longtext: 6t2h-pdb

content
HEADER    HYDROLASE                               08-OCT-19   6T2H
TITLE     FURANO[2,3-D]PRYMIDINE AMIDES AS NOTUM INHIBITORS
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: PALMITOLEOYL-PROTEIN CARBOXYLESTERASE NOTUM;
COMPND   3 CHAIN: A;
COMPND   4 SYNONYM: HNOTUM;
COMPND   5 EC: 3.1.1.98;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: NOTUM, OK/SW-CL.30;
SOURCE   6 EXPRESSION_SYSTEM: HOMO SAPIENS;
SOURCE   7 EXPRESSION_SYSTEM_COMMON: HUMAN;
SOURCE   8 EXPRESSION_SYSTEM_TAXID: 9606;
SOURCE   9 EXPRESSION_SYSTEM_CELL_LINE: HEK293S GNTI-
KEYWDS    SUBSTRATE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    Y.ZHAO,E.Y.JONES
REVDAT   1   01-JAN-20 6T2H    0
JRNL        AUTH   B.N.ATKINSON,D.STEADMAN,W.MAHY,Y.ZHAO,J.SIPTHORP,E.D.BAYLE,
JRNL        AUTH 2 F.SVENSSON,G.PAPAGEORGIOU,F.JEGANATHAN,S.FREW,A.MONAGHAN,
JRNL        AUTH 3 M.BICTASH,E.Y.JONES,P.V.FISH
JRNL        TITL   SCAFFOLD-HOPPING IDENTIFIES FURANO[2,3-D]PYRIMIDINE AMIDES
JRNL        TITL 2 AS POTENT NOTUM INHIBITORS.
JRNL        REF    BIOORG.MED.CHEM.LETT.                26751 2019
JRNL        REFN                   ESSN 1464-3405
JRNL        PMID   31862412
JRNL        DOI    10.1016/J.BMCL.2019.126751
REMARK   2
REMARK   2 RESOLUTION.    1.41 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX DEV_2645
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.41
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 32.69
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.340
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.9
REMARK   3   NUMBER OF REFLECTIONS             : 65466
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.178
REMARK   3   R VALUE            (WORKING SET) : 0.176
REMARK   3   FREE R VALUE                     : 0.198
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.110
REMARK   3   FREE R VALUE TEST SET COUNT      : 3348
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 32.6920 -  4.0647    1.00     2775   163  0.1768 0.1849
REMARK   3     2  4.0647 -  3.2272    1.00     2673   142  0.1678 0.1717
REMARK   3     3  3.2272 -  2.8195    1.00     2639   141  0.1781 0.2167
REMARK   3     4  2.8195 -  2.5618    1.00     2632   138  0.1799 0.2169
REMARK   3     5  2.5618 -  2.3782    1.00     2604   131  0.1733 0.1924
REMARK   3     6  2.3782 -  2.2381    1.00     2589   154  0.1676 0.1944
REMARK   3     7  2.2381 -  2.1260    1.00     2578   135  0.1613 0.1815
REMARK   3     8  2.1260 -  2.0335    1.00     2621   141  0.1629 0.1869
REMARK   3     9  2.0335 -  1.9552    1.00     2548   161  0.1587 0.2160
REMARK   3    10  1.9552 -  1.8877    1.00     2593   134  0.1604 0.1779
REMARK   3    11  1.8877 -  1.8287    1.00     2565   134  0.1587 0.1930
REMARK   3    12  1.8287 -  1.7764    1.00     2578   143  0.1554 0.1704
REMARK   3    13  1.7764 -  1.7297    1.00     2553   155  0.1703 0.1961
REMARK   3    14  1.7297 -  1.6875    1.00     2550   143  0.1842 0.2224
REMARK   3    15  1.6875 -  1.6491    1.00     2587   138  0.1878 0.2195
REMARK   3    16  1.6491 -  1.6140    1.00     2546   127  0.1904 0.2230
REMARK   3    17  1.6140 -  1.5817    1.00     2560   126  0.1977 0.2133
REMARK   3    18  1.5817 -  1.5519    1.00     2586   136  0.2034 0.2467
REMARK   3    19  1.5519 -  1.5242    1.00     2576   122  0.2096 0.2233
REMARK   3    20  1.5242 -  1.4983    1.00     2512   150  0.2341 0.2768
REMARK   3    21  1.4983 -  1.4742    1.00     2592   128  0.2340 0.2870
REMARK   3    22  1.4742 -  1.4515    1.00     2522   152  0.2628 0.2915
REMARK   3    23  1.4515 -  1.4302    1.00     2574   136  0.2665 0.2825
REMARK   3    24  1.4302 -  1.4100    1.00     2565   118  0.2944 0.3180
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : NULL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.140
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 20.380
