longtext: 8qdb-pdb

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HEADER    LYASE                                   28-AUG-23   8QDB
TITLE     WDYG1P SER256DHA (PSF)
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: YELLOWISH-GREEN 1-LIKE PROTEIN;
COMPND   3 CHAIN: A, B;
COMPND   4 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: EXOPHIALA DERMATITIDIS;
SOURCE   3 ORGANISM_TAXID: 5970;
SOURCE   4 GENE: YG1;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   7 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   8 EXPRESSION_SYSTEM_PLASMID: PCDFDUET
KEYWDS    NATURAL PRODUCT BIOSYNTHESIS, POLYKETIDE SYNTHASE SYSTEM, ALPHA,
KEYWDS   2 BETA-HYDROLASE FOLD, POLYKETIDE SHORTENING, RETRO CLAISEN REACTION,
KEYWDS   3 LYASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.SCHMALHOFER,A.L.VAGSTAD,Q.ZHOU,H.B.BODE,M.GROLL
REVDAT   1   13-MAR-24 8QDB    0
JRNL        AUTH   M.SCHMALHOFER,A.L.VAGSTAD,Q.ZHOU,H.B.BODE,M.GROLL
JRNL        TITL   POLYKETIDE TRIMMING SHAPES DIHYDROXYNAPHTHALENE-MELANIN AND
JRNL        TITL 2 ANTHRAQUINONE PIGMENTS
JRNL        REF    ADV SCI                                    2024
JRNL        REFN                   ESSN 2198-3844
JRNL        DOI    10.1002/ADVS.202400184
REMARK   2
REMARK   2 RESOLUTION.    1.85 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0267
REMARK   3   AUTHORS     : MURSHUDOV,SKUBAK,LEBEDEV,PANNU,STEINER,
REMARK   3               : NICHOLLS,WINN,LONG,VAGIN
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.85
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 43.29
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 98.7
REMARK   3   NUMBER OF REFLECTIONS             : 74434
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.153
REMARK   3   R VALUE            (WORKING SET) : 0.151
REMARK   3   FREE R VALUE                     : 0.190
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 3918
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.85
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.90
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 5459
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 99.48
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2030
REMARK   3   BIN FREE R VALUE SET COUNT          : 288
REMARK   3   BIN FREE R VALUE                    : 0.3070
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 6574
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 12
REMARK   3   SOLVENT ATOMS            : 739
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 30.16
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 1.90000
REMARK   3    B22 (A**2) : 5.51000
REMARK   3    B33 (A**2) : -7.41000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.056
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.024
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.064
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 4.600
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.973
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.961
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  6769 ; 0.003 ; 0.013
REMARK   3   BOND LENGTHS OTHERS               (A):  6330 ; 0.001 ; 0.016
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  9161 ; 1.200 ; 1.646
REMARK   3   BOND ANGLES OTHERS          (DEGREES): 14683 ; 1.172 ; 1.580
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   820 ; 6.317 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   332 ;31.571 ;22.831
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):  1173 ;12.804 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    30 ;16.548 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   845 ; 0.058 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  7534 ; 0.004 ; 0.020
REMARK   3   GENERAL PLANES OTHERS             (A):  1486 ; 0.001 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  3286 ; 0.857 ; 1.928
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  3285 ; 0.855 ; 1.927
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  4104 ; 1.145 ; 2.892
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  4105 ; 1.145 ; 2.893
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  3483 ; 1.004 ; 2.104
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  3484 ; 1.005 ; 2.104
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  5058 ; 1.264 ; 3.086
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  8084 ; 2.305 ;24.467
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  8085 ; 2.305 ;24.471
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2): 13099 ; 0.589 ; 3.000
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 2
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A     2        A   410
REMARK   3    ORIGIN FOR THE GROUP (A):    2.280   17.673   21.418
REMARK   3    T TENSOR
REMARK   3      T11:   0.0217 T22:   0.0314
REMARK   3      T33:   0.0623 T12:   0.0066
REMARK   3      T13:   0.0001 T23:  -0.0152
REMARK   3    L TENSOR
REMARK   3      L11:   0.9523 L22:   0.8864
REMARK   3      L33:   1.1987 L12:  -0.1796
REMARK   3      L13:   0.2202 L23:   0.0996
REMARK   3    S TENSOR
REMARK   3      S11:   0.0181 S12:   0.1355 S13:  -0.0883
REMARK   3      S21:  -0.0933 S22:  -0.0965 S23:   0.0810
REMARK   3      S31:   0.1038 S32:  -0.0040 S33:   0.0784
REMARK   3
REMARK   3   TLS GROUP : 2
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   B     2        B   412
REMARK   3    ORIGIN FOR THE GROUP (A):   42.669   23.404   43.772
REMARK   3    T TENSOR
REMARK   3      T11:   0.0042 T22:   0.1512
REMARK   3      T33:   0.1214 T12:  -0.0069
REMARK   3      T13:  -0.0083 T23:   0.0620
REMARK   3    L TENSOR
REMARK   3      L11:   1.1863 L22:   0.5394
REMARK   3      L33:   0.7190 L12:   0.1055
REMARK   3      L13:   0.0793 L23:   0.0212
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0013 S12:   0.0371 S13:   0.1425
REMARK   3      S21:  -0.0230 S22:  -0.0174 S23:  -0.1031
REMARK   3      S31:  -0.0265 S32:   0.1255 S33:   0.0186
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS
REMARK   4
REMARK   4 8QDB COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 28-AUG-23.
