longtext: 8f6u-pdb

content
HEADER    IMMUNE SYSTEM                           17-NOV-22   8F6U
TITLE     CRYSTAL STRUCTURE OF NANOBODY VHH113 BOUND TO ITS ANTIGEN PA14 CIF
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: CFTR INHIBITORY FACTOR;
COMPND   3 CHAIN: D, C;
COMPND   4 ENGINEERED: YES;
COMPND   5 MOL_ID: 2;
COMPND   6 MOLECULE: NANOBODY VHH113;
COMPND   7 CHAIN: B, A;
COMPND   8 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PSEUDOMONAS AERUGINOSA PA14;
SOURCE   3 ORGANISM_TAXID: 652611;
SOURCE   4 GENE: PA2394;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: TOP10;
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: PDPM73;
SOURCE  10 MOL_ID: 2;
SOURCE  11 ORGANISM_SCIENTIFIC: VICUGNA PACOS;
SOURCE  12 ORGANISM_TAXID: 30538;
SOURCE  13 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21;
SOURCE  14 EXPRESSION_SYSTEM_TAXID: 511693;
SOURCE  15 EXPRESSION_SYSTEM_VARIANT: DE3 RIL;
SOURCE  16 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  17 EXPRESSION_SYSTEM_PLASMID: PCOMB3X
KEYWDS    PSEUDOMONAS AERUGINOSA, NANOBODY VHH, IMMUNOGLOBULIN DOMAIN, CFTR
KEYWDS   2 INHIBITORY FACTOR (CIF), IMMUNE SYSTEM
EXPDTA    X-RAY DIFFRACTION
AUTHOR    A.R.SIMARD,N.M.TAHER,A.K.MISHRA,D.R.MADDEN
REVDAT   1   13-MAR-24 8F6U    0
JRNL        AUTH   A.R.SIMARD,D.R.MADDEN
JRNL        TITL   CRYSTAL STRUCTURE OF NANOBODY VHH113 BOUND TO ITS ANTIGEN
JRNL        TITL 2 PA14 CIF
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   1
REMARK   1 REFERENCE 1
REMARK   1  AUTH   N.VASYLIEVA,S.KITAMURA,J.DONG,B.BARNYCH,K.L.HVORECNY,
REMARK   1  AUTH 2 D.R.MADDEN,S.J.GEE,D.W.WOLAN,C.MORISSEAU,B.D.HAMMOCK
REMARK   1  TITL   NANOBODY-BASED BINDING ASSAY FOR THE DISCOVERY OF POTENT
REMARK   1  TITL 2 INHIBITORS OF CFTR INHIBITORY FACTOR (CIF).
REMARK   1  REF    ANAL CHIM ACTA                V.1057   106 2019
REMARK   1  REFN                   ISSN 1873-4324
REMARK   1  PMID   30832908
REMARK   1  DOI    10.1016/J.ACA.2018.12.060
REMARK   1 REFERENCE 2
REMARK   1  AUTH   P.V.AFONINE,R.W.GROSSE-KUNSTLEVE,N.ECHOLS,J.J.HEADD,
REMARK   1  AUTH 2 N.W.MORIARTY,M.MUSTYAKIMOV,T.C.TERWILLIGER,A.URZHUMTSEV,
REMARK   1  AUTH 3 P.H.ZWART,P.D.ADAMS
REMARK   1  TITL   TOWARDS AUTOMATED CRYSTALLOGRAPHIC STRUCTURE REFINEMENT WITH
REMARK   1  TITL 2 PHENIX.REFINE.
