longtext: 8qrj-pdb

content
HEADER    HYDROLASE                               09-OCT-23   8QRJ
TITLE     LCC-ICCG PETASE MUTANT H218Y
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LEAF-BRANCH COMPOST CUTINASE;
COMPND   3 CHAIN: A;
COMPND   4 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: UNIDENTIFIED PROKARYOTIC ORGANISM;
SOURCE   3 ORGANISM_TAXID: 2725;
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   5 EXPRESSION_SYSTEM_TAXID: 562
KEYWDS    LCC, PETASE, HYDROLASE, DIRECT ENZYME EVOLUTION
EXPDTA    X-RAY DIFFRACTION
AUTHOR    G.ORR,Y.NIV,M.BARAKAT,A.BOGINYA,M.DESSAU,L.AFRIAT-JURNOU
REVDAT   1   18-SEP-24 8QRJ    0
JRNL        AUTH   G.ORR,Y.NIV,M.BARAKAT,A.BOGINYA,M.DESSAU,L.AFRIAT-JURNOU
JRNL        TITL   STREAMLINED SCREENING OF EXTRACELLULARLY EXPRESSED PETASE
JRNL        TITL 2 LIBRARIES FOR IMPROVED POLYETHYLENE TEREPHTHALATE
JRNL        TITL 3 DEGRADATION.
JRNL        REF    BIOTECHNOL J                  V.  19 00021 2024
JRNL        REFN                   ESSN 1860-7314
JRNL        PMID   38987219
JRNL        DOI    10.1002/BIOT.202400021
REMARK   2
REMARK   2 RESOLUTION.    1.42 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (1.20.1_4487: ???)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.42
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 49.17
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.340
REMARK   3   COMPLETENESS FOR RANGE        (%) : 95.2
REMARK   3   NUMBER OF REFLECTIONS             : 56565
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.168
REMARK   3   R VALUE            (WORKING SET) : 0.167
REMARK   3   FREE R VALUE                     : 0.182
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.070
REMARK   3   FREE R VALUE TEST SET COUNT      : 2868
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 49.1700 -  3.8500    0.95     2851   146  0.1377 0.1273
REMARK   3     2  3.8500 -  3.0600    0.93     2716   125  0.1398 0.1426
REMARK   3     3  3.0600 -  2.6700    0.95     2701   154  0.1506 0.1793
REMARK   3     4  2.6700 -  2.4300    0.94     2658   162  0.1498 0.1732
REMARK   3     5  2.4300 -  2.2500    0.96     2710   153  0.1442 0.1456
REMARK   3     6  2.2500 -  2.1200    0.96     2724   136  0.1373 0.1564
REMARK   3     7  2.1200 -  2.0200    0.96     2714   146  0.1459 0.1744
REMARK   3     8  2.0100 -  1.9300    0.96     2701   148  0.1615 0.1850
REMARK   3     9  1.9300 -  1.8500    0.97     2724   144  0.1713 0.2018
REMARK   3    10  1.8500 -  1.7900    0.96     2698   132  0.1718 0.1797
REMARK   3    11  1.7900 -  1.7300    0.97     2729   142  0.1814 0.2217
REMARK   3    12  1.7300 -  1.6800    0.97     2704   149  0.1977 0.2056
REMARK   3    13  1.6800 -  1.6400    0.94     2625   150  0.2174 0.2567
REMARK   3    14  1.6400 -  1.6000    0.95     2653   140  0.2352 0.2693
REMARK   3    15  1.6000 -  1.5600    0.96     2693   143  0.2447 0.2669
REMARK   3    16  1.5600 -  1.5300    0.94     2628   144  0.2480 0.2592
REMARK   3    17  1.5300 -  1.5000    0.93     2601   137  0.2588 0.2631
REMARK   3    18  1.5000 -  1.4700    0.93     2599   153  0.2696 0.3125
REMARK   3    19  1.4400 -  1.4200    0.95     2631   134  0.3076 0.3559
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.10
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.160
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 18.930
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.006           2046
REMARK   3   ANGLE     :  0.826           2799
REMARK   3   CHIRALITY :  0.078            315
REMARK   3   PLANARITY :  0.010            365
REMARK   3   DIHEDRAL  : 11.490            738
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 7
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 36 THROUGH 87 )
REMARK   3    ORIGIN FOR THE GROUP (A):  25.4305  25.6475  23.3517
REMARK   3    T TENSOR
REMARK   3      T11:   0.0916 T22:   0.1026
REMARK   3      T33:   0.0922 T12:   0.0099
REMARK   3      T13:   0.0005 T23:   0.0171
REMARK   3    L TENSOR
REMARK   3      L11:   1.1431 L22:   1.7911
REMARK   3      L33:   1.4497 L12:  -0.1389
REMARK   3      L13:  -0.0013 L23:   0.3859
REMARK   3    S TENSOR
