longtext: 8bbx-pdb

content
HEADER    HYDROLASE                               14-OCT-22   8BBX
TITLE     STRUCTURE OF PROLYL ENDOPROTEASE FROM ASPERGILLUS NIGER CBS 109712 IN
TITLE    2 SPACE GROUP C222(1)
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ENDOPROTEASE ENDO-PRO;
COMPND   3 CHAIN: A, B;
COMPND   4 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: ASPERGILLUS NIGER;
SOURCE   3 ORGANISM_TAXID: 5061;
SOURCE   4 GENE: ATWU_08740;
SOURCE   5 EXPRESSION_SYSTEM: ASPERGILLUS NIGER;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 5061;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: CBS 109712
KEYWDS    ENDOPROTEASE, PROLINE-SPECIFIC, S28 PEPTIDASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    T.PIJNING,A.VUJICIC-ZAGAR,J.M.VAN DER LAAN,R.M.DE JONG,B.W.DIJKSTRA
REVDAT   1   20-DEC-23 8BBX    0
JRNL        AUTH   T.PIJNING,A.VUJICIC-ZAGAR,J.M.V.D.LAAN,R.M.D.JONG,
JRNL        AUTH 2 C.RAMIREZ-PALACIOS,A.VENTE,L.EDENS,B.W.DIJKSTRA
JRNL        TITL   STRUCTURAL AND TIME-RESOLVED MECHANISTIC INVESTIGATIONS OF
JRNL        TITL 2 PROTEIN HYDROLYSIS BY THE ACIDIC PROLINE-SPECIFIC
JRNL        TITL 3 ENDOPROTEASE FROM ASPERGILLUS NIGER
JRNL        REF    PROTEIN SCI.                               2023
JRNL        REFN                   ESSN 1469-896X
JRNL        DOI    10.1002/PRO.4856
REMARK   2
REMARK   2 RESOLUTION.    2.70 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.17.1_3660
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : GEOSTD + MONOMER LIBRARY + CDL V1.2
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.70
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 49.39
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.350
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.7
REMARK   3   NUMBER OF REFLECTIONS             : 42704
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.206
REMARK   3   R VALUE            (WORKING SET) : 0.204
REMARK   3   FREE R VALUE                     : 0.255
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.020
REMARK   3   FREE R VALUE TEST SET COUNT      : 2144
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 49.3900 -  6.6500    1.00     2868   138  0.1856 0.2297
REMARK   3     2  6.6500 -  5.2800    1.00     2763   158  0.1712 0.1899
REMARK   3     3  5.2800 -  4.6200    1.00     2727   142  0.1525 0.2211
REMARK   3     4  4.6200 -  4.1900    1.00     2715   151  0.1513 0.1874
REMARK   3     5  4.1900 -  3.8900    1.00     2684   154  0.1790 0.2442
REMARK   3     6  3.8900 -  3.6600    1.00     2731   136  0.1928 0.2588
REMARK   3     7  3.6600 -  3.4800    1.00     2710   125  0.2098 0.2786
REMARK   3     8  3.4800 -  3.3300    1.00     2691   152  0.2272 0.2791
REMARK   3     9  3.3300 -  3.2000    1.00     2685   145  0.2491 0.3099
REMARK   3    10  3.2000 -  3.0900    1.00     2689   130  0.2517 0.2671
REMARK   3    11  3.0900 -  2.9900    1.00     2682   141  0.2491 0.3416
REMARK   3    12  2.9900 -  2.9100    1.00     2655   150  0.2597 0.3103
REMARK   3    13  2.9100 -  2.8300    0.99     2665   123  0.2700 0.3598
REMARK   3    14  2.8300 -  2.7600    0.99     2648   161  0.3143 0.3520
REMARK   3    15  2.7600 -  2.7000    0.99     2647   138  0.3442 0.3996
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.442