REMARK   1  REF    ACTA CRYSTALLOGR D BIOL       V.  68   352 2012
REMARK   1  REF  2 CRYSTALLOGR
REMARK   1  REFN                   ESSN 1399-0047
REMARK   1  PMID   22505256
REMARK   1  DOI    10.1107/S0907444912001308
REMARK   2
REMARK   2 RESOLUTION.    1.70 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.20.1_4487
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : GEOSTD + MONOMER LIBRARY + CDL V1.2
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.70
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 46.49
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.330
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.1
REMARK   3   NUMBER OF REFLECTIONS             : 124297
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.177
REMARK   3   R VALUE            (WORKING SET) : 0.176
REMARK   3   FREE R VALUE                     : 0.199
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.010
REMARK   3   FREE R VALUE TEST SET COUNT      : 6228
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 46.4900 -  5.2800    0.99     4088   317  0.1774 0.1811
REMARK   3     2  5.2800 -  4.1900    0.99     4228    25  0.1344 0.1852
REMARK   3     3  4.1900 -  3.6600    0.99     3927   296  0.1471 0.1612
REMARK   3     4  3.6600 -  3.3300    1.00     3920   288  0.1560 0.1683
REMARK   3     5  3.3300 -  3.0900    0.99     4117    97  0.1666 0.1689
REMARK   3     6  3.0900 -  2.9100    0.99     3912   261  0.1836 0.1996
REMARK   3     7  2.9100 -  2.7600    0.99     3921   217  0.1818 0.2122
REMARK   3     8  2.7600 -  2.6400    0.99     3986   180  0.1774 0.2014
REMARK   3     9  2.6400 -  2.5400    0.99     3935   209  0.1771 0.1961
REMARK   3    10  2.5400 -  2.4500    0.99     3929   219  0.1812 0.2333
REMARK   3    11  2.4500 -  2.3800    0.99     3929   208  0.1787 0.1885
REMARK   3    12  2.3800 -  2.3100    0.99     3948   205  0.1708 0.2094
REMARK   3    13  2.3100 -  2.2500    0.99     3917   204  0.1712 0.1935
REMARK   3    14  2.2500 -  2.1900    0.99     3932   211  0.1722 0.1799
REMARK   3    15  2.1900 -  2.1400    0.99     3884   224  0.1760 0.1934
REMARK   3    16  2.1400 -  2.1000    0.99     3944   180  0.1810 0.2356
REMARK   3    17  2.1000 -  2.0500    0.99     3915   222  0.1864 0.2098
REMARK   3    18  2.0500 -  2.0200    0.99     3901   210  0.1919 0.2104
REMARK   3    19  2.0200 -  1.9800    0.99     3930   189  0.1945 0.2051
REMARK   3    20  1.9800 -  1.9500    0.99     3871   238  0.1952 0.2359
REMARK   3    21  1.9500 -  1.9100    0.99     3919   166  0.2124 0.2313
REMARK   3    22  1.9100 -  1.8900    0.99     3911   209  0.2269 0.2641
REMARK   3    23  1.8900 -  1.8600    0.99     3891   220  0.2460 0.2971
REMARK   3    24  1.8600 -  1.8300    0.99     3895   203  0.2573 0.2710
REMARK   3    25  1.8300 -  1.8100    0.99     3894   210  0.2580 0.2985
REMARK   3    26  1.8100 -  1.7800    0.99     3912   209  0.2548 0.2871
REMARK   3    27  1.7800 -  1.7600    0.98     3845   218  0.2648 0.3057
REMARK   3    28  1.7600 -  1.7400    0.99     3891   192  0.2844 0.2712
REMARK   3    29  1.7400 -  1.7200    0.99     3897   220  0.3018 0.3763
REMARK   3    30  1.7200 -  1.7000    0.98     3880   181  0.3142 0.3408
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.10
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.192
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 20.409
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 26.22
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 29.61
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.006           6812
REMARK   3   ANGLE     :  0.839           9259
REMARK   3   CHIRALITY :  0.055            965
REMARK   3   PLANARITY :  0.008           1221
REMARK   3   DIHEDRAL  : 12.868           2439
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: ATOMS MODELED WITH ZERO OCCUPANCY COULD
REMARK   3  NOT BE PLACED WITH CONFIDENCE AND WERE SELECTED FOR ZERO-
REMARK   3  OCCUPANCY FLAGGING AFTER MANUAL INSPECTION OF THE 2FO-FC MAP AT
REMARK   3  A 0.5-SIGMA CUTOFF.
