longtext: 8clp-pdb

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HEADER    HYDROLASE                               17-FEB-23   8CLP
TITLE     ZEARALENONE LACTONASE FROM STREPTOMYCES COELICOFLAVUS MUTANT H286Y
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: HYDROLASE;
COMPND   3 CHAIN: A, B, C, D;
COMPND   4 ENGINEERED: YES;
COMPND   5 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: STREPTOMYCES COELICOFLAVUS;
SOURCE   3 ORGANISM_TAXID: 285562;
SOURCE   4 GENE: SMCF_1294;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008
KEYWDS    ZEARALENONE, MYCOTOXIN, LACTONASE, CARBOXYLESTERASE, ESTERASE,
KEYWDS   2 HYDROLASE, BIODEGRADATION, STREPTOMYCES COELICOFLAVUS
EXPDTA    X-RAY DIFFRACTION
AUTHOR    D.PUEHRINGER,G.MLYNEK
REVDAT   1   21-FEB-24 8CLP    0
JRNL        AUTH   S.FRUHAUF,D.PUEHRINGER,M.THAMHESL,P.FAJTL,E.KUNZ-VEKIRO,
JRNL        AUTH 2 A.HOEBARTNER-GUSSL,J.PANHOELZL,K.PREM,V.KLINGENBRUNNER,
JRNL        AUTH 3 G.SCHATZMAYR,G.ADAM,J.DAMBORSKY,K.DJINOVIC-CARUGO,Z.PROKOP,
JRNL        AUTH 4 W.D.MOLL
JRNL        TITL   BACTERIAL LACTONASES ZENA WITH NONCANONICAL STRUCTURAL
JRNL        TITL 2 FEATURES HYDROLYSE THE MYCOTOXIN ZEARALENONE
JRNL        REF    ACS CATALYSIS                              2024
JRNL        REFN                   ESSN 2155-5435
JRNL        DOI    10.1021/ACSCATAL.4C00271
REMARK   2
REMARK   2 RESOLUTION.    1.92 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.20.1-4489
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.92
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 42.57
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.360
REMARK   3   COMPLETENESS FOR RANGE        (%) : 95.9
REMARK   3   NUMBER OF REFLECTIONS             : 76512
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.199
REMARK   3   R VALUE            (WORKING SET) : 0.197
REMARK   3   FREE R VALUE                     : 0.250
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 2.600
REMARK   3   FREE R VALUE TEST SET COUNT      : 1992
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 42.5700 -  4.6300    0.92     5372   140  0.2060 0.2445
REMARK   3     2  4.6300 -  3.6700    0.92     5190   143  0.1481 0.2078
REMARK   3     3  3.6700 -  3.2100    0.94     5287   138  0.1587 0.1936
REMARK   3     4  3.2100 -  2.9200    0.96     5353   142  0.1744 0.2519
REMARK   3     5  2.9100 -  2.7100    0.96     5352   141  0.1830 0.2413
REMARK   3     6  2.7100 -  2.5500    0.97     5319   146  0.1956 0.2465
REMARK   3     7  2.5500 -  2.4200    0.96     5335   140  0.1959 0.2468
REMARK   3     8  2.4200 -  2.3100    0.98     5391   149  0.2110 0.2881
REMARK   3     9  2.3100 -  2.2200    0.97     5347   144  0.2245 0.2568
REMARK   3    10  2.2200 -  2.1500    0.98     5364   143  0.2411 0.3264
REMARK   3    11  2.1500 -  2.0800    0.98     5344   134  0.2526 0.2928
REMARK   3    12  2.0800 -  2.0200    0.97     5362   151  0.2755 0.3417
REMARK   3    13  2.0200 -  1.9700    0.97     5272   145  0.2913 0.3546
REMARK   3    14  1.9700 -  1.9200    0.95     5232   136  0.2998 0.3165
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.10
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.240
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 25.880
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.002           9342
REMARK   3   ANGLE     :  0.506          12739
REMARK   3   CHIRALITY :  0.040           1336
REMARK   3   PLANARITY :  0.008           1681
REMARK   3   DIHEDRAL  :  4.980           1280
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 8CLP COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 17-FEB-23.