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 20.21
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 27.95
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.010           3116
REMARK   3   ANGLE     :  1.134           4233
REMARK   3   CHIRALITY :  0.102            439
REMARK   3   PLANARITY :  0.008            539
REMARK   3   DIHEDRAL  : 13.532           1776
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 1
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: ALL
REMARK   3    ORIGIN FOR THE GROUP (A):   4.8046  -1.4878  -2.0527
REMARK   3    T TENSOR
REMARK   3      T11:   0.1324 T22:   0.1491
REMARK   3      T33:   0.1406 T12:  -0.0021
REMARK   3      T13:  -0.0013 T23:   0.0185
REMARK   3    L TENSOR
REMARK   3      L11:   0.7816 L22:   1.0821
REMARK   3      L33:   0.7160 L12:   0.0475
REMARK   3      L13:  -0.0727 L23:   0.0547
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0186 S12:  -0.0094 S13:   0.0159
REMARK   3      S21:   0.0302 S22:   0.0053 S23:   0.0013
REMARK   3      S31:  -0.0428 S32:   0.0045 S33:   0.0125
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 6T2H COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 09-OCT-19.
REMARK 100 THE DEPOSITION ID IS D_1292104596.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 28-APR-17
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 4.2
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : DIAMOND
REMARK 200  BEAMLINE                       : I03
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9726
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 6M-F
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DIALS
REMARK 200  DATA SCALING SOFTWARE          : XIA2
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 72713
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.360
REMARK 200  RESOLUTION RANGE LOW       (A) : 78.250
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.6
REMARK 200  DATA REDUNDANCY                : 12.40
REMARK 200  R MERGE                    (I) : 0.08700
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 35.3000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.36
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.38
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: 6R8P
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 36.12
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 1.93
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 1.5M AMMONIUM SULFATE 0.1 M SODIUM
REMARK 280  CITRATE PH4.2, EVAPORATION, TEMPERATURE 300K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       29.86000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       39.06500
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       35.94500
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       39.06500
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       29.86000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       35.94500
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 4390 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 15940 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -35.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLU A    78
REMARK 465     THR A    79
REMARK 465     GLY A    80
REMARK 465     SER A    81
REMARK 465     ALA A    82
REMARK 465     GLN A    83
REMARK 465     GLN A    84
REMARK 465     LEU A    85
REMARK 465     ASN A    86
REMARK 465     ASP A   278
REMARK 465     CYS A   279
REMARK 465     VAL A   280
REMARK 465     ASP A   281
REMARK 465     ASP A   420
REMARK 465     SER A   421
REMARK 465     HIS A   422
REMARK 465     LYS A   423
REMARK 465     ALA A   424
REMARK 465     SER A   425
REMARK 465     LYS A   426
REMARK 465     GLY A   452
REMARK 465     THR A   453
REMARK 465     LYS A   454
REMARK 465     HIS A   455
REMARK 465     HIS A   456