REMARK   4
REMARK   4 8F6U COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 18-NOV-22.
REMARK 100 THE DEPOSITION ID IS D_1000270064.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 14-AUG-18
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : NSLS-II
REMARK 200  BEAMLINE                       : 17-ID-2
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.979339
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS EIGER X 16M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 124313
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.700
REMARK 200  RESOLUTION RANGE LOW       (A) : 46.490
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.1
REMARK 200  DATA REDUNDANCY                : 6.700
REMARK 200  R MERGE                    (I) : 0.10670
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 11.8000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.70
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.76
REMARK 200  COMPLETENESS FOR SHELL     (%) : 98.7
REMARK 200  DATA REDUNDANCY IN SHELL       : 6.60
REMARK 200  R MERGE FOR SHELL          (I) : 1.64600
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 1.260
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: 3KD2, 8E1C
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 56.63
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.83
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 200 MM POTASSIUM BROMIDE, 200 MM
REMARK 280  POTASSIUM THIOCYANATE, 100 MM SODIUM ACETATE PH 5, 3% (W/V)
REMARK 280  GAMMA-PGA (NA+ FORM, LM), 10% (W/V) PEG 2000 MME, PH 5.0, VAPOR
REMARK 280  DIFFUSION, SITTING DROP, TEMPERATURE 292.8K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 2 2 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z+1/2
REMARK 290       3555   -X,Y,-Z+1/2
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z
REMARK 290       6555   -X+1/2,-Y+1/2,Z+1/2
REMARK 290       7555   -X+1/2,Y+1/2,-Z+1/2
REMARK 290       8555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       53.37650
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       53.37650
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       57.59250
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       92.97050
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       57.59250
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       92.97050
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       53.37650
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       57.59250
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       92.97050
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       53.37650
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       57.59250
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000       92.97050
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: TETRAMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: TETRAMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 7020 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 29830 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -38.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: D, B, C, A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH C 402  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A 271  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A 329  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A 331  LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLY D   318