REMARK 100 THE DEPOSITION ID IS D_1292126824.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 03-OCT-16
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.9
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID30B
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 76545
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.920
REMARK 200  RESOLUTION RANGE LOW       (A) : 42.570
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 95.9
REMARK 200  DATA REDUNDANCY                : 3.300
REMARK 200  R MERGE                    (I) : 0.10960
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 8.2200
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.92
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.99
REMARK 200  COMPLETENESS FOR SHELL     (%) : 95.6
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.20
REMARK 200  R MERGE FOR SHELL          (I) : 0.76600
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 1.340
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 33.51
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 1.85
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1 M MG FORMATE DIHYDRATE, 15% PEG
REMARK 280  3350 (PH 6.9), VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 295.15K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       31.78450
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       64.35350
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       63.06400
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       64.35350
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       31.78450
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       63.06400
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: TETRAMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: TETRAMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 8210 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 38480 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -4.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     VAL A     2
REMARK 465     THR A     3
REMARK 465     ALA A   188
REMARK 465     SER A   189
REMARK 465     PRO A   190
REMARK 465     MET A   191
REMARK 465     ALA A   192
REMARK 465     ARG A   193
REMARK 465     SER A   194
REMARK 465     PHE A   195
REMARK 465     ALA A   306
REMARK 465     LEU A   307
REMARK 465     ALA A   308
REMARK 465     PRO A   309
REMARK 465     HIS A   310
REMARK 465     HIS A   311
REMARK 465     HIS A   312
REMARK 465     HIS A   313
REMARK 465     HIS A   314
REMARK 465     HIS A   315
REMARK 465     MET B     1
REMARK 465     VAL B     2
REMARK 465     THR B     3
REMARK 465     SER B     4
REMARK 465     PRO B     5
REMARK 465     ALA B     6
REMARK 465     LEU B     7
REMARK 465     ARG B     8
REMARK 465     ASP B     9
REMARK 465     VAL B    10
REMARK 465     HIS B    11
REMARK 465     PHE B   195
REMARK 465     VAL B   196
REMARK 465     ALA B   197
REMARK 465     ASP B   198
REMARK 465     GLY B   199
REMARK 465     PRO B   309
REMARK 465     HIS B   310
REMARK 465     HIS B   311
REMARK 465     HIS B   312
REMARK 465     HIS B   313
REMARK 465     HIS B   314
REMARK 465     HIS B   315
REMARK 465     MET C     1
REMARK 465     VAL C     2
REMARK 465     THR C     3
REMARK 465     SER C     4
REMARK 465     PRO C     5
