310  PHE A 314  TRP A 358  HIS A 408
SITE     3 AC1 12 HOH A 655  HOH A 726  HOH A 768  HOH A 775
SITE     1 AC2  6 HIS A 303  GLU A 305  HOH A 605  HOH A 722
SITE     2 AC2  6 HOH A 727  HOH A 802
SITE     1 AC3  1 HOH A 606
SITE     1 AC4  3 GLU A 131  ASP A 226  PRO A 254
SITE     1 AC5  6 PRO A  75  LEU A  76  ALA A  92  LEU A  94
SITE     2 AC5  6 GLY A  95  HOH A 640
SITE     1 AC6  6 LEU A 250  GLU A 251  ASP A 253  VAL A 256
SITE     2 AC6  6 HOH A 750  HOH A 778
CRYST1   43.215   44.189   50.172  75.78  65.43  70.85 P 1           1
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.023140 -0.008037 -0.009638        0.00000
SCALE2      0.000000  0.023956 -0.003072        0.00000
SCALE3      0.000000  0.000000  0.022096        0.00000
TER    6102      PRO A 431
MASTER      349    0    7   20    9    0   11    6 3326    1   77   34
END