REMARK   3      S11:   0.0079 S12:   0.0169 S13:   0.0950
REMARK   3      S21:  -0.0649 S22:  -0.0028 S23:   0.0721
REMARK   3      S31:  -0.0448 S32:  -0.0395 S33:  -0.0060
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 88 THROUGH 112 )
REMARK   3    ORIGIN FOR THE GROUP (A):  21.8365  16.4551  29.2542
REMARK   3    T TENSOR
REMARK   3      T11:   0.1120 T22:   0.1214
REMARK   3      T33:   0.1059 T12:   0.0020
REMARK   3      T13:   0.0006 T23:   0.0027
REMARK   3    L TENSOR
REMARK   3      L11:   0.7378 L22:   2.7892
REMARK   3      L33:   0.8028 L12:  -0.3049
REMARK   3      L13:   0.2825 L23:  -0.9622
REMARK   3    S TENSOR
REMARK   3      S11:   0.0170 S12:   0.0114 S13:  -0.0037
REMARK   3      S21:   0.0319 S22:   0.0609 S23:   0.1670
REMARK   3      S31:   0.0269 S32:  -0.1076 S33:  -0.0797
REMARK   3   TLS GROUP : 3
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 113 THROUGH 127 )
REMARK   3    ORIGIN FOR THE GROUP (A):  26.3844  10.0078  26.5078
REMARK   3    T TENSOR
REMARK   3      T11:   0.1143 T22:   0.1100
REMARK   3      T33:   0.1129 T12:   0.0002
REMARK   3      T13:   0.0013 T23:   0.0052
REMARK   3    L TENSOR
REMARK   3      L11:   0.2354 L22:   7.8782
REMARK   3      L33:   1.3097 L12:  -0.1130
REMARK   3      L13:   0.2802 L23:  -2.0080
REMARK   3    S TENSOR
REMARK   3      S11:   0.0377 S12:  -0.0196 S13:  -0.0468
REMARK   3      S21:  -0.1029 S22:   0.0308 S23:   0.2383
REMARK   3      S31:   0.1546 S32:  -0.0633 S33:  -0.0794
REMARK   3   TLS GROUP : 4
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 128 THROUGH 158 )
REMARK   3    ORIGIN FOR THE GROUP (A):  34.4081  16.6967  21.7311
REMARK   3    T TENSOR
REMARK   3      T11:   0.1066 T22:   0.1292
REMARK   3      T33:   0.1137 T12:   0.0060
REMARK   3      T13:  -0.0038 T23:   0.0042
REMARK   3    L TENSOR
REMARK   3      L11:   0.2709 L22:   1.2247
REMARK   3      L33:   0.5514 L12:   0.3191
REMARK   3      L13:  -0.3830 L23:  -0.3555
REMARK   3    S TENSOR
REMARK   3      S11:   0.0281 S12:   0.0165 S13:  -0.0150
REMARK   3      S21:  -0.0723 S22:  -0.0359 S23:  -0.0251
REMARK   3      S31:   0.0208 S32:   0.0186 S33:   0.0257
REMARK   3   TLS GROUP : 5
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 159 THROUGH 237 )
REMARK   3    ORIGIN FOR THE GROUP (A):  32.2367  14.4275  42.0210
REMARK   3    T TENSOR
REMARK   3      T11:   0.0958 T22:   0.0993
REMARK   3      T33:   0.1077 T12:   0.0044
REMARK   3      T13:  -0.0046 T23:   0.0028
REMARK   3    L TENSOR
REMARK   3      L11:   0.9619 L22:   0.8518
REMARK   3      L33:   1.5944 L12:   0.0529
REMARK   3      L13:  -0.5955 L23:  -0.0637
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0034 S12:  -0.0673 S13:  -0.0189
REMARK   3      S21:   0.0612 S22:  -0.0036 S23:  -0.0332
REMARK   3      S31:   0.0685 S32:   0.0955 S33:   0.0139
REMARK   3   TLS GROUP : 6
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 238 THROUGH 249 )
REMARK   3    ORIGIN FOR THE GROUP (A):  16.0059  15.9893  40.8143
REMARK   3    T TENSOR
REMARK   3      T11:   0.0732 T22:   0.1128
REMARK   3      T33:   0.1320 T12:  -0.0113
REMARK   3      T13:   0.0107 T23:  -0.0072
REMARK   3    L TENSOR
REMARK   3      L11:   0.7408 L22:   1.8108
REMARK   3      L33:   2.6709 L12:   0.3195
REMARK   3      L13:   0.3545 L23:  -0.0077
REMARK   3    S TENSOR
REMARK   3      S11:   0.0500 S12:  -0.0575 S13:   0.1307
REMARK   3      S21:  -0.0309 S22:  -0.0217 S23:   0.1116
REMARK   3      S31:   0.0836 S32:  -0.2210 S33:  -0.0054
REMARK   3   TLS GROUP : 7
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 250 THROUGH 292 )
REMARK   3    ORIGIN FOR THE GROUP (A):  28.5148  29.2065  41.9501
REMARK   3    T TENSOR
REMARK   3      T11:   0.1162 T22:   0.1074
REMARK   3      T33:   0.1169 T12:  -0.0062
REMARK   3      T13:   0.0120 T23:  -0.0190
REMARK   3    L TENSOR
REMARK   3      L11:   0.6051 L22:   0.8719
REMARK   3      L33:   2.2565 L12:   0.2911
REMARK   3      L13:  -0.7132 L23:  -0.6465
REMARK   3    S TENSOR
REMARK   3      S11:   0.0861 S12:  -0.0962 S13:   0.1067
REMARK   3      S21:   0.0920 S22:  -0.0367 S23:  -0.0190
REMARK   3      S31:  -0.2223 S32:   0.1221 S33:  -0.0578