REMARK 465     ALA C     6
REMARK 465     LEU C     7
REMARK 465     ARG C     8
REMARK 465     ASP C     9
REMARK 465     GLY C   185
REMARK 465     ALA C   186
REMARK 465     SER C   187
REMARK 465     ALA C   188
REMARK 465     SER C   189
REMARK 465     PRO C   190
REMARK 465     MET C   191
REMARK 465     ALA C   192
REMARK 465     ARG C   193
REMARK 465     SER C   194
REMARK 465     PHE C   195
REMARK 465     VAL C   196
REMARK 465     ALA C   197
REMARK 465     ASP C   198
REMARK 465     ALA C   308
REMARK 465     PRO C   309
REMARK 465     HIS C   310
REMARK 465     HIS C   311
REMARK 465     HIS C   312
REMARK 465     HIS C   313
REMARK 465     HIS C   314
REMARK 465     HIS C   315
REMARK 465     MET D     1
REMARK 465     VAL D     2
REMARK 465     GLY D   185
REMARK 465     ALA D   186
REMARK 465     SER D   187
REMARK 465     ALA D   188
REMARK 465     SER D   189
REMARK 465     PRO D   190
REMARK 465     MET D   191
REMARK 465     ALA D   192
REMARK 465     ARG D   193
REMARK 465     SER D   194
REMARK 465     PHE D   195
REMARK 465     VAL D   196
REMARK 465     ALA D   197
REMARK 465     ASP D   198
REMARK 465     ALA D   308
REMARK 465     PRO D   309
REMARK 465     HIS D   310
REMARK 465     HIS D   311
REMARK 465     HIS D   312
REMARK 465     HIS D   313
REMARK 465     HIS D   314
REMARK 465     HIS D   315
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASP A   9       73.19   -106.14
REMARK 500    VAL A  10      -11.00   -144.23
REMARK 500    THR A  48       -4.98     77.50
REMARK 500    SER A 112     -130.06     55.48
REMARK 500    PHE A 141      -44.51     70.63
REMARK 500    ALA A 197       22.15   -163.47
REMARK 500    ASP A 198       30.97   -170.66
REMARK 500    HIS A 242     -145.59    -94.05
REMARK 500    MET A 287       61.03   -105.18
REMARK 500    THR B  48       -8.58     75.05
REMARK 500    SER B 112     -120.97     56.67
REMARK 500    PHE B 141      -51.51     68.96
REMARK 500    HIS B 242     -153.52    -94.21
REMARK 500    MET B 287       75.43   -111.38
REMARK 500    LEU B 307      -74.57    -77.18
REMARK 500    SER C 112     -120.99     52.04
REMARK 500    PHE C 141      -50.40     73.98
REMARK 500    CYS C 224       65.43   -151.29
REMARK 500    HIS C 242     -149.24    -93.59
REMARK 500    MET C 287       74.10   -105.27
REMARK 500    THR D  48       -0.23     74.66
REMARK 500    VAL D 101      -63.15   -101.37
REMARK 500    SER D 112     -126.74     54.31
REMARK 500    PHE D 141      -48.95     71.27
REMARK 500    HIS D 242     -151.20    -83.86
REMARK 500    MET D 287       66.92   -112.54
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: PLANAR GROUPS
REMARK 500
REMARK 500 PLANAR GROUPS IN THE FOLLOWING RESIDUES HAVE A TOTAL
REMARK 500 RMS DISTANCE OF ALL ATOMS FROM THE BEST-FIT PLANE
REMARK 500 BY MORE THAN AN EXPECTED VALUE OF 6*RMSD, WITH AN
REMARK 500 RMSD 0.02 ANGSTROMS, OR AT LEAST ONE ATOM HAS
REMARK 500 AN RMSD GREATER THAN THIS VALUE
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500  M RES CSSEQI        RMS     TYPE
REMARK 500    ARG B 193         0.08    SIDE CHAIN
REMARK 500    ARG C  38         0.10    SIDE CHAIN
REMARK 500    ARG C 104         0.09    SIDE CHAIN
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH D 570        DISTANCE =  6.89 ANGSTROMS