REMARK 100 THE DEPOSITION ID IS D_1292133002.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 20-MAR-21
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 9.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SLS
REMARK 200  BEAMLINE                       : X06SA
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS EIGER X 16M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 78363
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.850
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.6
REMARK 200  DATA REDUNDANCY                : 5.300
REMARK 200  R MERGE                    (I) : 0.05900
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 16.0000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.85
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.95
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.5
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : 0.63100
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.800
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: REFMAC
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 47.44
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.34
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 100 MM TRIS PH 9.0, 23% PEG 3350,
REMARK 280  VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       42.75000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       61.00500
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       44.16000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       61.00500
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       42.75000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       44.16000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A   -16
REMARK 465     GLY A   -15
REMARK 465     TRP A   -14
REMARK 465     SER A   -13
REMARK 465     HIS A   -12
REMARK 465     PRO A   -11
REMARK 465     GLN A   -10
REMARK 465     PHE A    -9
REMARK 465     GLU A    -8
REMARK 465     LYS A    -7
REMARK 465     GLU A    -6
REMARK 465     ASN A    -5
REMARK 465     LEU A    -4
REMARK 465     TYR A    -3
REMARK 465     PHE A    -2
REMARK 465     GLN A    -1
REMARK 465     GLY A     0
REMARK 465     SER A     1
REMARK 465     ARG A   412
REMARK 465     THR A   413
REMARK 465     LYS A   414
REMARK 465     MET B   -16
REMARK 465     GLY B   -15
REMARK 465     TRP B   -14
REMARK 465     SER B   -13
REMARK 465     HIS B   -12
REMARK 465     PRO B   -11
REMARK 465     GLN B   -10
REMARK 465     PHE B    -9
REMARK 465     GLU B    -8
REMARK 465     LYS B    -7
REMARK 465     GLU B    -6
REMARK 465     ASN B    -5
REMARK 465     LEU B    -4
REMARK 465     TYR B    -3
REMARK 465     PHE B    -2
REMARK 465     GLN B    -1
REMARK 465     GLY B     0
REMARK 465     SER B     1
REMARK 465     LYS B   414
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     GLU A   4    CG   CD   OE1  OE2
REMARK 470     LYS A 144    CG   CD   CE   NZ
REMARK 470     LYS B 403    CG   CD   CE   NZ
REMARK 470     GLU B 404    CG   CD   OE1  OE2
REMARK 470     ARG B 412    CG   CD   NE   CZ   NH1  NH2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ARG A   9     -124.88     56.19
REMARK 500    LYS A  29      -46.38   -134.93
REMARK 500    ASN A 170       -0.72     75.94
REMARK 500    TYR A 189     -164.83   -111.24
REMARK 500    ALA A 283      -34.37   -157.19
REMARK 500    HIS A 284      -70.40   -152.89
REMARK 500    LEU A 298     -147.24   -108.20
REMARK 500    PHE A 302     -142.89   -118.84
REMARK 500    LYS A 377     -151.98    -95.88
REMARK 500    MET A 379      -11.48     83.57
REMARK 500    ARG B   9     -121.03     57.53