REMARK 465     ARG D   319
REMARK 465     HIS D   320
REMARK 465     HIS D   321
REMARK 465     HIS D   322
REMARK 465     HIS D   323
REMARK 465     HIS D   324
REMARK 465     HIS D   325
REMARK 465     MET B     1
REMARK 465     ALA B     2
REMARK 465     GLU B     3
REMARK 465     VAL B     4
REMARK 465     GLN B     5
REMARK 465     GLY B   130
REMARK 465     GLN B   131
REMARK 465     ALA B   132
REMARK 465     GLY B   133
REMARK 465     GLN B   134
REMARK 465     HIS B   135
REMARK 465     HIS B   136
REMARK 465     HIS B   137
REMARK 465     HIS B   138
REMARK 465     HIS B   139
REMARK 465     HIS B   140
REMARK 465     GLY B   141
REMARK 465     ALA B   142
REMARK 465     TYR B   143
REMARK 465     PRO B   144
REMARK 465     TYR B   145
REMARK 465     ASP B   146
REMARK 465     VAL B   147
REMARK 465     PRO B   148
REMARK 465     ASP B   149
REMARK 465     TYR B   150
REMARK 465     ALA B   151
REMARK 465     SER B   152
REMARK 465     GLY B   153
REMARK 465     SER B   154
REMARK 465     ALA C    25
REMARK 465     ARG C   319
REMARK 465     HIS C   320
REMARK 465     HIS C   321
REMARK 465     HIS C   322
REMARK 465     HIS C   323
REMARK 465     HIS C   324
REMARK 465     HIS C   325
REMARK 465     MET A     1
REMARK 465     ALA A     2
REMARK 465     GLU A     3
REMARK 465     GLY A   130
REMARK 465     GLN A   131
REMARK 465     ALA A   132
REMARK 465     GLY A   133
REMARK 465     GLN A   134
REMARK 465     HIS A   135
REMARK 465     HIS A   136
REMARK 465     HIS A   137
REMARK 465     HIS A   138
REMARK 465     HIS A   139
REMARK 465     HIS A   140
REMARK 465     GLY A   141
REMARK 465     ALA A   142
REMARK 465     TYR A   143
REMARK 465     PRO A   144
REMARK 465     TYR A   145
REMARK 465     ASP A   146
REMARK 465     VAL A   147
REMARK 465     PRO A   148
REMARK 465     ASP A   149
REMARK 465     TYR A   150
REMARK 465     ALA A   151
REMARK 465     SER A   152
REMARK 465     GLY A   153
REMARK 465     SER A   154
REMARK 480
REMARK 480 ZERO OCCUPANCY ATOM
REMARK 480 THE FOLLOWING RESIDUES HAVE ATOMS MODELED WITH ZERO
REMARK 480 OCCUPANCY. THE LOCATION AND PROPERTIES OF THESE ATOMS
REMARK 480 MAY NOT BE RELIABLE. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 480 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 480   M RES C SSEQI ATOMS
REMARK 480     GLN D   53   CD   OE1  NE2
REMARK 480     GLU D  212   CG   CD   OE1  OE2
REMARK 480     LYS C   45   CE   NZ
REMARK 480     ARG C  121   NE   CZ   NH1  NH2
REMARK 480     GLU C  212   CG   CD   OE1  OE2
REMARK 480     GLU C  285   CD   OE1  OE2
REMARK 480     ARG C  309   NE   CZ   NH1  NH2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASP D 129     -131.82     57.07
REMARK 500    TRP D 298       68.73   -100.69
REMARK 500    ASP C 129     -131.86     58.00