DBREF  8CLP A    1   309  UNP    H1Q8U7   H1Q8U7_9ACTN     1    309
DBREF  8CLP B    1   309  UNP    H1Q8U7   H1Q8U7_9ACTN     1    309
DBREF  8CLP C    1   309  UNP    H1Q8U7   H1Q8U7_9ACTN     1    309
DBREF  8CLP D    1   309  UNP    H1Q8U7   H1Q8U7_9ACTN     1    309
SEQADV 8CLP TYR A  286  UNP  H1Q8U7    HIS   286 ENGINEERED MUTATION
SEQADV 8CLP HIS A  310  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS A  311  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS A  312  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS A  313  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS A  314  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS A  315  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP TYR B  286  UNP  H1Q8U7    HIS   286 ENGINEERED MUTATION
SEQADV 8CLP HIS B  310  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS B  311  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS B  312  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS B  313  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS B  314  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS B  315  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP TYR C  286  UNP  H1Q8U7    HIS   286 ENGINEERED MUTATION
SEQADV 8CLP HIS C  310  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS C  311  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS C  312  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS C  313  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS C  314  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS C  315  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP TYR D  286  UNP  H1Q8U7    HIS   286 ENGINEERED MUTATION
SEQADV 8CLP HIS D  310  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS D  311  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS D  312  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS D  313  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS D  314  UNP  H1Q8U7              EXPRESSION TAG
SEQADV 8CLP HIS D  315  UNP  H1Q8U7              EXPRESSION TAG
SEQRES   1 A  315  MET VAL THR SER PRO ALA LEU ARG ASP VAL HIS VAL PRO
SEQRES   2 A  315  HIS ALA TYR PRO GLU GLN GLN VAL ASP LEU GLY GLU ILE
SEQRES   3 A  315  THR MET ASN TYR ALA GLU ALA GLY ASP PRO ASP ARG PRO
SEQRES   4 A  315  ALA VAL LEU LEU ILE PRO GLU GLN THR GLY SER TRP TRP
SEQRES   5 A  315  SER TYR GLU GLU ALA MET GLY LEU LEU SER GLU HIS PHE
SEQRES   6 A  315  HIS VAL TYR ALA VAL ASP LEU ARG GLY GLN GLY ARG SER
SEQRES   7 A  315  SER TRP THR PRO LYS ARG TYR SER LEU ASP ASN PHE GLY
SEQRES   8 A  315  ASN ASP LEU VAL ARG PHE ILE ALA LEU VAL VAL LYS ARG
SEQRES   9 A  315  PRO VAL VAL VAL ALA GLY ASN SER SER GLY GLY VAL LEU
SEQRES  10 A  315  ALA ALA TRP LEU SER ALA TYR SER MET PRO GLY GLN LEU
SEQRES  11 A  315  ARG GLY VAL LEU CYS GLU ASP PRO PRO PHE PHE ALA SER
SEQRES  12 A  315  GLU LEU VAL PRO ALA HIS GLY HIS SER VAL ARG GLN GLY
SEQRES  13 A  315  ALA GLY PRO VAL PHE GLU LEU PHE ARG THR TYR LEU GLY
SEQRES  14 A  315  ASP GLN TRP SER VAL GLY ASP TRP GLU GLY PHE CYS ARG
SEQRES  15 A  315  ALA ALA GLY ALA SER ALA SER PRO MET ALA ARG SER PHE
SEQRES  16 A  315  VAL ALA ASP GLY ILE PRO GLN HIS LEU GLN GLU TYR ASP
SEQRES  17 A  315  PRO GLU TRP ALA ARG VAL PHE TYR GLU GLY THR VAL GLY
SEQRES  18 A  315  LEU SER CYS PRO HIS GLU ARG MET LEU GLY GLN VAL LYS
