longtext: 8clu-pdb

content
HEADER    HYDROLASE                               17-FEB-23   8CLU
TITLE     ZEARALENONE LACTONASE FROM RHODOCOCCUS ERYTHROPOLIS IN COMPLEX WITH
TITLE    2 ZEARALACTAMENONE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ZEARALENONE LACTONASE;
COMPND   3 CHAIN: A, B;
COMPND   4 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: RHODOCOCCUS ERYTHROPOLIS;
SOURCE   3 ORGANISM_TAXID: 1833;
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   5 EXPRESSION_SYSTEM_TAXID: 469008
KEYWDS    ZEARALENONE, MYCOTOXIN, LACTONASE, CARBOXYLESTERASE, ESTERASE,
KEYWDS   2 HYDROLASE, BIODEGRADATION, RHODOCOCCUS ERYTHROPOLIS,
KEYWDS   3 ZEARALACTAMENONE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    D.PUEHRINGER
REVDAT   1   21-FEB-24 8CLU    0
JRNL        AUTH   S.FRUHAUF,D.PUEHRINGER,M.THAMHESL,P.FAJTL,E.KUNZ-VEKIRO,
JRNL        AUTH 2 A.HOEBARTNER-GUSSL,J.PANHOELZL,K.PREM,V.KLINGENBRUNNER,
JRNL        AUTH 3 G.SCHATZMAYR,G.ADAM,J.DAMBORSKY,K.DJINOVIC-CARUGO,Z.PROKOP,
JRNL        AUTH 4 W.D.MOLL
JRNL        TITL   BACTERIAL LACTONASES ZENA WITH NONCANONICAL STRUCTURAL
JRNL        TITL 2 FEATURES HYDROLYSE THE MYCOTOXIN ZEARALENONE
JRNL        REF    ACS CATALYSIS                              2024
JRNL        REFN                   ESSN 2155-5435
JRNL        DOI    10.1021/ACSCATAL.4C00271
REMARK   2
REMARK   2 RESOLUTION.    1.80 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.20.1-4487
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.80
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 47.59
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.360
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.8
REMARK   3   NUMBER OF REFLECTIONS             : 63294
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.167
REMARK   3   R VALUE            (WORKING SET) : 0.165
REMARK   3   FREE R VALUE                     : 0.198
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.960
REMARK   3   FREE R VALUE TEST SET COUNT      : 3139
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 47.5900 -  5.0400    1.00     2913   165  0.1802 0.1806
REMARK   3     2  5.0400 -  4.0000    1.00     2775   173  0.1230 0.1476
REMARK   3     3  4.0000 -  3.5000    1.00     2816   116  0.1384 0.1701
REMARK   3     4  3.5000 -  3.1800    1.00     2775   129  0.1487 0.1906
REMARK   3     5  3.1800 -  2.9500    1.00     2722   167  0.1595 0.2015
REMARK   3     6  2.9500 -  2.7700    1.00     2708   170  0.1484 0.1967
REMARK   3     7  2.7700 -  2.6400    1.00     2737   172  0.1500 0.1911
REMARK   3     8  2.6400 -  2.5200    1.00     2727   150  0.1561 0.2138
REMARK   3     9  2.5200 -  2.4200    1.00     2709   156  0.1558 0.1887
REMARK   3    10  2.4200 -  2.3400    1.00     2731   146  0.1636 0.1920
REMARK   3    11  2.3400 -  2.2700    1.00     2723   131  0.1612 0.2232
REMARK   3    12  2.2700 -  2.2000    1.00     2753   117  0.1733 0.2208
REMARK   3    13  2.2000 -  2.1400    1.00     2724   133  0.1766 0.2197
REMARK   3    14  2.1400 -  2.0900    1.00     2700   138  0.1815 0.1925
REMARK   3    15  2.0900 -  2.0400    1.00     2724   137  0.1824 0.2130
REMARK   3    16  2.0400 -  2.0000    1.00     2733   143  0.1970 0.2306