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 27.464
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 49.49
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 44.82
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.010           8362
REMARK   3   ANGLE     :  1.163          11447
REMARK   3   CHIRALITY :  0.059           1268
REMARK   3   PLANARITY :  0.008           1435
REMARK   3   DIHEDRAL  :  9.916           1325
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : 3
REMARK   3   NCS GROUP : 1
REMARK   3    NCS OPERATOR : 1
REMARK   3     REFERENCE SELECTION: CHAIN 'A'
REMARK   3     SELECTION          : (CHAIN 'B' AND RESID 43 THROUGH 525)
REMARK   3     ATOM PAIRS NUMBER  : NULL
REMARK   3     RMSD               : NULL
REMARK   3   NCS GROUP : 2
REMARK   3    NCS OPERATOR : 1
REMARK   3     REFERENCE SELECTION: CHAIN 'C'
REMARK   3     SELECTION          : CHAIN 'E'
REMARK   3     ATOM PAIRS NUMBER  : NULL
REMARK   3     RMSD               : NULL
REMARK   3    NCS OPERATOR : 2
REMARK   3     REFERENCE SELECTION: CHAIN 'C'
REMARK   3     SELECTION          : CHAIN 'F'
REMARK   3     ATOM PAIRS NUMBER  : NULL
REMARK   3     RMSD               : NULL
REMARK   3    NCS OPERATOR : 3
REMARK   3     REFERENCE SELECTION: CHAIN 'C'
REMARK   3     SELECTION          : CHAIN 'G'
REMARK   3     ATOM PAIRS NUMBER  : NULL
REMARK   3     RMSD               : NULL
REMARK   3    NCS OPERATOR : 4
REMARK   3     REFERENCE SELECTION: CHAIN 'C'
REMARK   3     SELECTION          : CHAIN 'H'
REMARK   3     ATOM PAIRS NUMBER  : NULL
REMARK   3     RMSD               : NULL
REMARK   3    NCS OPERATOR : 5
REMARK   3     REFERENCE SELECTION: CHAIN 'C'
REMARK   3     SELECTION          : CHAIN 'I'
REMARK   3     ATOM PAIRS NUMBER  : NULL
REMARK   3     RMSD               : NULL
REMARK   3    NCS OPERATOR : 6
REMARK   3     REFERENCE SELECTION: CHAIN 'C'
REMARK   3     SELECTION          : CHAIN 'K'
REMARK   3     ATOM PAIRS NUMBER  : NULL
REMARK   3     RMSD               : NULL
REMARK   3   NCS GROUP : 3
REMARK   3    NCS OPERATOR : 1
REMARK   3     REFERENCE SELECTION: (CHAIN 'D' AND (RESID 2 OR RESID 5
REMARK   3                          THROUGH 10))
REMARK   3     SELECTION          : (CHAIN 'J' AND (RESID 1 OR RESID 4
REMARK   3                          THROUGH 9))
REMARK   3     ATOM PAIRS NUMBER  : NULL
REMARK   3     RMSD               : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 8BBX COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 24-OCT-22.
REMARK 100 THE DEPOSITION ID IS D_1292125952.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 20-NOV-08
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID23-2
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.8726
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MARMOSAIC 225 MM CCD
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : AIMLESS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 42923
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.690
REMARK 200  RESOLUTION RANGE LOW       (A) : 49.390
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.4
REMARK 200  DATA REDUNDANCY                : 4.000
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 4.9000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.69
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.79