REMARK 465     HIS A   457
REMARK 465     HIS A   458
REMARK 465     HIS A   459
REMARK 465     HIS A   460
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     GLU A  87    CB   CG   CD   OE1  OE2
REMARK 470     THR A 277    OG1  CG2
REMARK 470     THR A 427    OG1  CG2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   OE2  GLU A   167     OG1  THR A   282     2554     2.18
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    TRP A 128     -141.86     62.08
REMARK 500    SER A 232     -124.13     63.12
REMARK 500    SER A 232     -125.84     65.48
REMARK 500    HIS A 276       75.43   -103.88
REMARK 500    HIS A 276       74.73   -103.30
REMARK 500    GLN A 311     -175.72     68.00
REMARK 500    THR A 352      -70.51   -112.45
REMARK 500    SER A 388     -169.12   -164.97
REMARK 500    GLU A 390      161.44     61.86
REMARK 500    ILE A 391      -39.43   -146.92
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue DMS A 501
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue DMS A 502
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 503
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 504
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 505
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 506
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 507
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 508
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 A 509
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 510
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 511
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 512
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 513
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 514
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 515
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 516
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 517
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 518
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 519
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 520
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 521
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 522
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue M9N A 524
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Mono-Saccharide NAG A 523 bound
REMARK 800  to ASN A 96
DBREF  6T2H A   81   451  UNP    Q6P988   NOTUM_HUMAN     81    451
SEQADV 6T2H GLU A   78  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H THR A   79  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H GLY A   80  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H SER A  330  UNP  Q6P988    CYS   330 ENGINEERED MUTATION
SEQADV 6T2H GLY A  452  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H THR A  453  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H LYS A  454  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H HIS A  455  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H HIS A  456  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H HIS A  457  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H HIS A  458  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H HIS A  459  UNP  Q6P988              EXPRESSION TAG
SEQADV 6T2H HIS A  460  UNP  Q6P988              EXPRESSION TAG
SEQRES   1 A  383  GLU THR GLY SER ALA GLN GLN LEU ASN GLU ASP LEU ARG
SEQRES   2 A  383  LEU HIS LEU LEU LEU ASN THR SER VAL THR CYS ASN ASP
SEQRES   3 A  383  GLY SER PRO ALA GLY TYR TYR LEU LYS GLU SER ARG GLY
SEQRES   4 A  383  SER ARG ARG TRP LEU LEU PHE LEU GLU GLY GLY TRP TYR
SEQRES   5 A  383  CYS PHE ASN ARG GLU ASN CYS ASP SER ARG TYR ASP THR
SEQRES   6 A  383  MET ARG ARG LEU MET SER SER ARG ASP TRP PRO ARG THR
SEQRES   7 A  383  ARG THR GLY THR GLY ILE LEU SER SER GLN PRO GLU GLU
SEQRES   8 A  383  ASN PRO TYR TRP TRP ASN ALA ASN MET VAL PHE ILE PRO