REMARK   3    17  2.0000 -  1.9600    1.00     2691   148  0.2017 0.2787
REMARK   3    18  1.9600 -  1.9200    1.00     2726   128  0.2237 0.2854
REMARK   3    19  1.9200 -  1.8900    1.00     2727   126  0.2399 0.3205
REMARK   3    20  1.8900 -  1.8600    1.00     2694   150  0.2554 0.2827
REMARK   3    21  1.8600 -  1.8300    1.00     2738   113  0.2611 0.2890
REMARK   3    22  1.8300 -  1.8000    0.97     2622   131  0.2734 0.3407
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.10
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.210
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 21.410
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 22.25
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.010           5082
REMARK   3   ANGLE     :  1.126           6937
REMARK   3   CHIRALITY :  0.058            727
REMARK   3   PLANARITY :  0.012            917
REMARK   3   DIHEDRAL  :  8.705            699
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 2
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: (CHAIN 'A' AND RESID 24 THROUGH 332)
REMARK   3    ORIGIN FOR THE GROUP (A):  21.6451  15.5540  22.8444
REMARK   3    T TENSOR
REMARK   3      T11:   0.2116 T22:   0.1830
REMARK   3      T33:   0.1970 T12:  -0.0255
REMARK   3      T13:  -0.0111 T23:   0.0064
REMARK   3    L TENSOR
REMARK   3      L11:   0.5560 L22:   1.0581
REMARK   3      L33:   0.9348 L12:  -0.0368
REMARK   3      L13:   0.0069 L23:   0.0815
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0341 S12:   0.0406 S13:   0.0439
REMARK   3      S21:   0.0365 S22:   0.0078 S23:  -0.0333
REMARK   3      S31:  -0.0703 S32:   0.0361 S33:   0.0245
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: (CHAIN 'B' AND RESID 21 THROUGH 326)
REMARK   3    ORIGIN FOR THE GROUP (A):   7.6098 -15.6899  20.2094
REMARK   3    T TENSOR
REMARK   3      T11:   0.2140 T22:   0.1974
REMARK   3      T33:   0.2045 T12:  -0.0308
REMARK   3      T13:   0.0055 T23:   0.0149
REMARK   3    L TENSOR
REMARK   3      L11:   0.5740 L22:   1.0284
REMARK   3      L33:   1.1595 L12:  -0.1394
REMARK   3      L13:   0.0443 L23:  -0.0877
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0268 S12:   0.0384 S13:  -0.0334
REMARK   3      S21:   0.0078 S22:   0.0288 S23:   0.0832
REMARK   3      S31:   0.0876 S32:  -0.1019 S33:  -0.0028
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 8CLU COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 20-FEB-23.
REMARK 100 THE DEPOSITION ID IS D_1292126827.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 12-APR-19
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : DIAMOND
REMARK 200  BEAMLINE                       : I03
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.976250
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS EIGER X 16M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XDS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 63294
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.800
REMARK 200  RESOLUTION RANGE LOW       (A) : 47.590
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.9
REMARK 200  DATA REDUNDANCY                : 6.700
REMARK 200  R MERGE                    (I) : 0.15800