REMARK 200  COMPLETENESS FOR SHELL     (%) : 95.3
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.60
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 1.400
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: 8B57
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 65.43
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.56
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 28-32% (W/V) PEG 3000, 0.1 M SODIUM
REMARK 280  CITRATE BUFFER PH 5.25, PH 6.0, VAPOR DIFFUSION, HANGING DROP,
REMARK 280  TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 2 2 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z+1/2
REMARK 290       3555   -X,Y,-Z+1/2
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z
REMARK 290       6555   -X+1/2,-Y+1/2,Z+1/2
REMARK 290       7555   -X+1/2,Y+1/2,-Z+1/2
REMARK 290       8555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       96.70000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       96.70000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       51.08500
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       78.10300
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       51.08500
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       78.10300
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       96.70000
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       51.08500
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       78.10300
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       96.70000
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       51.08500
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000       78.10300
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, C, D, E, F
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B, G, H, I, J, K
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     ALA A    42
REMARK 465     ALA A   526
REMARK 465     ALA B   526
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   ND2  ASN B   446     O5   NAG I     1              2.17
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    GLU A  59   CA  -  CB  -  CG  ANGL. DEV. = -14.7 DEGREES
REMARK 500    GLU A 101   CA  -  CB  -  CG  ANGL. DEV. = -24.1 DEGREES
REMARK 500    ARG B 121   NE  -  CZ  -  NH1 ANGL. DEV. =   3.4 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    PRO A  58       -9.80    -59.61
REMARK 500    GLU A  88       39.77    -72.37
REMARK 500    THR A 155      -18.22   -141.96
REMARK 500    ASP A 161       87.82   -168.20
REMARK 500    SER A 179     -131.51     55.10
REMARK 500    THR A 195      -60.82   -100.74
REMARK 500    TYR A 212       58.66    -95.54
REMARK 500    ASP A 266      -65.72    -22.51
REMARK 500    ASP A 283      156.57    176.94
REMARK 500    VAL A 348       18.35   -142.01
REMARK 500    PHE A 384      -80.35     62.47
REMARK 500    PRO A 391     -177.82    -64.37
REMARK 500    VAL A 402       92.78    -68.40
REMARK 500    LEU A 475      131.64    -39.76
REMARK 500    PHE A 490     -148.72   -110.60
REMARK 500    CYS A 492       54.78     30.90
REMARK 500    GLU B  88       38.11    -72.29
REMARK 500    ASP B 161       86.99   -168.69
REMARK 500    SER B 179     -129.20     51.08
REMARK 500    TYR B 212       59.57    -94.48