SEQRES   9 A  383  TYR CYS SER SER ASP VAL TRP SER GLY ALA SER SER LYS
SEQRES  10 A  383  SER GLU LYS ASN GLU TYR ALA PHE MET GLY ALA LEU ILE
SEQRES  11 A  383  ILE GLN GLU VAL VAL ARG GLU LEU LEU GLY ARG GLY LEU
SEQRES  12 A  383  SER GLY ALA LYS VAL LEU LEU LEU ALA GLY SER SER ALA
SEQRES  13 A  383  GLY GLY THR GLY VAL LEU LEU ASN VAL ASP ARG VAL ALA
SEQRES  14 A  383  GLU GLN LEU GLU LYS LEU GLY TYR PRO ALA ILE GLN VAL
SEQRES  15 A  383  ARG GLY LEU ALA ASP SER GLY TRP PHE LEU ASP ASN LYS
SEQRES  16 A  383  GLN TYR ARG HIS THR ASP CYS VAL ASP THR ILE THR CYS
SEQRES  17 A  383  ALA PRO THR GLU ALA ILE ARG ARG GLY ILE ARG TYR TRP
SEQRES  18 A  383  ASN GLY VAL VAL PRO GLU ARG CYS ARG ARG GLN PHE GLN
SEQRES  19 A  383  GLU GLY GLU GLU TRP ASN CYS PHE PHE GLY TYR LYS VAL
SEQRES  20 A  383  TYR PRO THR LEU ARG SER PRO VAL PHE VAL VAL GLN TRP
SEQRES  21 A  383  LEU PHE ASP GLU ALA GLN LEU THR VAL ASP ASN VAL HIS
SEQRES  22 A  383  LEU THR GLY GLN PRO VAL GLN GLU GLY LEU ARG LEU TYR
SEQRES  23 A  383  ILE GLN ASN LEU GLY ARG GLU LEU ARG HIS THR LEU LYS
SEQRES  24 A  383  ASP VAL PRO ALA SER PHE ALA PRO ALA CYS LEU SER HIS
SEQRES  25 A  383  GLU ILE ILE ILE ARG SER HIS TRP THR ASP VAL GLN VAL
SEQRES  26 A  383  LYS GLY THR SER LEU PRO ARG ALA LEU HIS CYS TRP ASP
SEQRES  27 A  383  ARG SER LEU HIS ASP SER HIS LYS ALA SER LYS THR PRO
SEQRES  28 A  383  LEU LYS GLY CYS PRO VAL HIS LEU VAL ASP SER CYS PRO
SEQRES  29 A  383  TRP PRO HIS CYS ASN PRO SER CYS PRO THR GLY THR LYS
SEQRES  30 A  383  HIS HIS HIS HIS HIS HIS
HET    DMS  A 501       4
HET    DMS  A 502       4
HET    SO4  A 503       5
HET    SO4  A 504       5
HET    SO4  A 505       5
HET    SO4  A 506       5
HET    SO4  A 507       5
HET    SO4  A 508       5
HET    SO4  A 509       5
HET    EDO  A 510       4
HET    EDO  A 511       4
HET    EDO  A 512       4
HET    EDO  A 513       4
HET    EDO  A 514       4
HET    EDO  A 515       4
HET    EDO  A 516       4
HET    EDO  A 517       4
HET    EDO  A 518       4
HET    EDO  A 519       4
HET    EDO  A 520       4
HET    EDO  A 521       4
HET    EDO  A 522       4
HET    NAG  A 523      14
HET    M9N  A 524      16
HETNAM     DMS DIMETHYL SULFOXIDE
HETNAM     SO4 SULFATE ION
HETNAM     EDO 1,2-ETHANEDIOL
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     M9N 2-[[(4~{S})-5-CHLORANYL-6-METHYL-1,2,3,4-
HETNAM   2 M9N  TETRAHYDROTHIENO[2,3-D]PYRIMIDIN-4-
HETNAM   3 M9N  YL]SULFANYL]ETHANOIC ACID
HETSYN     EDO ETHYLENE GLYCOL
FORMUL   2  DMS    2(C2 H6 O S)
FORMUL   4  SO4    7(O4 S 2-)
FORMUL  11  EDO    13(C2 H6 O2)
FORMUL  24  NAG    C8 H15 N O6
FORMUL  25  M9N    C9 H11 CL N2 O2 S2
FORMUL  26  HOH   *109(H2 O)
HELIX    1 AA1 ASN A  132  MET A  143  1                                  12
HELIX    2 AA2 ARG A  144  MET A  147  5                                   4
HELIX    3 AA3 THR A  159  SER A  163  5                                   5
HELIX    4 AA4 MET A  203  GLY A  217  1                                  15
HELIX    5 AA5 ARG A  218  ALA A  223  5                                   6
HELIX    6 AA6 SER A  232  LEU A  252  1                                  21
HELIX    7 AA7 ALA A  286  ASN A  299  1                                  14
HELIX    8 AA8 PRO A  303  GLN A  311  1                                   9
HELIX    9 AA9 GLU A  314  PHE A  319  5                                   6
HELIX   10 AB1 PHE A  320  TYR A  325  1                                   6
HELIX   11 AB2 PRO A  326  LEU A  328  5                                   3
HELIX   12 AB3 GLU A  341  ASP A  347  1                                   7
HELIX   13 AB4 GLN A  357  LEU A  375  1                                  19
HELIX   14 AB5 LEU A  407  LEU A  418  1                                  12
SHEET    1 AA110 THR A 155  ARG A 156  0
SHEET    2 AA110 LEU A  89  LEU A  93 -1  N  LEU A  89   O  ARG A 156