REMARK 500    LYS B  29      -48.93   -141.60
REMARK 500    PHE B  43       -0.94     72.08
REMARK 500    TYR B 189     -162.53   -111.65
REMARK 500    ALA B 283      -31.22   -157.30
REMARK 500    HIS B 284      -68.41   -155.14
REMARK 500    LEU B 298     -146.41   -102.54
REMARK 500    PHE B 302     -144.86   -120.14
REMARK 500    THR B 340     -178.18    -68.80
REMARK 500    LYS B 377     -156.68    -95.54
REMARK 500    MET B 379       -9.93     82.79
REMARK 500
REMARK 500 REMARK: NULL
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 6HXA   RELATED DB: PDB
REMARK 900 ANTI FROM P. LUMINESCENS CATALYSES TERMINAL POLYKETIDE SHORTENING
REMARK 900 IN THE BIOSYNTHESIS OF ANTHRAQUINONES
DBREF  8QDB A    2   414  UNP    Q6BCB5   Q6BCB5_EXODE     2    414
DBREF  8QDB B    2   414  UNP    Q6BCB5   Q6BCB5_EXODE     2    414
SEQADV 8QDB MET A  -16  UNP  Q6BCB5              INITIATING METHIONINE
SEQADV 8QDB GLY A  -15  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB TRP A  -14  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB SER A  -13  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB HIS A  -12  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB PRO A  -11  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB GLN A  -10  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB PHE A   -9  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB GLU A   -8  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB LYS A   -7  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB GLU A   -6  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB ASN A   -5  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB LEU A   -4  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB TYR A   -3  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB PHE A   -2  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB GLN A   -1  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB GLY A    0  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB SER A    1  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB MET B  -16  UNP  Q6BCB5              INITIATING METHIONINE
SEQADV 8QDB GLY B  -15  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB TRP B  -14  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB SER B  -13  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB HIS B  -12  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB PRO B  -11  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB GLN B  -10  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB PHE B   -9  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB GLU B   -8  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB LYS B   -7  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB GLU B   -6  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB ASN B   -5  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB LEU B   -4  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB TYR B   -3  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB PHE B   -2  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB GLN B   -1  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB GLY B    0  UNP  Q6BCB5              EXPRESSION TAG
SEQADV 8QDB SER B    1  UNP  Q6BCB5              EXPRESSION TAG
SEQRES   1 A  431  MET GLY TRP SER HIS PRO GLN PHE GLU LYS GLU ASN LEU
SEQRES   2 A  431  TYR PHE GLN GLY SER ALA THR GLU LYS TYR TYR ILE ARG
SEQRES   3 A  431  ASP ALA ILE THR LYS PRO ALA VAL HIS HIS GLU SER TYR
SEQRES   4 A  431  GLN LYS LEU TRP GLU THR LYS TRP LYS LYS PRO CYS GLU
SEQRES   5 A  431  MET GLY VAL TYR PRO PHE MET PHE GLY SER ILE LYS ASP