REMARK 500    ASP B 266      -68.00    -19.88
REMARK 500    ASP B 283      158.53    179.03
REMARK 500    VAL B 348       20.57   -140.81
REMARK 500    PHE B 384      -82.44     62.35
REMARK 500    VAL B 402       90.44    -64.32
REMARK 500    PHE B 490     -149.09   -105.90
REMARK 500    CYS B 492       54.79     35.59
REMARK 500
REMARK 500 REMARK: NULL
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 8B57   RELATED DB: PDB
REMARK 900 THE SAME PROTEIN CRYSTALLIZED IN A DIFFERENT SPACE GROUP
DBREF1 8BBX A   42   526  UNP                  A0A8H3Y1T9_ASPTU
DBREF2 8BBX A     A0A8H3Y1T9                         42         526
DBREF1 8BBX B   42   526  UNP                  A0A8H3Y1T9_ASPTU
DBREF2 8BBX B     A0A8H3Y1T9                         42         526
SEQRES   1 A  485  ALA THR THR GLY GLU ALA TYR PHE GLU GLN LEU LEU ASP
SEQRES   2 A  485  HIS HIS ASN PRO GLU LYS GLY THR PHE SER GLN ARG TYR
SEQRES   3 A  485  TRP TRP SER THR GLU TYR TRP GLY GLY PRO GLY SER PRO
SEQRES   4 A  485  VAL VAL LEU PHE THR PRO GLY GLU VAL SER ALA ASP GLY
SEQRES   5 A  485  TYR GLU GLY TYR LEU THR ASN GLU THR LEU THR GLY VAL
SEQRES   6 A  485  TYR ALA GLN GLU ILE GLN GLY ALA VAL ILE LEU ILE GLU
SEQRES   7 A  485  HIS ARG TYR TRP GLY ASP SER SER PRO TYR GLU VAL LEU
SEQRES   8 A  485  ASN ALA GLU THR LEU GLN TYR LEU THR LEU ASP GLN ALA
SEQRES   9 A  485  ILE LEU ASP MET THR TYR PHE ALA GLU THR VAL LYS LEU
SEQRES  10 A  485  GLN PHE ASP ASN SER THR ARG SER ASN ALA GLN ASN ALA
SEQRES  11 A  485  PRO TRP VAL MET VAL GLY GLY SER TYR SER GLY ALA LEU
SEQRES  12 A  485  THR ALA TRP THR GLU SER VAL ALA PRO GLY THR PHE TRP
SEQRES  13 A  485  ALA TYR HIS ALA THR SER ALA PRO VAL GLU ALA ILE TYR
SEQRES  14 A  485  ASP TYR TRP GLN TYR PHE TYR PRO ILE GLN GLN GLY MET
SEQRES  15 A  485  ALA GLN ASN CYS SER LYS ASP VAL SER LEU VAL ALA GLU
SEQRES  16 A  485  TYR VAL ASP LYS ILE GLY LYS ASN GLY THR ALA LYS GLU
SEQRES  17 A  485  GLN GLN ALA LEU LYS GLU LEU PHE GLY LEU GLY ALA VAL
SEQRES  18 A  485  GLU HIS PHE ASP ASP PHE ALA ALA VAL LEU PRO ASN GLY
SEQRES  19 A  485  PRO TYR LEU TRP GLN ASP ASN ASP PHE ALA THR GLY TYR
SEQRES  20 A  485  SER SER PHE PHE GLN PHE CYS ASP ALA VAL GLU GLY VAL
SEQRES  21 A  485  GLU ALA GLY ALA ALA VAL THR PRO GLY PRO GLU GLY VAL
SEQRES  22 A  485  GLY LEU GLU LYS ALA LEU ALA ASN TYR ALA ASN TRP PHE
SEQRES  23 A  485  ASN SER THR ILE LEU PRO ASP TYR CYS ALA SER TYR GLY
SEQRES  24 A  485  TYR TRP THR ASP GLU TRP SER VAL ALA CYS PHE ASP SER
SEQRES  25 A  485  TYR ASN ALA SER SER PRO ILE TYR THR ASP THR SER VAL
SEQRES  26 A  485  GLY ASN ALA VAL ASP ARG GLN TRP GLU TRP PHE LEU CYS
SEQRES  27 A  485  ASN GLU PRO PHE PHE TYR TRP GLN ASP GLY ALA PRO GLU
SEQRES  28 A  485  GLY THR SER THR ILE VAL PRO ARG LEU VAL SER ALA SER
SEQRES  29 A  485  TYR TRP GLN ARG GLN CYS PRO LEU TYR PHE PRO GLU THR
SEQRES  30 A  485  ASN GLY TYR THR TYR GLY SER ALA LYS GLY LYS ASN ALA
SEQRES  31 A  485  ALA THR VAL ASN SER TRP THR GLY GLY TRP ASP MET THR
SEQRES  32 A  485  ARG ASN THR THR ARG LEU ILE TRP THR ASN GLY GLN TYR
SEQRES  33 A  485  ASP PRO TRP ARG ASP SER GLY VAL SER SER THR PHE ARG
SEQRES  34 A  485  PRO GLY GLY PRO LEU ALA SER THR ALA ASN GLU PRO VAL
SEQRES  35 A  485  GLN ILE ILE PRO GLY GLY PHE HIS CYS SER ASP LEU TYR
SEQRES  36 A  485  MET ALA ASP TYR TYR ALA ASN GLU GLY VAL LYS LYS VAL