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 8.8700
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.80
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.86
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.4
REMARK 200  DATA REDUNDANCY IN SHELL       : 6.90
REMARK 200  R MERGE FOR SHELL          (I) : 3.11200
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 0.500
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 47.58
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.35
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.2 M NACL, 0.1 M BIS-TRIS 5.5 PH, 25%
REMARK 280  W/V PEG 3350, PH 5.5, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE
REMARK 280  295.15K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 2 2 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z+1/2
REMARK 290       3555   -X,Y,-Z+1/2
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z
REMARK 290       6555   -X+1/2,-Y+1/2,Z+1/2
REMARK 290       7555   -X+1/2,Y+1/2,-Z+1/2
REMARK 290       8555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       86.31250
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       86.31250
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       32.27800
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       60.92000
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       32.27800
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       60.92000
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       86.31250
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       32.27800
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       60.92000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       86.31250
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       32.27800
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000       60.92000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 2580 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 21800 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -5.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     ALA A     2
REMARK 465     GLU A     3
REMARK 465     GLU A     4
REMARK 465     GLY A     5
REMARK 465     THR A     6
REMARK 465     ARG A     7
REMARK 465     SER A     8
REMARK 465     GLU A     9
REMARK 465     ALA A    10
REMARK 465     ALA A    11
REMARK 465     ASP A    12
REMARK 465     ALA A    13
REMARK 465     ALA A    14
REMARK 465     THR A    15
REMARK 465     GLN A    16
REMARK 465     ALA A    17
REMARK 465     ARG A    18
REMARK 465     GLN A    19
REMARK 465     LEU A    20
REMARK 465     PRO A    21
REMARK 465     ASP A    22
REMARK 465     SER A    23
REMARK 465     HIS A   333
REMARK 465     HIS A   334
REMARK 465     MET B     1
REMARK 465     ALA B     2
REMARK 465     GLU B     3
REMARK 465     GLU B     4
REMARK 465     GLY B     5
REMARK 465     THR B     6
REMARK 465     ARG B     7
REMARK 465     SER B     8
REMARK 465     GLU B     9
REMARK 465     ALA B    10
REMARK 465     ALA B    11
REMARK 465     ASP B    12
REMARK 465     ALA B    13
REMARK 465     ALA B    14
REMARK 465     THR B    15
REMARK 465     GLN B    16
REMARK 465     ALA B    17
REMARK 465     ARG B    18