SEQRES  19 A  315  THR PRO VAL LEU LEU THR HIS HIS MET ARG GLY ILE ASP
SEQRES  20 A  315  PRO GLU THR GLY ASN LEU LEU GLY ALA LEU SER ASP GLU
SEQRES  21 A  315  GLN ALA LEU ARG ALA ARG ARG LEU MET ASP SER ALA GLY
SEQRES  22 A  315  VAL THR VAL ASP TYR GLU SER VAL PRO ASP ALA SER TYR
SEQRES  23 A  315  MET MET HIS GLN SER ALA PRO ALA ARG TYR VAL GLU ILE
SEQRES  24 A  315  PHE THR ARG TRP ALA ALA ALA LEU ALA PRO HIS HIS HIS
SEQRES  25 A  315  HIS HIS HIS
SEQRES   1 B  315  MET VAL THR SER PRO ALA LEU ARG ASP VAL HIS VAL PRO
SEQRES   2 B  315  HIS ALA TYR PRO GLU GLN GLN VAL ASP LEU GLY GLU ILE
SEQRES   3 B  315  THR MET ASN TYR ALA GLU ALA GLY ASP PRO ASP ARG PRO
SEQRES   4 B  315  ALA VAL LEU LEU ILE PRO GLU GLN THR GLY SER TRP TRP
SEQRES   5 B  315  SER TYR GLU GLU ALA MET GLY LEU LEU SER GLU HIS PHE
SEQRES   6 B  315  HIS VAL TYR ALA VAL ASP LEU ARG GLY GLN GLY ARG SER
SEQRES   7 B  315  SER TRP THR PRO LYS ARG TYR SER LEU ASP ASN PHE GLY
SEQRES   8 B  315  ASN ASP LEU VAL ARG PHE ILE ALA LEU VAL VAL LYS ARG
SEQRES   9 B  315  PRO VAL VAL VAL ALA GLY ASN SER SER GLY GLY VAL LEU
SEQRES  10 B  315  ALA ALA TRP LEU SER ALA TYR SER MET PRO GLY GLN LEU
SEQRES  11 B  315  ARG GLY VAL LEU CYS GLU ASP PRO PRO PHE PHE ALA SER
SEQRES  12 B  315  GLU LEU VAL PRO ALA HIS GLY HIS SER VAL ARG GLN GLY
SEQRES  13 B  315  ALA GLY PRO VAL PHE GLU LEU PHE ARG THR TYR LEU GLY
SEQRES  14 B  315  ASP GLN TRP SER VAL GLY ASP TRP GLU GLY PHE CYS ARG
SEQRES  15 B  315  ALA ALA GLY ALA SER ALA SER PRO MET ALA ARG SER PHE
SEQRES  16 B  315  VAL ALA ASP GLY ILE PRO GLN HIS LEU GLN GLU TYR ASP
SEQRES  17 B  315  PRO GLU TRP ALA ARG VAL PHE TYR GLU GLY THR VAL GLY
SEQRES  18 B  315  LEU SER CYS PRO HIS GLU ARG MET LEU GLY GLN VAL LYS
SEQRES  19 B  315  THR PRO VAL LEU LEU THR HIS HIS MET ARG GLY ILE ASP
SEQRES  20 B  315  PRO GLU THR GLY ASN LEU LEU GLY ALA LEU SER ASP GLU
SEQRES  21 B  315  GLN ALA LEU ARG ALA ARG ARG LEU MET ASP SER ALA GLY
SEQRES  22 B  315  VAL THR VAL ASP TYR GLU SER VAL PRO ASP ALA SER TYR
SEQRES  23 B  315  MET MET HIS GLN SER ALA PRO ALA ARG TYR VAL GLU ILE
SEQRES  24 B  315  PHE THR ARG TRP ALA ALA ALA LEU ALA PRO HIS HIS HIS
SEQRES  25 B  315  HIS HIS HIS
SEQRES   1 C  315  MET VAL THR SER PRO ALA LEU ARG ASP VAL HIS VAL PRO
SEQRES   2 C  315  HIS ALA TYR PRO GLU GLN GLN VAL ASP LEU GLY GLU ILE
SEQRES   3 C  315  THR MET ASN TYR ALA GLU ALA GLY ASP PRO ASP ARG PRO
SEQRES   4 C  315  ALA VAL LEU LEU ILE PRO GLU GLN THR GLY SER TRP TRP
SEQRES   5 C  315  SER TYR GLU GLU ALA MET GLY LEU LEU SER GLU HIS PHE
SEQRES   6 C  315  HIS VAL TYR ALA VAL ASP LEU ARG GLY GLN GLY ARG SER
SEQRES   7 C  315  SER TRP THR PRO LYS ARG TYR SER LEU ASP ASN PHE GLY
SEQRES   8 C  315  ASN ASP LEU VAL ARG PHE ILE ALA LEU VAL VAL LYS ARG
SEQRES   9 C  315  PRO VAL VAL VAL ALA GLY ASN SER SER GLY GLY VAL LEU
SEQRES  10 C  315  ALA ALA TRP LEU SER ALA TYR SER MET PRO GLY GLN LEU
SEQRES  11 C  315  ARG GLY VAL LEU CYS GLU ASP PRO PRO PHE PHE ALA SER
SEQRES  12 C  315  GLU LEU VAL PRO ALA HIS GLY HIS SER VAL ARG GLN GLY
SEQRES  13 C  315  ALA GLY PRO VAL PHE GLU LEU PHE ARG THR TYR LEU GLY
SEQRES  14 C  315  ASP GLN TRP SER VAL GLY ASP TRP GLU GLY PHE CYS ARG
SEQRES  15 C  315  ALA ALA GLY ALA SER ALA SER PRO MET ALA ARG SER PHE