SHEET    3 AA110 GLY A 108  LYS A 112 -1  O  TYR A 109   N  HIS A  92
SHEET    4 AA110 ASN A 176  ILE A 180 -1  O  PHE A 179   N  TYR A 110
SHEET    5 AA110 ARG A 119  LEU A 124  1  N  PHE A 123   O  ILE A 180
SHEET    6 AA110 VAL A 225  SER A 231  1  O  LEU A 227   N  TRP A 120
SHEET    7 AA110 GLN A 258  ASP A 264  1  O  ARG A 260   N  LEU A 228
SHEET    8 AA110 VAL A 332  VAL A 335  1  O  VAL A 335   N  ALA A 263
SHEET    9 AA110 SER A 381  ALA A 383  1  O  PHE A 382   N  VAL A 334
SHEET   10 AA110 HIS A 435  VAL A 437  1  O  LEU A 436   N  SER A 381
SHEET    1 AA2 2 PHE A 339  ASP A 340  0
SHEET    2 AA2 2 LEU A 387  SER A 388  1  O  SER A 388   N  PHE A 339
SHEET    1 AA3 2 GLN A 401  VAL A 402  0
SHEET    2 AA3 2 THR A 405  SER A 406 -1  O  THR A 405   N  VAL A 402
SSBOND   1 CYS A  101    CYS A  183                          1555   1555  2.07
SSBOND   2 CYS A  130    CYS A  136                          1555   1555  2.04
SSBOND   3 CYS A  306    CYS A  318                          1555   1555  2.10
SSBOND   4 CYS A  386    CYS A  449                          1555   1555  2.04
SSBOND   5 CYS A  413    CYS A  432                          1555   1555  2.05
SSBOND   6 CYS A  440    CYS A  445                          1555   1555  2.03
LINK         ND2 ASN A  96                 C1  NAG A 523     1555   1555  1.43
SITE     1 AC1  7 TYR A 110  GLU A 125  TRP A 152  PRO A 153
SITE     2 AC1  7 ARG A 156  PHE A 179  PRO A 181
SITE     1 AC2  2 SER A 149  TRP A 152
SITE     1 AC3  9 ARG A 133  LYS A 197  ASN A 198  GLU A 199
SITE     2 AC3  9 ARG A 275  TYR A 297  EDO A 518  HOH A 668
SITE     3 AC3  9 HOH A 678
SITE     1 AC4  5 ARG A 305  VAL A 356  GLN A 357  GLU A 358
SITE     2 AC4  5 ARG A 361
SITE     1 AC5  4 TYR A 274  ARG A 275  LEU A 360  HOH A 621
SITE     1 AC6  3 ARG A 145  ARG A 416  HIS A 419
SITE     1 AC7  8 THR A 142  MET A 143  ARG A 144  ARG A 145
SITE     2 AC7  8 ARG A 409  HIS A 412  ARG A 416  HOH A 632
SITE     1 AC8  3 ARG A 244  GLU A 247  GLU A 304
SITE     1 AC9  4 ARG A  90  LYS A 112  GLU A 113  HOH A 643
SITE     1 AD1  7 LEU A 252  PRO A 433  VAL A 434  HIS A 435
SITE     2 AD1  7 HOH A 630  HOH A 641  HOH A 685
SITE     1 AD2  6 TYR A 325  SER A 330  PRO A 331  VAL A 332
SITE     2 AD2  6 HOH A 617  HOH A 644
SITE     1 AD3  6 ARG A 119  TYR A 171  TRP A 172  TRP A 173
SITE     2 AD3  6 ASN A 174  EDO A 515
SITE     1 AD4  6 ARG A 156  THR A 157  GLY A 158  THR A 159
SITE     2 AD4  6 PHE A 179  EDO A 520
SITE     1 AD5  5 PRO A 153  ARG A 154  GLN A 209  LYS A 430
SITE     2 AD5  5 NAG A 523
SITE     1 AD6  5 PRO A 170  TYR A 171  HIS A 412  ASP A 415
SITE     2 AD6  5 EDO A 512
SITE     1 AD7  5 ARG A 372  HIS A 373  LYS A 376  HOH A 654
SITE     2 AD7  5 HOH A 658
SITE     1 AD8  4 SER A 117  ARG A 118  ARG A 119  LYS A 224
SITE     1 AD9  6 ARG A 133  TYR A 274  GLY A 359  SO4 A 503
SITE     2 AD9  6 HOH A 645  HOH A 697
SITE     1 AE1  5 ASP A 270  ASN A 271  TYR A 322  HIS A 444
SITE     2 AE1  5 PRO A 447
SITE     1 AE2  3 ARG A 156  THR A 157  EDO A 513
SITE     1 AE3  3 ARG A 394  SER A 395  HIS A 396
SITE     1 AE4  4 TYR A 322  TYR A 325  PRO A 326  THR A 374
SITE     1 AE5 12 GLY A 127  TRP A 128  VAL A 187  SER A 232
SITE     2 AE5 12 ALA A 233  PHE A 268  PRO A 287  PHE A 319
SITE     3 AE5 12 ALA A 342  VAL A 346  HIS A 389  HOH A 602
SITE     1 AE6  7 ASN A  96  SER A  98  VAL A  99  PRO A 153
SITE     2 AE6  7 ARG A 154  THR A 155  EDO A 514
CRYST1   59.720   71.890   78.130  90.00  90.00  90.00 P 21 21 21    4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.016745  0.000000  0.000000        0.00000
SCALE2      0.000000  0.013910  0.000000        0.00000
SCALE3      0.000000  0.000000  0.012799        0.00000
TER    2905      THR A 451
MASTER      419    0   24   14   14    0   41    6 3062    1  138   30
END