REMARK 500    ALA C 154      147.97   -176.91
REMARK 500    ALA A  97      165.80    175.75
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 338        DISTANCE =  5.86 ANGSTROMS
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3KD2   RELATED DB: PDB
REMARK 900 3KD2 CONTAINS CIF
REMARK 900 RELATED ID: 8E2N   RELATED DB: PDB
REMARK 900 8E2N CONTAINS NANOBODY VHH113
REMARK 900 RELATED ID: 8ELN   RELATED DB: PDB
REMARK 900 8ELN CONTAINS CIF CO-CRYSTALIZED WITH RELATED NANOBODY VHH222
REMARK 900 RELATED ID: 8EVD   RELATED DB: PDB
REMARK 900 8EVD CONTAINS CIF CO-CRYSTALIZED WITH RELATED NANOBODY VHH101
REMARK 900 RELATED ID: 8EE2   RELATED DB: PDB
REMARK 900 8EE2 ONTAINS CIF CO-CRYSTALIZED WITH RELATED NANOBODY VHH219
DBREF1 8F6U D   25   325  UNP                  A0A0M3KL26_PSEAB
DBREF2 8F6U D     A0A0M3KL26                          1         301
DBREF  8F6U B    1   154  PDB    8F6U     8F6U             1    154
DBREF1 8F6U C   25   325  UNP                  A0A0M3KL26_PSEAB
DBREF2 8F6U C     A0A0M3KL26                          1         301
DBREF  8F6U A    1   154  PDB    8F6U     8F6U             1    154
SEQRES   1 D  301  ALA GLU GLU PHE PRO VAL PRO ASN GLY PHE GLU SER ALA
SEQRES   2 D  301  TYR ARG GLU VAL ASP GLY VAL LYS LEU HIS TYR VAL LYS
SEQRES   3 D  301  GLY GLY GLN GLY PRO LEU VAL MET LEU VAL HIS GLY PHE
SEQRES   4 D  301  GLY GLN THR TRP TYR GLU TRP HIS GLN LEU MET PRO GLU
SEQRES   5 D  301  LEU ALA LYS ARG PHE THR VAL ILE ALA PRO ASP LEU PRO
SEQRES   6 D  301  GLY LEU GLY GLN SER GLU PRO PRO LYS THR GLY TYR SER
SEQRES   7 D  301  GLY GLU GLN VAL ALA VAL TYR LEU HIS LYS LEU ALA ARG
SEQRES   8 D  301  GLN PHE SER PRO ASP ARG PRO PHE ASP LEU VAL ALA HIS
SEQRES   9 D  301  ASP ILE GLY ILE TRP ASN THR TYR PRO MET VAL VAL LYS
SEQRES  10 D  301  ASN GLN ALA ASP ILE ALA ARG LEU VAL TYR MET GLU ALA
SEQRES  11 D  301  PRO ILE PRO ASP ALA ARG ILE TYR ARG PHE PRO ALA PHE
SEQRES  12 D  301  THR ALA GLN GLY GLU SER LEU VAL TRP HIS PHE SER PHE
SEQRES  13 D  301  PHE ALA ALA ASP ASP ARG LEU ALA GLU THR LEU ILE ALA
SEQRES  14 D  301  GLY LYS GLU ARG PHE PHE LEU GLU HIS PHE ILE LYS SER
SEQRES  15 D  301  HIS ALA SER ASN THR GLU VAL PHE SER GLU ARG LEU LEU
SEQRES  16 D  301  ASP LEU TYR ALA ARG SER TYR ALA LYS PRO HIS SER LEU
SEQRES  17 D  301  ASN ALA SER PHE GLU TYR TYR ARG ALA LEU ASN GLU SER
SEQRES  18 D  301  VAL ARG GLN ASN ALA GLU LEU ALA LYS THR ARG LEU GLN
SEQRES  19 D  301  MET PRO THR MET THR LEU ALA GLY GLY GLY HIS GLY GLY
SEQRES  20 D  301  MET GLY THR PHE GLN LEU GLU GLN MET LYS ALA TYR ALA
SEQRES  21 D  301  GLU ASP VAL GLU GLY HIS VAL LEU PRO GLY CYS GLY HIS
SEQRES  22 D  301  TRP LEU PRO GLU GLU CYS ALA ALA PRO MET ASN ARG LEU
SEQRES  23 D  301  VAL ILE ASP PHE LEU SER ARG GLY ARG HIS HIS HIS HIS
SEQRES  24 D  301  HIS HIS
SEQRES   1 B  154  MET ALA GLU VAL GLN LEU VAL GLU SER GLY GLY GLY LEU
SEQRES   2 B  154  VAL PRO ALA GLY GLY SER LEU ARG LEU SER CYS THR THR
SEQRES   3 B  154  SER GLU ARG ALA PHE ARG SER ASN ALA MET GLY TRP PHE
SEQRES   4 B  154  ARG GLN ALA PRO GLY LYS GLU ARG GLU PHE VAL ALA ALA
SEQRES   5 B  154  VAL SER VAL LEU SER TRP SER GLY ASP SER ALA VAL VAL