REMARK 465     GLN B    19
REMARK 465     LEU B    20
REMARK 465     ASN B   327
REMARK 465     ASP B   328
REMARK 465     HIS B   329
REMARK 465     HIS B   330
REMARK 465     HIS B   331
REMARK 465     HIS B   332
REMARK 465     HIS B   333
REMARK 465     HIS B   334
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASN A  25       44.83   -150.20
REMARK 500    SER A 128     -124.31     65.43
REMARK 500    PHE A 157      -51.26     69.64
REMARK 500    ALA A 172     -110.69   -141.78
REMARK 500    ALA A 172     -110.69   -137.87
REMARK 500    HIS A 259     -145.06    -92.73
REMARK 500    ASN B  25       34.72   -141.42
REMARK 500    THR B  64       -0.52     74.74
REMARK 500    SER B 128     -126.24     66.35
REMARK 500    PHE B 157      -49.29     70.71
REMARK 500    ALA B 172     -101.50   -142.10
REMARK 500    HIS B 259     -146.58    -95.55
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: PLANAR GROUPS
REMARK 500
REMARK 500 PLANAR GROUPS IN THE FOLLOWING RESIDUES HAVE A TOTAL
REMARK 500 RMS DISTANCE OF ALL ATOMS FROM THE BEST-FIT PLANE
REMARK 500 BY MORE THAN AN EXPECTED VALUE OF 6*RMSD, WITH AN
REMARK 500 RMSD 0.02 ANGSTROMS, OR AT LEAST ONE ATOM HAS
REMARK 500 AN RMSD GREATER THAN THIS VALUE
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500  M RES CSSEQI        RMS     TYPE
REMARK 500    ARG B  99         0.08    SIDE CHAIN
REMARK 500
REMARK 500 REMARK: NULL
DBREF  8CLU A    1   334  PDB    8CLU     8CLU             1    334
DBREF  8CLU B    1   334  PDB    8CLU     8CLU             1    334
SEQRES   1 A  334  MET ALA GLU GLU GLY THR ARG SER GLU ALA ALA ASP ALA
SEQRES   2 A  334  ALA THR GLN ALA ARG GLN LEU PRO ASP SER ARG ASN ILE
SEQRES   3 A  334  PHE VAL SER HIS ARG PHE PRO GLU ARG GLN VAL ASP LEU
SEQRES   4 A  334  GLY GLU VAL VAL MET ASN PHE ALA GLU ALA GLY SER PRO
SEQRES   5 A  334  ASP ASN PRO ALA LEU LEU LEU LEU PRO GLU GLN THR GLY
SEQRES   6 A  334  SER TRP TRP SER TYR GLU PRO VAL MET GLY LEU LEU ALA
SEQRES   7 A  334  GLU ASN PHE HIS VAL PHE ALA VAL ASP ILE ARG GLY GLN
SEQRES   8 A  334  GLY ARG SER THR TRP THR PRO ARG ARG TYR SER LEU ASP
SEQRES   9 A  334  ASN PHE GLY ASN ASP LEU VAL ARG PHE ILE ALA LEU VAL
SEQRES  10 A  334  ILE LYS ARG PRO VAL VAL VAL ALA GLY ASN SER SER GLY
SEQRES  11 A  334  GLY LEU LEU ALA ALA TRP LEU SER ALA TYR ALA MET PRO
SEQRES  12 A  334  GLY GLN ILE ARG ALA ALA LEU CYS GLU ASP ALA PRO PHE
SEQRES  13 A  334  PHE ALA SER GLU LEU VAL PRO ALA TYR GLY HIS SER VAL
SEQRES  14 A  334  LEU GLN ALA ALA GLY PRO ALA PHE GLU LEU TYR ARG ASP
SEQRES  15 A  334  PHE LEU GLY ASP GLN TRP SER ILE GLY ASP TRP LYS GLY
SEQRES  16 A  334  PHE VAL GLU ALA ALA LYS ALA SER PRO ALA LYS ALA MET
SEQRES  17 A  334  GLN LEU PHE PRO THR PRO ASP GLU ALA PRO GLN ASN LEU
SEQRES  18 A  334  LYS GLU TYR ASP PRO GLU TRP GLY ARG ALA PHE PHE GLU
SEQRES  19 A  334  GLY THR VAL ALA LEU HIS CYS PRO HIS ASP ARG MET LEU
SEQRES  20 A  334  SER GLN VAL LYS THR PRO ILE LEU ILE THR HIS HIS ALA
SEQRES  21 A  334  ARG THR ILE ASP PRO GLU THR GLY GLU LEU LEU GLY ALA
SEQRES  22 A  334  LEU SER ASP LEU GLN ALA GLU HIS ALA GLN ASP ILE ILE
SEQRES  23 A  334  ARG SER ALA GLY VAL ARG VAL ASP TYR GLN SER HIS PRO