SEQRES  37 A  485  VAL ASP ASN GLU VAL LYS GLN ILE LYS GLU TRP VAL GLU
SEQRES  38 A  485  GLU TYR TYR ALA
SEQRES   1 B  485  ALA THR THR GLY GLU ALA TYR PHE GLU GLN LEU LEU ASP
SEQRES   2 B  485  HIS HIS ASN PRO GLU LYS GLY THR PHE SER GLN ARG TYR
SEQRES   3 B  485  TRP TRP SER THR GLU TYR TRP GLY GLY PRO GLY SER PRO
SEQRES   4 B  485  VAL VAL LEU PHE THR PRO GLY GLU VAL SER ALA ASP GLY
SEQRES   5 B  485  TYR GLU GLY TYR LEU THR ASN GLU THR LEU THR GLY VAL
SEQRES   6 B  485  TYR ALA GLN GLU ILE GLN GLY ALA VAL ILE LEU ILE GLU
SEQRES   7 B  485  HIS ARG TYR TRP GLY ASP SER SER PRO TYR GLU VAL LEU
SEQRES   8 B  485  ASN ALA GLU THR LEU GLN TYR LEU THR LEU ASP GLN ALA
SEQRES   9 B  485  ILE LEU ASP MET THR TYR PHE ALA GLU THR VAL LYS LEU
SEQRES  10 B  485  GLN PHE ASP ASN SER THR ARG SER ASN ALA GLN ASN ALA
SEQRES  11 B  485  PRO TRP VAL MET VAL GLY GLY SER TYR SER GLY ALA LEU
SEQRES  12 B  485  THR ALA TRP THR GLU SER VAL ALA PRO GLY THR PHE TRP
SEQRES  13 B  485  ALA TYR HIS ALA THR SER ALA PRO VAL GLU ALA ILE TYR
SEQRES  14 B  485  ASP TYR TRP GLN TYR PHE TYR PRO ILE GLN GLN GLY MET
SEQRES  15 B  485  ALA GLN ASN CYS SER LYS ASP VAL SER LEU VAL ALA GLU
SEQRES  16 B  485  TYR VAL ASP LYS ILE GLY LYS ASN GLY THR ALA LYS GLU
SEQRES  17 B  485  GLN GLN ALA LEU LYS GLU LEU PHE GLY LEU GLY ALA VAL
SEQRES  18 B  485  GLU HIS PHE ASP ASP PHE ALA ALA VAL LEU PRO ASN GLY
SEQRES  19 B  485  PRO TYR LEU TRP GLN ASP ASN ASP PHE ALA THR GLY TYR
SEQRES  20 B  485  SER SER PHE PHE GLN PHE CYS ASP ALA VAL GLU GLY VAL
SEQRES  21 B  485  GLU ALA GLY ALA ALA VAL THR PRO GLY PRO GLU GLY VAL
SEQRES  22 B  485  GLY LEU GLU LYS ALA LEU ALA ASN TYR ALA ASN TRP PHE
SEQRES  23 B  485  ASN SER THR ILE LEU PRO ASP TYR CYS ALA SER TYR GLY
SEQRES  24 B  485  TYR TRP THR ASP GLU TRP SER VAL ALA CYS PHE ASP SER
SEQRES  25 B  485  TYR ASN ALA SER SER PRO ILE TYR THR ASP THR SER VAL
SEQRES  26 B  485  GLY ASN ALA VAL ASP ARG GLN TRP GLU TRP PHE LEU CYS
SEQRES  27 B  485  ASN GLU PRO PHE PHE TYR TRP GLN ASP GLY ALA PRO GLU
SEQRES  28 B  485  GLY THR SER THR ILE VAL PRO ARG LEU VAL SER ALA SER
SEQRES  29 B  485  TYR TRP GLN ARG GLN CYS PRO LEU TYR PHE PRO GLU THR
SEQRES  30 B  485  ASN GLY TYR THR TYR GLY SER ALA LYS GLY LYS ASN ALA
SEQRES  31 B  485  ALA THR VAL ASN SER TRP THR GLY GLY TRP ASP MET THR
SEQRES  32 B  485  ARG ASN THR THR ARG LEU ILE TRP THR ASN GLY GLN TYR
SEQRES  33 B  485  ASP PRO TRP ARG ASP SER GLY VAL SER SER THR PHE ARG
SEQRES  34 B  485  PRO GLY GLY PRO LEU ALA SER THR ALA ASN GLU PRO VAL
SEQRES  35 B  485  GLN ILE ILE PRO GLY GLY PHE HIS CYS SER ASP LEU TYR
SEQRES  36 B  485  MET ALA ASP TYR TYR ALA ASN GLU GLY VAL LYS LYS VAL
SEQRES  37 B  485  VAL ASP ASN GLU VAL LYS GLN ILE LYS GLU TRP VAL GLU
SEQRES  38 B  485  GLU TYR TYR ALA
HET    NAG  C   1      14
HET    NAG  C   2      14
HET    NAG  D   1      14
HET    NAG  D   2      14
HET    BMA  D   3      11
HET    MAN  D   4      11
HET    MAN  D   5      11
HET    MAN  D   6      11
HET    MAN  D   7      11
HET    MAN  D   8      11
HET    MAN  D   9      11
HET    MAN  D  10      11
HET    NAG  E   1      14
HET    NAG  E   2      14
HET    NAG  F   1      14
HET    NAG  F   2      14
HET    NAG  G   1      14
HET    NAG  G   2      14
HET    NAG  H   1      14
HET    NAG  H   2      14