SEQRES   6 A  431  PHE GLU PRO VAL ALA GLN GLU ILE ILE LYS LYS GLY LEU
SEQRES   7 A  431  LYS GLU PRO TYR ASP TRP ASP GLU TYR ALA GLN MET TYR
SEQRES   8 A  431  PHE PRO LYS ALA GLU GLU LEU ALA LYS ILE ALA GLU GLU
SEQRES   9 A  431  ALA GLU ALA ALA GLY GLU LYS GLU LYS ALA SER GLU TYR
SEQRES  10 A  431  TYR LEU ARG SER SER ALA VAL TYR ARG ILE SER ARG PHE
SEQRES  11 A  431  PRO THR PRO ARG SER GLU LYS GLN LYS TYR ALA TRP ARG
SEQRES  12 A  431  LYS GLY CYS GLU VAL PHE TYR LYS GLY ALA ALA LEU MET
SEQRES  13 A  431  GLU TYR PRO ILE LYS GLU VAL ARG ILE PRO HIS LYS HIS
SEQRES  14 A  431  GLY ILE GLU GLY GLU GLY ASP VAL VAL PRO VAL ASN PHE
SEQRES  15 A  431  LEU LEU PRO PRO ASN ALA SER GLU THR SER PRO VAL PRO
SEQRES  16 A  431  CYS VAL LEU ILE ILE THR GLY LEU ASP GLY TYR ARG THR
SEQRES  17 A  431  GLU LEU ALA VAL TRP GLN GLN GLY TRP ARG SER LYS GLY
SEQRES  18 A  431  VAL ALA THR VAL ILE ALA GLU ILE PRO GLY THR GLY ASP
SEQRES  19 A  431  SER PRO ALA LEU ARG GLN ASP PRO THR SER PRO ASP ARG
SEQRES  20 A  431  GLN TRP SER SER VAL LEU ASP TRP ILE GLU SER GLN LYS
SEQRES  21 A  431  ALA VAL ASP SER LYS LYS ILE VAL ALA TRP GLY PHE DHA
SEQRES  22 A  431  THR GLY GLY TYR TYR ALA LEU ARG MET ALA HIS THR HIS
SEQRES  23 A  431  LYS ASP ARG LEU LEU ALA THR ILE SER LEU GLY GLY GLY
SEQRES  24 A  431  ALA HIS HIS MET PHE ASP ARG GLU TRP LEU GLU HIS ALA
SEQRES  25 A  431  ASN LYS LEU GLU TYR PRO PHE ASP LEU SER ASN THR LEU
SEQRES  26 A  431  ALA TYR LYS PHE GLY TYR PRO ASP LEU GLU SER PHE ILE
SEQRES  27 A  431  GLU GLU SER SER LYS PHE SER LEU LEU ASN ASP GLY THR
SEQRES  28 A  431  LEU GLN LYS PRO CYS THR LYS VAL LEU LEU VAL ASN GLY
SEQRES  29 A  431  ASN ASP ASP GLU ILE PHE PRO ILE ASP ASP MET PHE VAL
SEQRES  30 A  431  SER LEU GLU ASN GLY GLN PRO LYS LEU ALA ARG MET VAL
SEQRES  31 A  431  LYS GLY LYS LYS HIS MET GLY GLU PRO GLU SER PHE SER
SEQRES  32 A  431  ILE ILE LEU GLU TRP ILE HIS LYS LEU LEU GLY LEU ASP
SEQRES  33 A  431  GLY LYS ILE LYS GLU GLN LEU ALA MET ILE PRO SER ARG
SEQRES  34 A  431  THR LYS
SEQRES   1 B  431  MET GLY TRP SER HIS PRO GLN PHE GLU LYS GLU ASN LEU
SEQRES   2 B  431  TYR PHE GLN GLY SER ALA THR GLU LYS TYR TYR ILE ARG
SEQRES   3 B  431  ASP ALA ILE THR LYS PRO ALA VAL HIS HIS GLU SER TYR
SEQRES   4 B  431  GLN LYS LEU TRP GLU THR LYS TRP LYS LYS PRO CYS GLU
SEQRES   5 B  431  MET GLY VAL TYR PRO PHE MET PHE GLY SER ILE LYS ASP
SEQRES   6 B  431  PHE GLU PRO VAL ALA GLN GLU ILE ILE LYS LYS GLY LEU
SEQRES   7 B  431  LYS GLU PRO TYR ASP TRP ASP GLU TYR ALA GLN MET TYR
SEQRES   8 B  431  PHE PRO LYS ALA GLU GLU LEU ALA LYS ILE ALA GLU GLU
SEQRES   9 B  431  ALA GLU ALA ALA GLY GLU LYS GLU LYS ALA SER GLU TYR
SEQRES  10 B  431  TYR LEU ARG SER SER ALA VAL TYR ARG ILE SER ARG PHE
SEQRES  11 B  431  PRO THR PRO ARG SER GLU LYS GLN LYS TYR ALA TRP ARG
SEQRES  12 B  431  LYS GLY CYS GLU VAL PHE TYR LYS GLY ALA ALA LEU MET
SEQRES  13 B  431  GLU TYR PRO ILE LYS GLU VAL ARG ILE PRO HIS LYS HIS
SEQRES  14 B  431  GLY ILE GLU GLY GLU GLY ASP VAL VAL PRO VAL ASN PHE
SEQRES  15 B  431  LEU LEU PRO PRO ASN ALA SER GLU THR SER PRO VAL PRO
SEQRES  16 B  431  CYS VAL LEU ILE ILE THR GLY LEU ASP GLY TYR ARG THR
SEQRES  17 B  431  GLU LEU ALA VAL TRP GLN GLN GLY TRP ARG SER LYS GLY
SEQRES  18 B  431  VAL ALA THR VAL ILE ALA GLU ILE PRO GLY THR GLY ASP
SEQRES  19 B  431  SER PRO ALA LEU ARG GLN ASP PRO THR SER PRO ASP ARG
SEQRES  20 B  431  GLN TRP SER SER VAL LEU ASP TRP ILE GLU SER GLN LYS
SEQRES  21 B  431  ALA VAL ASP SER LYS LYS ILE VAL ALA TRP GLY PHE DHA
SEQRES  22 B  431  THR GLY GLY TYR TYR ALA LEU ARG MET ALA HIS THR HIS