SEQRES  16 C  315  VAL ALA ASP GLY ILE PRO GLN HIS LEU GLN GLU TYR ASP
SEQRES  17 C  315  PRO GLU TRP ALA ARG VAL PHE TYR GLU GLY THR VAL GLY
SEQRES  18 C  315  LEU SER CYS PRO HIS GLU ARG MET LEU GLY GLN VAL LYS
SEQRES  19 C  315  THR PRO VAL LEU LEU THR HIS HIS MET ARG GLY ILE ASP
SEQRES  20 C  315  PRO GLU THR GLY ASN LEU LEU GLY ALA LEU SER ASP GLU
SEQRES  21 C  315  GLN ALA LEU ARG ALA ARG ARG LEU MET ASP SER ALA GLY
SEQRES  22 C  315  VAL THR VAL ASP TYR GLU SER VAL PRO ASP ALA SER TYR
SEQRES  23 C  315  MET MET HIS GLN SER ALA PRO ALA ARG TYR VAL GLU ILE
SEQRES  24 C  315  PHE THR ARG TRP ALA ALA ALA LEU ALA PRO HIS HIS HIS
SEQRES  25 C  315  HIS HIS HIS
SEQRES   1 D  315  MET VAL THR SER PRO ALA LEU ARG ASP VAL HIS VAL PRO
SEQRES   2 D  315  HIS ALA TYR PRO GLU GLN GLN VAL ASP LEU GLY GLU ILE
SEQRES   3 D  315  THR MET ASN TYR ALA GLU ALA GLY ASP PRO ASP ARG PRO
SEQRES   4 D  315  ALA VAL LEU LEU ILE PRO GLU GLN THR GLY SER TRP TRP
SEQRES   5 D  315  SER TYR GLU GLU ALA MET GLY LEU LEU SER GLU HIS PHE
SEQRES   6 D  315  HIS VAL TYR ALA VAL ASP LEU ARG GLY GLN GLY ARG SER
SEQRES   7 D  315  SER TRP THR PRO LYS ARG TYR SER LEU ASP ASN PHE GLY
SEQRES   8 D  315  ASN ASP LEU VAL ARG PHE ILE ALA LEU VAL VAL LYS ARG
SEQRES   9 D  315  PRO VAL VAL VAL ALA GLY ASN SER SER GLY GLY VAL LEU
SEQRES  10 D  315  ALA ALA TRP LEU SER ALA TYR SER MET PRO GLY GLN LEU
SEQRES  11 D  315  ARG GLY VAL LEU CYS GLU ASP PRO PRO PHE PHE ALA SER
SEQRES  12 D  315  GLU LEU VAL PRO ALA HIS GLY HIS SER VAL ARG GLN GLY
SEQRES  13 D  315  ALA GLY PRO VAL PHE GLU LEU PHE ARG THR TYR LEU GLY
SEQRES  14 D  315  ASP GLN TRP SER VAL GLY ASP TRP GLU GLY PHE CYS ARG
SEQRES  15 D  315  ALA ALA GLY ALA SER ALA SER PRO MET ALA ARG SER PHE
SEQRES  16 D  315  VAL ALA ASP GLY ILE PRO GLN HIS LEU GLN GLU TYR ASP
SEQRES  17 D  315  PRO GLU TRP ALA ARG VAL PHE TYR GLU GLY THR VAL GLY
SEQRES  18 D  315  LEU SER CYS PRO HIS GLU ARG MET LEU GLY GLN VAL LYS
SEQRES  19 D  315  THR PRO VAL LEU LEU THR HIS HIS MET ARG GLY ILE ASP
SEQRES  20 D  315  PRO GLU THR GLY ASN LEU LEU GLY ALA LEU SER ASP GLU
SEQRES  21 D  315  GLN ALA LEU ARG ALA ARG ARG LEU MET ASP SER ALA GLY
SEQRES  22 D  315  VAL THR VAL ASP TYR GLU SER VAL PRO ASP ALA SER TYR
SEQRES  23 D  315  MET MET HIS GLN SER ALA PRO ALA ARG TYR VAL GLU ILE
SEQRES  24 D  315  PHE THR ARG TRP ALA ALA ALA LEU ALA PRO HIS HIS HIS
SEQRES  25 D  315  HIS HIS HIS
FORMUL   5  HOH   *589(H2 O)
HELIX    1 AA1 SER A   50  SER A   53  5                                   4
HELIX    2 AA2 TYR A   54  SER A   62  1                                   9
HELIX    3 AA3 SER A   86  VAL A  101  1                                  16
HELIX    4 AA4 SER A  112  SER A  125  1                                  14
HELIX    5 AA5 SER A  152  GLY A  156  5                                   5
HELIX    6 AA6 ALA A  157  LEU A  168  1                                  12
HELIX    7 AA7 GLY A  169  SER A  173  5                                   5
HELIX    8 AA8 ASP A  176  GLY A  185  1                                  10
HELIX    9 AA9 PRO A  201  GLU A  206  1                                   6
HELIX   10 AB1 PRO A  209  GLU A  217  1                                   9
HELIX   11 AB2 PRO A  225  GLY A  231  1                                   7