HET    NAG  I   1      14
HET    NAG  I   2      14
HET    NAG  J   1      14
HET    NAG  J   2      14
HET    BMA  J   3      11
HET    MAN  J   4      11
HET    MAN  J   5      11
HET    MAN  J   6      11
HET    MAN  J   7      11
HET    MAN  J   8      11
HET    MAN  J   9      11
HET    NAG  K   1      14
HET    NAG  K   2      14
HET    PG4  B 601      13
HETNAM     NAG 2-ACETAMIDO-2-DEOXY-BETA-D-GLUCOPYRANOSE
HETNAM     BMA BETA-D-MANNOPYRANOSE
HETNAM     MAN ALPHA-D-MANNOPYRANOSE
HETNAM     PG4 TETRAETHYLENE GLYCOL
HETSYN     NAG N-ACETYL-BETA-D-GLUCOSAMINE; 2-ACETAMIDO-2-DEOXY-BETA-
HETSYN   2 NAG  D-GLUCOSE; 2-ACETAMIDO-2-DEOXY-D-GLUCOSE; 2-ACETAMIDO-
HETSYN   3 NAG  2-DEOXY-GLUCOSE; N-ACETYL-D-GLUCOSAMINE
HETSYN     BMA BETA-D-MANNOSE; D-MANNOSE; MANNOSE
HETSYN     MAN ALPHA-D-MANNOSE; D-MANNOSE; MANNOSE
FORMUL   3  NAG    18(C8 H15 N O6)
FORMUL   4  BMA    2(C6 H12 O6)
FORMUL   4  MAN    13(C6 H12 O6)
FORMUL  12  PG4    C8 H18 O5
FORMUL  13  HOH   *16(H2 O)
HELIX    1 AA1 TYR A   94  THR A   99  5                                   6
HELIX    2 AA2 THR A  102  GLN A  112  1                                  11
HELIX    3 AA3 GLU A  135  LEU A  140  5                                   6
HELIX    4 AA4 THR A  141  VAL A  156  1                                  16
HELIX    5 AA5 SER A  163  ASN A  167  5                                   5
HELIX    6 AA6 SER A  179  ALA A  192  1                                  14
HELIX    7 AA7 TRP A  213  TYR A  215  5                                   3
HELIX    8 AA8 PHE A  216  MET A  223  1                                   8
HELIX    9 AA9 ALA A  224  GLY A  245  1                                  22
HELIX   10 AB1 THR A  246  PHE A  257  1                                  12
HELIX   11 AB2 HIS A  264  ALA A  270  1                                   7
HELIX   12 AB3 PRO A  273  LEU A  278  1                                   6
HELIX   13 AB4 TRP A  279  ASN A  282  5                                   4
HELIX   14 AB5 SER A  289  GLU A  299  1                                  11
HELIX   15 AB6 GLY A  315  ILE A  331  1                                  17
HELIX   16 AB7 TYR A  335  GLY A  340  5                                   6
HELIX   17 AB8 SER A  358  ASP A  363  1                                   6
HELIX   18 AB9 ASP A  371  GLU A  381  1                                  11
HELIX   19 AC1 SER A  403  ARG A  409  1                                   7
HELIX   20 AC2 GLN A  410  PHE A  415  1                                   6
HELIX   21 AC3 TYR A  423  LYS A  427  5                                   5
HELIX   22 AC4 ASN A  430  GLY A  439  1                                  10
HELIX   23 AC5 GLY A  440  ASN A  446  5                                   7
HELIX   24 AC6 TRP A  460  GLY A  464  5                                   5
HELIX   25 AC7 TYR A  496  ASN A  503  1                                   8
HELIX   26 AC8 ASN A  503  GLU A  523  1                                  21
HELIX   27 AC9 ASN B   57  LYS B   60  5                                   4
HELIX   28 AD1 TYR B   94  THR B   99  5                                   6
HELIX   29 AD2 THR B  102  GLN B  112  1                                  11
HELIX   30 AD3 GLU B  135  LEU B  140  5                                   6