SEQRES  23 B  431  LYS ASP ARG LEU LEU ALA THR ILE SER LEU GLY GLY GLY
SEQRES  24 B  431  ALA HIS HIS MET PHE ASP ARG GLU TRP LEU GLU HIS ALA
SEQRES  25 B  431  ASN LYS LEU GLU TYR PRO PHE ASP LEU SER ASN THR LEU
SEQRES  26 B  431  ALA TYR LYS PHE GLY TYR PRO ASP LEU GLU SER PHE ILE
SEQRES  27 B  431  GLU GLU SER SER LYS PHE SER LEU LEU ASN ASP GLY THR
SEQRES  28 B  431  LEU GLN LYS PRO CYS THR LYS VAL LEU LEU VAL ASN GLY
SEQRES  29 B  431  ASN ASP ASP GLU ILE PHE PRO ILE ASP ASP MET PHE VAL
SEQRES  30 B  431  SER LEU GLU ASN GLY GLN PRO LYS LEU ALA ARG MET VAL
SEQRES  31 B  431  LYS GLY LYS LYS HIS MET GLY GLU PRO GLU SER PHE SER
SEQRES  32 B  431  ILE ILE LEU GLU TRP ILE HIS LYS LEU LEU GLY LEU ASP
SEQRES  33 B  431  GLY LYS ILE LYS GLU GLN LEU ALA MET ILE PRO SER ARG
SEQRES  34 B  431  THR LYS
MODRES 8QDB DHA A  256  SER  MODIFIED RESIDUE
MODRES 8QDB DHA B  256  SER  MODIFIED RESIDUE
HET    DHA  A 256       5
HET    DHA  B 256       5
HET    EDO  A 501       4
HET    EDO  A 502       4
HET    EDO  A 503       4
HETNAM     DHA 2-AMINO-ACRYLIC ACID
HETNAM     EDO 1,2-ETHANEDIOL
HETSYN     DHA 2,3-DIDEHYDROALANINE
HETSYN     EDO ETHYLENE GLYCOL
FORMUL   1  DHA    2(C3 H5 N O2)
FORMUL   3  EDO    3(C2 H6 O2)
FORMUL   6  HOH   *739(H2 O)
HELIX    1 AA1 ILE A    8  LYS A   14  5                                   7
HELIX    2 AA2 ALA A   16  GLU A   20  5                                   5
HELIX    3 AA3 SER A   21  LYS A   29  1                                   9
HELIX    4 AA4 TRP A   30  GLY A   37  1                                   8
HELIX    5 AA5 SER A   45  GLY A   60  1                                  16
HELIX    6 AA6 ASP A   66  MET A   73  1                                   8
HELIX    7 AA7 TYR A   74  ALA A   91  1                                  18
HELIX    8 AA8 GLU A   93  ARG A  112  1                                  20
HELIX    9 AA9 SER A  118  LEU A  138  1                                  21
HELIX   10 AB1 TYR A  189  ALA A  194  1                                   6
HELIX   11 AB2 TRP A  196  LYS A  203  1                                   8
HELIX   12 AB3 THR A  226  GLN A  242  1                                  17
HELIX   13 AB4 THR A  257  HIS A  269  1                                  13
HELIX   14 AB5 LYS A  270  LEU A  273  5                                   4
HELIX   15 AB6 HIS A  284  PHE A  287  5                                   4
HELIX   16 AB7 ASP A  288  ASN A  296  1                                   9
HELIX   17 AB8 ASP A  303  PHE A  312  1                                  10
HELIX   18 AB9 ASP A  316  SER A  324  1                                   9
HELIX   19 AC1 SER A  325  SER A  328  5                                   4
HELIX   20 AC2 GLY A  333  LYS A  337  5                                   5
HELIX   21 AC3 ILE A  355  GLU A  363  1                                   9
HELIX   22 AC4 LYS A  377  GLU A  381  5                                   5
HELIX   23 AC5 PRO A  382  GLY A  397  1                                  16
HELIX   24 AC6 LYS A  401  ALA A  407  1                                   7
HELIX   25 AC7 ILE B    8  LYS B   14  5                                   7
HELIX   26 AC8 ALA B   16  GLU B   20  5                                   5
HELIX   27 AC9 SER B   21  LYS B   29  1                                   9
HELIX   28 AD1 TRP B   30  GLY B   37  1                                   8
HELIX   29 AD2 SER B   45  LYS B   59  1                                  15
HELIX   30 AD3 ASP B   66  MET B   73  1                                   8
HELIX   31 AD4 TYR B   74  ALA B   91  1                                  18