HELIX   12 AB3 SER A  258  ALA A  272  1                                  15
HELIX   13 AB4 MET A  287  ALA A  292  1                                   6
HELIX   14 AB5 ALA A  292  ALA A  305  1                                  14
HELIX   15 AB6 SER B   50  SER B   53  5                                   4
HELIX   16 AB7 TYR B   54  SER B   62  1                                   9
HELIX   17 AB8 SER B   86  VAL B  101  1                                  16
HELIX   18 AB9 SER B  112  SER B  125  1                                  14
HELIX   19 AC1 SER B  152  GLY B  156  5                                   5
HELIX   20 AC2 ALA B  157  LEU B  168  1                                  12
HELIX   21 AC3 GLY B  169  SER B  173  5                                   5
HELIX   22 AC4 ASP B  176  ALA B  186  1                                  11
HELIX   23 AC5 SER B  189  SER B  194  1                                   6
HELIX   24 AC6 PRO B  201  GLU B  206  1                                   6
HELIX   25 AC7 PRO B  209  GLU B  217  1                                   9
HELIX   26 AC8 PRO B  225  GLY B  231  1                                   7
HELIX   27 AC9 SER B  258  ALA B  272  1                                  15
HELIX   28 AD1 MET B  287  ALA B  292  1                                   6
HELIX   29 AD2 ALA B  292  ALA B  305  1                                  14
HELIX   30 AD3 SER C   50  SER C   53  5                                   4
HELIX   31 AD4 TYR C   54  SER C   62  1                                   9
HELIX   32 AD5 SER C   86  VAL C  101  1                                  16
HELIX   33 AD6 SER C  112  SER C  125  1                                  14
HELIX   34 AD7 SER C  152  GLY C  156  5                                   5
HELIX   35 AD8 ALA C  157  LEU C  168  1                                  12
HELIX   36 AD9 GLY C  169  SER C  173  5                                   5
HELIX   37 AE1 ASP C  176  ALA C  184  1                                   9
HELIX   38 AE2 PRO C  201  GLU C  206  1                                   6
HELIX   39 AE3 PRO C  209  GLU C  217  1                                   9
HELIX   40 AE4 PRO C  225  GLY C  231  1                                   7
HELIX   41 AE5 SER C  258  GLY C  273  1                                  16
HELIX   42 AE6 MET C  287  ALA C  292  1                                   6
HELIX   43 AE7 ALA C  292  ALA C  306  1                                  15
HELIX   44 AE8 SER D    4  ARG D    8  5                                   5
HELIX   45 AE9 SER D   50  SER D   53  5                                   4
HELIX   46 AF1 TYR D   54  SER D   62  1                                   9
HELIX   47 AF2 SER D   86  VAL D  101  1                                  16
HELIX   48 AF3 SER D  112  SER D  125  1                                  14
HELIX   49 AF4 SER D  152  GLY D  156  5                                   5
HELIX   50 AF5 ALA D  157  LEU D  168  1                                  12
HELIX   51 AF6 GLY D  169  SER D  173  5                                   5
HELIX   52 AF7 ASP D  176  ALA D  184  1                                   9
HELIX   53 AF8 PRO D  201  GLU D  206  1                                   6
HELIX   54 AF9 PRO D  209  GLU D  217  1                                   9