HELIX   31 AD4 THR B  141  VAL B  156  1                                  16
HELIX   32 AD5 SER B  179  ALA B  192  1                                  14
HELIX   33 AD6 TRP B  213  TYR B  215  5                                   3
HELIX   34 AD7 PHE B  216  MET B  223  1                                   8
HELIX   35 AD8 ALA B  224  ASN B  244  1                                  21
HELIX   36 AD9 THR B  246  LEU B  256  1                                  11
HELIX   37 AE1 HIS B  264  ALA B  270  1                                   7
HELIX   38 AE2 PRO B  273  LEU B  278  1                                   6
HELIX   39 AE3 TRP B  279  ASN B  282  5                                   4
HELIX   40 AE4 SER B  289  GLU B  299  1                                  11
HELIX   41 AE5 GLY B  315  ILE B  331  1                                  17
HELIX   42 AE6 TYR B  335  GLY B  340  5                                   6
HELIX   43 AE7 SER B  358  ASP B  363  1                                   6
HELIX   44 AE8 ASP B  371  GLU B  381  1                                  11
HELIX   45 AE9 SER B  403  ARG B  409  1                                   7
HELIX   46 AF1 GLN B  410  PHE B  415  1                                   6
HELIX   47 AF2 TYR B  423  LYS B  427  5                                   5
HELIX   48 AF3 ASN B  430  GLY B  439  1                                  10
HELIX   49 AF4 GLY B  440  ASN B  446  5                                   7
HELIX   50 AF5 TRP B  460  GLY B  464  5                                   5
HELIX   51 AF6 TYR B  496  ASN B  503  1                                   8
HELIX   52 AF7 ASN B  503  GLU B  523  1                                  21
SHEET    1 AA1 8 THR A  44  LEU A  52  0
SHEET    2 AA1 8 THR A  62  SER A  70 -1  O  TYR A  67   N  ALA A  47
SHEET    3 AA1 8 ALA A 114  ILE A 118 -1  O  VAL A 115   N  SER A  70
SHEET    4 AA1 8 VAL A  81  PHE A  84  1  N  VAL A  82   O  ALA A 114
SHEET    5 AA1 8 TRP A 173  GLY A 178  1  O  VAL A 174   N  VAL A  81
SHEET    6 AA1 8 ALA A 198  THR A 202  1  O  HIS A 200   N  MET A 175
SHEET    7 AA1 8 LEU A 450  GLY A 455  1  O  ILE A 451   N  ALA A 201
SHEET    8 AA1 8 VAL A 483  ILE A 486  1  O  ILE A 486   N  ASN A 454
SHEET    1 AA2 2 ILE A 209  ASP A 211  0
SHEET    2 AA2 2 PHE A 384  TRP A 386  1  O  TYR A 385   N  ILE A 209
SHEET    1 AA3 8 THR B  44  LEU B  52  0
SHEET    2 AA3 8 THR B  62  SER B  70 -1  O  GLN B  65   N  PHE B  49
SHEET    3 AA3 8 ALA B 114  ILE B 118 -1  O  LEU B 117   N  TRP B  68
SHEET    4 AA3 8 VAL B  81  PHE B  84  1  N  VAL B  82   O  ALA B 114
SHEET    5 AA3 8 TRP B 173  GLY B 178  1  O  VAL B 174   N  VAL B  81
SHEET    6 AA3 8 ALA B 198  THR B 202  1  O  HIS B 200   N  MET B 175
SHEET    7 AA3 8 LEU B 450  GLY B 455  1  O  ILE B 451   N  TYR B 199
SHEET    8 AA3 8 VAL B 483  ILE B 486  1  O  ILE B 486   N  ASN B 454
SHEET    1 AA4 2 ILE B 209  ASP B 211  0
SHEET    2 AA4 2 PHE B 384  TRP B 386  1  O  TYR B 385   N  ILE B 209
SSBOND   1 CYS A  227    CYS A  295                          1555   1555  2.09
SSBOND   2 CYS A  336    CYS A  350                          1555   1555  2.08
SSBOND   3 CYS A  379    CYS A  411                          1555   1555  2.06
SSBOND   4 CYS B  227    CYS B  295                          1555   1555  2.02