HELIX   32 AD5 GLU B   93  ARG B  112  1                                  20
HELIX   33 AD6 SER B  118  ALA B  137  1                                  20
HELIX   34 AD7 TYR B  189  ALA B  194  1                                   6
HELIX   35 AD8 TRP B  196  LYS B  203  1                                   8
HELIX   36 AD9 THR B  226  GLN B  242  1                                  17
HELIX   37 AE1 THR B  257  HIS B  269  1                                  13
HELIX   38 AE2 LYS B  270  LEU B  273  5                                   4
HELIX   39 AE3 HIS B  284  PHE B  287  5                                   4
HELIX   40 AE4 ASP B  288  ASN B  296  1                                   9
HELIX   41 AE5 ASP B  303  PHE B  312  1                                  10
HELIX   42 AE6 ASP B  316  SER B  324  1                                   9
HELIX   43 AE7 SER B  325  SER B  328  5                                   4
HELIX   44 AE8 GLY B  333  LYS B  337  5                                   5
HELIX   45 AE9 ILE B  355  GLU B  363  1                                   9
HELIX   46 AF1 LYS B  377  GLU B  381  5                                   5
HELIX   47 AF2 PRO B  382  GLY B  397  1                                  16
HELIX   48 AF3 LYS B  401  MET B  408  1                                   8
SHEET    1 AA116 LYS A 144  PRO A 149  0
SHEET    2 AA116 VAL A 160  LEU A 166 -1  O  VAL A 161   N  ILE A 148
SHEET    3 AA116 ALA A 206  ALA A 210 -1  O  THR A 207   N  LEU A 166
SHEET    4 AA116 VAL A 177  ILE A 183  1  N  ILE A 182   O  VAL A 208
SHEET    5 AA116 VAL A 245  PHE A 255  1  O  VAL A 251   N  LEU A 181
SHEET    6 AA116 ALA A 275  LEU A 279  1  O  ILE A 277   N  GLY A 254
SHEET    7 AA116 LYS A 341  GLY A 347  1  O  LEU A 343   N  SER A 278
SHEET    8 AA116 LYS A 368  VAL A 373  1  O  LEU A 369   N  VAL A 342
SHEET    9 AA116 LYS B 368  VAL B 373 -1  O  LYS B 368   N  ALA A 370
SHEET   10 AA116 LYS B 341  GLY B 347  1  N  VAL B 342   O  LEU B 369
SHEET   11 AA116 ALA B 275  LEU B 279  1  N  SER B 278   O  VAL B 345
SHEET   12 AA116 VAL B 245  PHE B 255  1  N  GLY B 254   O  ILE B 277
SHEET   13 AA116 VAL B 177  ILE B 183  1  N  VAL B 177   O  ASP B 246
SHEET   14 AA116 ALA B 206  ALA B 210  1  O  VAL B 208   N  ILE B 182
SHEET   15 AA116 VAL B 160  LEU B 166 -1  N  LEU B 166   O  THR B 207
SHEET   16 AA116 LYS B 144  PRO B 149 -1  N  ILE B 148   O  VAL B 161
LINK         C   PHE A 255                 N   DHA A 256     1555   1555  1.33
LINK         C   DHA A 256                 N   THR A 257     1555   1555  1.34
LINK         C   PHE B 255                 N   DHA B 256     1555   1555  1.34
LINK         C   DHA B 256                 N   THR B 257     1555   1555  1.34
CISPEP   1 TYR A   39    PRO A   40          0        -0.21
CISPEP   2 GLU A   63    PRO A   64          0        -4.68
CISPEP   3 PHE A  113    PRO A  114          0         0.38
CISPEP   4 GLU A  381    PRO A  382          0        -1.41
CISPEP   5 TYR B   39    PRO B   40          0        -0.84
CISPEP   6 GLU B   63    PRO B   64          0        -3.64
CISPEP   7 PHE B  113    PRO B  114          0        -1.59
CISPEP   8 GLU B  381    PRO B  382          0        -2.79
CRYST1   85.500   88.320  122.010  90.00  90.00  90.00 P 21 21 21    8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.011696  0.000000  0.000000        0.00000
SCALE2      0.000000  0.011322  0.000000        0.00000
SCALE3      0.000000  0.000000  0.008196        0.00000
TER    3282      SER A 411
TER    6576      THR B 413
MASTER      378    0    5   48   16    0    0    6 7325    2   26   68
END