HELIX   55 AG1 PRO D  225  GLY D  231  1                                   7
HELIX   56 AG2 SER D  258  GLY D  273  1                                  16
HELIX   57 AG3 MET D  287  ALA D  292  1                                   6
HELIX   58 AG4 ALA D  292  ALA D  306  1                                  15
SHEET    1 AA1 8 GLU A  18  ASP A  22  0
SHEET    2 AA1 8 THR A  27  ALA A  33 -1  O  TYR A  30   N  GLN A  19
SHEET    3 AA1 8 HIS A  66  VAL A  70 -1  O  ALA A  69   N  ALA A  31
SHEET    4 AA1 8 ALA A  40  ILE A  44  1  N  LEU A  43   O  TYR A  68
SHEET    5 AA1 8 VAL A 106  ASN A 111  1  O  ALA A 109   N  ILE A  44
SHEET    6 AA1 8 LEU A 130  GLU A 136  1  O  LEU A 134   N  VAL A 108
SHEET    7 AA1 8 VAL A 237  HIS A 241  1  O  LEU A 238   N  CYS A 135
SHEET    8 AA1 8 VAL A 276  SER A 280  1  O  ASP A 277   N  VAL A 237
SHEET    1 AA2 2 ARG A 244  ILE A 246  0
SHEET    2 AA2 2 LEU A 253  GLY A 255 -1  O  LEU A 254   N  GLY A 245
SHEET    1 AA3 8 GLU B  18  ASP B  22  0
SHEET    2 AA3 8 THR B  27  ALA B  33 -1  O  TYR B  30   N  GLN B  19
SHEET    3 AA3 8 HIS B  66  VAL B  70 -1  O  ALA B  69   N  ALA B  31
SHEET    4 AA3 8 ALA B  40  ILE B  44  1  N  LEU B  43   O  TYR B  68
SHEET    5 AA3 8 VAL B 106  ASN B 111  1  O  ALA B 109   N  ILE B  44
SHEET    6 AA3 8 LEU B 130  GLU B 136  1  O  LEU B 134   N  VAL B 108
SHEET    7 AA3 8 VAL B 237  HIS B 241  1  O  LEU B 238   N  CYS B 135
SHEET    8 AA3 8 VAL B 276  SER B 280  1  O  ASP B 277   N  LEU B 239
SHEET    1 AA4 2 ARG B 244  ILE B 246  0
SHEET    2 AA4 2 LEU B 253  GLY B 255 -1  O  LEU B 254   N  GLY B 245
SHEET    1 AA5 8 GLU C  18  ASP C  22  0
SHEET    2 AA5 8 THR C  27  ALA C  33 -1  O  TYR C  30   N  GLN C  19
SHEET    3 AA5 8 HIS C  66  VAL C  70 -1  O  ALA C  69   N  ALA C  31
SHEET    4 AA5 8 ALA C  40  ILE C  44  1  N  LEU C  43   O  TYR C  68
SHEET    5 AA5 8 VAL C 106  ASN C 111  1  O  ALA C 109   N  ILE C  44
SHEET    6 AA5 8 LEU C 130  GLU C 136  1  O  GLU C 136   N  GLY C 110
SHEET    7 AA5 8 VAL C 237  HIS C 241  1  O  LEU C 238   N  CYS C 135
SHEET    8 AA5 8 VAL C 276  SER C 280  1  O  ASP C 277   N  LEU C 239
SHEET    1 AA6 2 ARG C 244  ILE C 246  0
SHEET    2 AA6 2 LEU C 253  GLY C 255 -1  O  LEU C 254   N  GLY C 245
SHEET    1 AA7 8 GLU D  18  ASP D  22  0
SHEET    2 AA7 8 THR D  27  ALA D  33 -1  O  TYR D  30   N  GLN D  19
SHEET    3 AA7 8 HIS D  66  VAL D  70 -1  O  VAL D  67   N  ALA D  33
SHEET    4 AA7 8 ALA D  40  ILE D  44  1  N  VAL D  41   O  TYR D  68
SHEET    5 AA7 8 VAL D 106  ASN D 111  1  O  ALA D 109   N  LEU D  42
SHEET    6 AA7 8 LEU D 130  GLU D 136  1  O  LEU D 134   N  VAL D 108
SHEET    7 AA7 8 VAL D 237  HIS D 241  1  O  LEU D 238   N  CYS D 135
SHEET    8 AA7 8 VAL D 276  SER D 280  1  O  ASP D 277   N  LEU D 239
SHEET    1 AA8 2 ARG D 244  ILE D 246  0
SHEET    2 AA8 2 LEU D 253  GLY D 255 -1  O  LEU D 254   N  GLY D 245
CRYST1   63.569  126.128  128.707  90.00  90.00  90.00 P 21 21 21   16
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.015731  0.000000  0.000000        0.00000
SCALE2      0.000000  0.007928  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007770        0.00000
TER    4443      ALA A 305
TER    8902      ALA B 308
TER   13244      LEU C 307
TER   17704      LEU D 307
MASTER      385    0    0   58   40    0    0    6 9651    4    0  100
END