SEQRES   6 B  154  ALA ASP SER VAL ALA GLY ARG PHE THR ILE PHE ARG ASP
SEQRES   7 B  154  ASN ALA LYS ASN THR VAL TYR LEU GLN MET ASN SER LEU
SEQRES   8 B  154  LYS PRO GLU ASP THR ALA VAL TYR TYR CYS ASN GLY ALA
SEQRES   9 B  154  SER ASP ILE GLY ALA LEU GLN SER GLY ALA SER SER TRP
SEQRES  10 B  154  SER TRP GLY HIS GLY THR GLN VAL THR VAL SER SER GLY
SEQRES  11 B  154  GLN ALA GLY GLN HIS HIS HIS HIS HIS HIS GLY ALA TYR
SEQRES  12 B  154  PRO TYR ASP VAL PRO ASP TYR ALA SER GLY SER
SEQRES   1 C  301  ALA GLU GLU PHE PRO VAL PRO ASN GLY PHE GLU SER ALA
SEQRES   2 C  301  TYR ARG GLU VAL ASP GLY VAL LYS LEU HIS TYR VAL LYS
SEQRES   3 C  301  GLY GLY GLN GLY PRO LEU VAL MET LEU VAL HIS GLY PHE
SEQRES   4 C  301  GLY GLN THR TRP TYR GLU TRP HIS GLN LEU MET PRO GLU
SEQRES   5 C  301  LEU ALA LYS ARG PHE THR VAL ILE ALA PRO ASP LEU PRO
SEQRES   6 C  301  GLY LEU GLY GLN SER GLU PRO PRO LYS THR GLY TYR SER
SEQRES   7 C  301  GLY GLU GLN VAL ALA VAL TYR LEU HIS LYS LEU ALA ARG
SEQRES   8 C  301  GLN PHE SER PRO ASP ARG PRO PHE ASP LEU VAL ALA HIS
SEQRES   9 C  301  ASP ILE GLY ILE TRP ASN THR TYR PRO MET VAL VAL LYS
SEQRES  10 C  301  ASN GLN ALA ASP ILE ALA ARG LEU VAL TYR MET GLU ALA
SEQRES  11 C  301  PRO ILE PRO ASP ALA ARG ILE TYR ARG PHE PRO ALA PHE
SEQRES  12 C  301  THR ALA GLN GLY GLU SER LEU VAL TRP HIS PHE SER PHE
SEQRES  13 C  301  PHE ALA ALA ASP ASP ARG LEU ALA GLU THR LEU ILE ALA
SEQRES  14 C  301  GLY LYS GLU ARG PHE PHE LEU GLU HIS PHE ILE LYS SER
SEQRES  15 C  301  HIS ALA SER ASN THR GLU VAL PHE SER GLU ARG LEU LEU
SEQRES  16 C  301  ASP LEU TYR ALA ARG SER TYR ALA LYS PRO HIS SER LEU
SEQRES  17 C  301  ASN ALA SER PHE GLU TYR TYR ARG ALA LEU ASN GLU SER
SEQRES  18 C  301  VAL ARG GLN ASN ALA GLU LEU ALA LYS THR ARG LEU GLN
SEQRES  19 C  301  MET PRO THR MET THR LEU ALA GLY GLY GLY HIS GLY GLY
SEQRES  20 C  301  MET GLY THR PHE GLN LEU GLU GLN MET LYS ALA TYR ALA
SEQRES  21 C  301  GLU ASP VAL GLU GLY HIS VAL LEU PRO GLY CYS GLY HIS
SEQRES  22 C  301  TRP LEU PRO GLU GLU CYS ALA ALA PRO MET ASN ARG LEU
SEQRES  23 C  301  VAL ILE ASP PHE LEU SER ARG GLY ARG HIS HIS HIS HIS
SEQRES  24 C  301  HIS HIS
SEQRES   1 A  154  MET ALA GLU VAL GLN LEU VAL GLU SER GLY GLY GLY LEU
SEQRES   2 A  154  VAL PRO ALA GLY GLY SER LEU ARG LEU SER CYS THR THR
SEQRES   3 A  154  SER GLU ARG ALA PHE ARG SER ASN ALA MET GLY TRP PHE
SEQRES   4 A  154  ARG GLN ALA PRO GLY LYS GLU ARG GLU PHE VAL ALA ALA
SEQRES   5 A  154  VAL SER VAL LEU SER TRP SER GLY ASP SER ALA VAL VAL
SEQRES   6 A  154  ALA ASP SER VAL ALA GLY ARG PHE THR ILE PHE ARG ASP
SEQRES   7 A  154  ASN ALA LYS ASN THR VAL TYR LEU GLN MET ASN SER LEU
SEQRES   8 A  154  LYS PRO GLU ASP THR ALA VAL TYR TYR CYS ASN GLY ALA
SEQRES   9 A  154  SER ASP ILE GLY ALA LEU GLN SER GLY ALA SER SER TRP
SEQRES  10 A  154  SER TRP GLY HIS GLY THR GLN VAL THR VAL SER SER GLY
SEQRES  11 A  154  GLN ALA GLY GLN HIS HIS HIS HIS HIS HIS GLY ALA TYR
SEQRES  12 A  154  PRO TYR ASP VAL PRO ASP TYR ALA SER GLY SER
FORMUL   5  HOH   *784(H2 O)
HELIX    1 AA1 THR D   66  HIS D   71  5                                   6