SEQRES  24 A  334  ASP ALA LEU HIS MET MET HIS LEU PHE ASP PRO ALA ARG
SEQRES  25 A  334  TYR ALA GLU ILE LEU THR SER TRP SER ALA THR LEU PRO
SEQRES  26 A  334  ALA ASN ASP HIS HIS HIS HIS HIS HIS
SEQRES   1 B  334  MET ALA GLU GLU GLY THR ARG SER GLU ALA ALA ASP ALA
SEQRES   2 B  334  ALA THR GLN ALA ARG GLN LEU PRO ASP SER ARG ASN ILE
SEQRES   3 B  334  PHE VAL SER HIS ARG PHE PRO GLU ARG GLN VAL ASP LEU
SEQRES   4 B  334  GLY GLU VAL VAL MET ASN PHE ALA GLU ALA GLY SER PRO
SEQRES   5 B  334  ASP ASN PRO ALA LEU LEU LEU LEU PRO GLU GLN THR GLY
SEQRES   6 B  334  SER TRP TRP SER TYR GLU PRO VAL MET GLY LEU LEU ALA
SEQRES   7 B  334  GLU ASN PHE HIS VAL PHE ALA VAL ASP ILE ARG GLY GLN
SEQRES   8 B  334  GLY ARG SER THR TRP THR PRO ARG ARG TYR SER LEU ASP
SEQRES   9 B  334  ASN PHE GLY ASN ASP LEU VAL ARG PHE ILE ALA LEU VAL
SEQRES  10 B  334  ILE LYS ARG PRO VAL VAL VAL ALA GLY ASN SER SER GLY
SEQRES  11 B  334  GLY LEU LEU ALA ALA TRP LEU SER ALA TYR ALA MET PRO
SEQRES  12 B  334  GLY GLN ILE ARG ALA ALA LEU CYS GLU ASP ALA PRO PHE
SEQRES  13 B  334  PHE ALA SER GLU LEU VAL PRO ALA TYR GLY HIS SER VAL
SEQRES  14 B  334  LEU GLN ALA ALA GLY PRO ALA PHE GLU LEU TYR ARG ASP
SEQRES  15 B  334  PHE LEU GLY ASP GLN TRP SER ILE GLY ASP TRP LYS GLY
SEQRES  16 B  334  PHE VAL GLU ALA ALA LYS ALA SER PRO ALA LYS ALA MET
SEQRES  17 B  334  GLN LEU PHE PRO THR PRO ASP GLU ALA PRO GLN ASN LEU
SEQRES  18 B  334  LYS GLU TYR ASP PRO GLU TRP GLY ARG ALA PHE PHE GLU
SEQRES  19 B  334  GLY THR VAL ALA LEU HIS CYS PRO HIS ASP ARG MET LEU
SEQRES  20 B  334  SER GLN VAL LYS THR PRO ILE LEU ILE THR HIS HIS ALA
SEQRES  21 B  334  ARG THR ILE ASP PRO GLU THR GLY GLU LEU LEU GLY ALA
SEQRES  22 B  334  LEU SER ASP LEU GLN ALA GLU HIS ALA GLN ASP ILE ILE
SEQRES  23 B  334  ARG SER ALA GLY VAL ARG VAL ASP TYR GLN SER HIS PRO
SEQRES  24 B  334  ASP ALA LEU HIS MET MET HIS LEU PHE ASP PRO ALA ARG
SEQRES  25 B  334  TYR ALA GLU ILE LEU THR SER TRP SER ALA THR LEU PRO
SEQRES  26 B  334  ALA ASN ASP HIS HIS HIS HIS HIS HIS
HET    V0F  A 600      46
HET    GOL  A 601      14
HET    V0F  B 600      46
HETNAM     V0F (4~{S})-4-METHYL-16,18-BIS(OXIDANYL)-3-
HETNAM   2 V0F  AZABICYCLO[12.4.0]OCTADECA-1(18),12,14,16-TETRAENE-2,
HETNAM   3 V0F  8-DIONE
HETNAM     GOL GLYCEROL
HETSYN     GOL GLYCERIN; PROPANE-1,2,3-TRIOL
FORMUL   3  V0F    2(C18 H23 N O4)
FORMUL   4  GOL    C3 H8 O3
FORMUL   6  HOH   *379(H2 O)
HELIX    1 AA1 SER A   66  SER A   69  5                                   4
HELIX    2 AA2 TYR A   70  GLU A   79  1                                  10
HELIX    3 AA3 SER A  102  VAL A  117  1                                  16
HELIX    4 AA4 SER A  128  ALA A  141  1                                  14
HELIX    5 AA5 SER A  168  ALA A  172  5                                   5
HELIX    6 AA6 ALA A  173  LEU A  184  1                                  12
HELIX    7 AA7 GLY A  185  SER A  189  5                                   5
HELIX    8 AA8 ASP A  192  ALA A  202  1                                  11
HELIX    9 AA9 ALA A  205  PHE A  211  5                                   7
HELIX   10 AB1 PRO A  218  GLU A  223  1                                   6