SSBOND   5 CYS B  336    CYS B  350                          1555   1555  2.07
SSBOND   6 CYS B  379    CYS B  411                          1555   1555  2.07
LINK         ND2 ASN A 100                 C1  NAG C   1     1555   1555  1.46
LINK         ND2 ASN A 226                 C1  NAG D   1     1555   1555  1.46
LINK         ND2 ASN A 355                 C1  NAG E   1     1555   1555  1.45
LINK         ND2 ASN A 446                 C1  NAG F   1     1555   1555  1.44
LINK         ND2 ASN B 100                 C1  NAG G   1     1555   1555  1.45
LINK         ND2 ASN B 226                 C1  NAG J   1     1555   1555  1.44
LINK         ND2 ASN B 328                 C1  NAG K   1     1555   1555  1.45
LINK         ND2 ASN B 355                 C1  NAG H   1     1555   1555  1.45
LINK         ND2 ASN B 446                 C1  NAG I   1     1555   1555  1.44
LINK         O4  NAG C   1                 C1  NAG C   2     1555   1555  1.45
LINK         O4  NAG D   1                 C1  NAG D   2     1555   1555  1.43
LINK         O4  NAG D   2                 C1  BMA D   3     1555   1555  1.43
LINK         O3  BMA D   3                 C1  MAN D   4     1555   1555  1.43
LINK         O6  BMA D   3                 C1  MAN D   7     1555   1555  1.44
LINK         O2  MAN D   4                 C1  MAN D   5     1555   1555  1.45
LINK         O2  MAN D   5                 C1  MAN D   6     1555   1555  1.44
LINK         O3  MAN D   7                 C1  MAN D   8     1555   1555  1.45
LINK         O6  MAN D   7                 C1  MAN D  10     1555   1555  1.44
LINK         O3  MAN D   8                 C1  MAN D   9     1555   1555  1.45
LINK         O4  NAG E   1                 C1  NAG E   2     1555   1555  1.45
LINK         O4  NAG F   1                 C1  NAG F   2     1555   1555  1.48
LINK         O4  NAG G   1                 C1  NAG G   2     1555   1555  1.45
LINK         O4  NAG H   1                 C1  NAG H   2     1555   1555  1.45
LINK         O4  NAG I   1                 C1  NAG I   2     1555   1555  1.45
LINK         O4  NAG J   1                 C1  NAG J   2     1555   1555  1.43
LINK         O4  NAG J   2                 C1  BMA J   3     1555   1555  1.44
LINK         O3  BMA J   3                 C1  MAN J   4     1555   1555  1.45
LINK         O6  BMA J   3                 C1  MAN J   7     1555   1555  1.45
LINK         O2  MAN J   4                 C1  MAN J   5     1555   1555  1.44
LINK         O2  MAN J   5                 C1  MAN J   6     1555   1555  1.44
LINK         O3  MAN J   7                 C1  MAN J   8     1555   1555  1.44
LINK         O6  MAN J   7                 C1  MAN J   9     1555   1555  1.45
LINK         O4  NAG K   1                 C1  NAG K   2     1555   1555  1.45
CISPEP   1 GLU A  381    PRO A  382          0        -7.21
CISPEP   2 GLU B  381    PRO B  382          0        -8.49
CRYST1  102.170  156.206  193.400  90.00  90.00  90.00 C 2 2 21     16
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.009788  0.000000  0.000000        0.00000
SCALE2      0.000000  0.006402  0.000000        0.00000
SCALE3      0.000000  0.000000  0.005171        0.00000
TER    3835      TYR A 525
TER    7675      TYR B 525
MASTER      359    0   34   52   20    0    0    6 8119    2  451   76
END