HELIX    2 AA2 LEU D   73  ALA D   78  1                                   6
HELIX    3 AA3 SER D  102  SER D  118  1                                  17
HELIX    4 AA4 ASP D  129  ASN D  134  1                                   6
HELIX    5 AA5 THR D  135  ASN D  142  1                                   8
HELIX    6 AA6 ASP D  158  PHE D  164  5                                   7
HELIX    7 AA7 TRP D  176  ALA D  183  1                                   8
HELIX    8 AA8 ARG D  186  ALA D  193  1                                   8
HELIX    9 AA9 LYS D  195  HIS D  207  1                                  13
HELIX   10 AB1 ASN D  210  PHE D  214  5                                   5
HELIX   11 AB2 SER D  215  ALA D  227  1                                  13
HELIX   12 AB3 LYS D  228  ALA D  241  1                                  14
HELIX   13 AB4 ALA D  241  ALA D  253  1                                  13
HELIX   14 AB5 THR D  274  ALA D  282  1                                   9
HELIX   15 AB6 TRP D  298  CYS D  303  1                                   6
HELIX   16 AB7 CYS D  303  ARG D  317  1                                  15
HELIX   17 AB8 SER B   27  SER B   33  1                                   7
HELIX   18 AB9 LYS B   92  THR B   96  5                                   5
HELIX   19 AC1 ASP B  106  GLY B  113  1                                   8
HELIX   20 AC2 THR C   66  HIS C   71  5                                   6
HELIX   21 AC3 LEU C   73  ALA C   78  1                                   6
HELIX   22 AC4 SER C  102  SER C  118  1                                  17
HELIX   23 AC5 ASP C  129  ASN C  134  1                                   6
HELIX   24 AC6 THR C  135  ASN C  142  1                                   8
HELIX   25 AC7 ASP C  158  PHE C  164  5                                   7
HELIX   26 AC8 TRP C  176  ALA C  183  1                                   8
HELIX   27 AC9 ARG C  186  ALA C  193  1                                   8
HELIX   28 AD1 LYS C  195  HIS C  207  1                                  13
HELIX   29 AD2 ASN C  210  PHE C  214  5                                   5
HELIX   30 AD3 SER C  215  ALA C  227  1                                  13
HELIX   31 AD4 LYS C  228  ALA C  241  1                                  14
HELIX   32 AD5 ALA C  241  ALA C  253  1                                  13
HELIX   33 AD6 THR C  274  LYS C  281  1                                   8
HELIX   34 AD7 TRP C  298  CYS C  303  1                                   6
HELIX   35 AD8 CYS C  303  SER C  316  1                                  14
HELIX   36 AD9 SER A   27  SER A   33  1                                   7
HELIX   37 AE1 ASN A   79  LYS A   81  5                                   3
HELIX   38 AE2 LYS A   92  THR A   96  5                                   5
HELIX   39 AE3 ASP A  106  GLY A  113  1                                   8
SHEET    1 AA1 8 GLU D  35  VAL D  41  0
SHEET    2 AA1 8 VAL D  44  GLY D  52 -1  O  LYS D  50   N  GLU D  35
SHEET    3 AA1 8 THR D  82  PRO D  86 -1  O  VAL D  83   N  GLY D  51
SHEET    4 AA1 8 LEU D  56  VAL D  60  1  N  VAL D  57   O  THR D  82