HELIX   11 AB2 PRO A  226  GLU A  234  1                                   9
HELIX   12 AB3 PRO A  242  GLN A  249  1                                   8
HELIX   13 AB4 SER A  275  ALA A  289  1                                  15
HELIX   14 AB5 MET A  304  ASP A  309  1                                   6
HELIX   15 AB6 ASP A  309  ALA A  322  1                                  14
HELIX   16 AB7 SER B   66  SER B   69  5                                   4
HELIX   17 AB8 TYR B   70  GLU B   79  1                                  10
HELIX   18 AB9 SER B  102  VAL B  117  1                                  16
HELIX   19 AC1 SER B  128  ALA B  141  1                                  14
HELIX   20 AC2 SER B  168  ALA B  172  5                                   5
HELIX   21 AC3 ALA B  173  LEU B  184  1                                  12
HELIX   22 AC4 GLY B  185  SER B  189  5                                   5
HELIX   23 AC5 ASP B  192  SER B  203  1                                  12
HELIX   24 AC6 ALA B  205  PHE B  211  5                                   7
HELIX   25 AC7 PRO B  218  GLU B  223  1                                   6
HELIX   26 AC8 PRO B  226  GLU B  234  1                                   9
HELIX   27 AC9 PRO B  242  GLN B  249  1                                   8
HELIX   28 AD1 SER B  275  ALA B  289  1                                  15
HELIX   29 AD2 MET B  304  ASP B  309  1                                   6
HELIX   30 AD3 ASP B  309  ALA B  322  1                                  14
SHEET    1 AA1 8 GLU A  34  ASP A  38  0
SHEET    2 AA1 8 VAL A  43  ALA A  49 -1  O  MET A  44   N  VAL A  37
SHEET    3 AA1 8 HIS A  82  VAL A  86 -1  O  VAL A  83   N  ALA A  49
SHEET    4 AA1 8 ALA A  56  LEU A  60  1  N  LEU A  57   O  PHE A  84
SHEET    5 AA1 8 VAL A 122  ASN A 127  1  O  ALA A 125   N  LEU A  60
SHEET    6 AA1 8 ILE A 146  GLU A 152  1  O  GLU A 152   N  GLY A 126
SHEET    7 AA1 8 ILE A 254  HIS A 258  1  O  LEU A 255   N  CYS A 151
SHEET    8 AA1 8 VAL A 293  SER A 297  1  O  ASP A 294   N  ILE A 256
SHEET    1 AA2 2 ARG A 261  ILE A 263  0
SHEET    2 AA2 2 LEU A 270  GLY A 272 -1  O  LEU A 271   N  THR A 262
SHEET    1 AA3 8 GLU B  34  ASP B  38  0
SHEET    2 AA3 8 VAL B  43  ALA B  49 -1  O  MET B  44   N  VAL B  37
SHEET    3 AA3 8 HIS B  82  VAL B  86 -1  O  ALA B  85   N  ALA B  47
SHEET    4 AA3 8 ALA B  56  LEU B  60  1  N  LEU B  59   O  PHE B  84
SHEET    5 AA3 8 VAL B 122  ASN B 127  1  O  VAL B 123   N  LEU B  58
SHEET    6 AA3 8 ILE B 146  GLU B 152  1  O  GLU B 152   N  GLY B 126
SHEET    7 AA3 8 ILE B 254  HIS B 258  1  O  LEU B 255   N  CYS B 151
SHEET    8 AA3 8 VAL B 293  SER B 297  1  O  ASP B 294   N  ILE B 256
SHEET    1 AA4 2 ARG B 261  ILE B 263  0
SHEET    2 AA4 2 LEU B 270  GLY B 272 -1  O  LEU B 271   N  THR B 262
CRYST1   64.556  121.840  172.625  90.00  90.00  90.00 C 2 2 21     16
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.015490  0.000000  0.000000        0.00000
SCALE2      0.000000  0.008207  0.000000        0.00000
SCALE3      0.000000  0.000000  0.005793        0.00000
TER    4833      HIS A 332
TER    9567      ALA B 326
MASTER      365    0    3   30   20    0    0    6 5252    2  106   52
END