SHEET    5 AA1 8 PHE D 123  HIS D 128  1  O  ASP D 124   N  LEU D  56
SHEET    6 AA1 8 ILE D 146  MET D 152  1  O  VAL D 150   N  LEU D 125
SHEET    7 AA1 8 THR D 261  GLY D 266  1  O  MET D 262   N  LEU D 149
SHEET    8 AA1 8 VAL D 287  LEU D 292  1  O  LEU D 292   N  ALA D 265
SHEET    1 AA2 2 PHE D 167  THR D 168  0
SHEET    2 AA2 2 GLY D 171  GLU D 172 -1  O  GLY D 171   N  THR D 168
SHEET    1 AA3 4 VAL B   7  SER B   9  0
SHEET    2 AA3 4 LEU B  20  THR B  25 -1  O  THR B  25   N  VAL B   7
SHEET    3 AA3 4 THR B  83  MET B  88 -1  O  MET B  88   N  LEU B  20
SHEET    4 AA3 4 PHE B  73  ASP B  78 -1  N  PHE B  76   O  TYR B  85
SHEET    1 AA4 6 GLY B  12  PRO B  15  0
SHEET    2 AA4 6 THR B 123  SER B 128  1  O  THR B 126   N  VAL B  14
SHEET    3 AA4 6 ALA B  97  ALA B 104 -1  N  TYR B  99   O  THR B 123
SHEET    4 AA4 6 ALA B  35  GLN B  41 -1  N  PHE B  39   O  TYR B 100
SHEET    5 AA4 6 GLU B  48  VAL B  53 -1  O  GLU B  48   N  ARG B  40
SHEET    6 AA4 6 ALA B  63  VAL B  65 -1  O  VAL B  64   N  ALA B  52
SHEET    1 AA5 8 PHE C  34  VAL C  41  0
SHEET    2 AA5 8 VAL C  44  GLY C  52 -1  O  LEU C  46   N  ARG C  39
SHEET    3 AA5 8 THR C  82  PRO C  86 -1  O  VAL C  83   N  GLY C  51
SHEET    4 AA5 8 LEU C  56  VAL C  60  1  N  VAL C  57   O  THR C  82
SHEET    5 AA5 8 PHE C 123  HIS C 128  1  O  ASP C 124   N  LEU C  56
SHEET    6 AA5 8 ILE C 146  MET C 152  1  O  VAL C 150   N  LEU C 125
SHEET    7 AA5 8 THR C 261  GLY C 266  1  O  MET C 262   N  TYR C 151
SHEET    8 AA5 8 VAL C 287  LEU C 292  1  O  LEU C 292   N  ALA C 265
SHEET    1 AA6 2 PHE C 167  THR C 168  0
SHEET    2 AA6 2 GLY C 171  GLU C 172 -1  O  GLY C 171   N  THR C 168
SHEET    1 AA7 4 VAL A   7  SER A   9  0
SHEET    2 AA7 4 LEU A  20  THR A  25 -1  O  SER A  23   N  SER A   9
SHEET    3 AA7 4 THR A  83  MET A  88 -1  O  MET A  88   N  LEU A  20
SHEET    4 AA7 4 PHE A  73  ASP A  78 -1  N  ASP A  78   O  THR A  83
SHEET    1 AA8 6 LEU A  13  PRO A  15  0
SHEET    2 AA8 6 THR A 123  SER A 128  1  O  THR A 126   N  VAL A  14
SHEET    3 AA8 6 ALA A  97  ALA A 104 -1  N  TYR A  99   O  THR A 123
SHEET    4 AA8 6 ALA A  35  GLN A  41 -1  N  ALA A  35   O  ALA A 104
SHEET    5 AA8 6 GLU A  48  VAL A  53 -1  O  ALA A  51   N  TRP A  38
SHEET    6 AA8 6 ALA A  63  VAL A  65 -1  O  VAL A  64   N  ALA A  52
SSBOND   1 CYS D  295    CYS D  303                          1555   1555  2.02
SSBOND   2 CYS B   24    CYS B  101                          1555   1555  2.04
SSBOND   3 CYS C  295    CYS C  303                          1555   1555  2.21
SSBOND   4 CYS A   24    CYS A  101                          1555   1555  2.02
CRYST1  115.185  185.941  106.753  90.00  90.00  90.00 C 2 2 21     16
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008682  0.000000  0.000000        0.00000
SCALE2      0.000000  0.005378  0.000000        0.00000
SCALE3      0.000000  0.000000  0.009367        0.00000
TER    2373      ARG D 317
TER    3303      SER B 129
TER    5682      GLY C 318
TER    6624      SER A 129
MASTER      416    0    0   39   40    0    0    6 7311    4    8   72
END