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 8QRJ COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 10-OCT-23.
REMARK 100 THE DEPOSITION ID IS D_1292133744.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 04-JUL-23
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 7.3
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID30B
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.8731
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 93953
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.420
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 82.0
REMARK 200  DATA REDUNDANCY                : 1.770
REMARK 200  R MERGE                    (I) : 0.09900
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 4.6500
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.42
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.46
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : 0.89400
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHENIX
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 50.67
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.49
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 20% PEG 3350, 0.2 M NA-FORMATE PH=7.3,
REMARK 280  PH 7.3, VAPOR DIFFUSION, TEMPERATURE 289K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 2 2 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z+1/2
REMARK 290       3555   -X,Y,-Z+1/2
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z
REMARK 290       6555   -X+1/2,-Y+1/2,Z+1/2
REMARK 290       7555   -X+1/2,Y+1/2,-Z+1/2
REMARK 290       8555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       40.47500
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       40.47500
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       40.95000
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       47.25500
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       40.95000
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       47.25500
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       40.47500
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       40.95000
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       47.25500
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       40.47500
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       40.95000
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000       47.25500
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH A 682  LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     ASP A     2
REMARK 465     GLY A     3
REMARK 465     VAL A     4
REMARK 465     LEU A     5
REMARK 465     TRP A     6
REMARK 465     ARG A     7
REMARK 465     VAL A     8
REMARK 465     ARG A     9
REMARK 465     THR A    10
REMARK 465     ALA A    11
REMARK 465     ALA A    12
REMARK 465     LEU A    13
REMARK 465     MET A    14
REMARK 465     ALA A    15
REMARK 465     ALA A    16
REMARK 465     LEU A    17
REMARK 465     LEU A    18
REMARK 465     ALA A    19
REMARK 465     LEU A    20
REMARK 465     ALA A    21
REMARK 465     ALA A    22
REMARK 465     TRP A    23
REMARK 465     ALA A    24
REMARK 465     LEU A    25
REMARK 465     VAL A    26
REMARK 465     TRP A    27
REMARK 465     ALA A    28
REMARK 465     SER A    29
REMARK 465     PRO A    30
REMARK 465     SER A    31
REMARK 465     VAL A    32
REMARK 465     GLU A    33
REMARK 465     ALA A    34
REMARK 465     GLN A    35
REMARK 465     GLN A   293
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    HOH A   808     O    HOH A   827              1.87
REMARK 500   O    HOH A   769     O    HOH A   804              1.91
REMARK 500   O    HOH A   414     O    HOH A   441              1.93
REMARK 500   O    HOH A   575     O    HOH A   615              1.99
REMARK 500   O    HOH A   703     O    HOH A   753              2.01
REMARK 500   O    HOH A   686     O    HOH A   721              2.04
REMARK 500   O    HOH A   683     O    HOH A   821              2.06
REMARK 500   O    HOH A   516     O    HOH A   768              2.06
REMARK 500   O    HOH A   618     O    HOH A   641              2.07
REMARK 500   O    HOH A   602     O    HOH A   805              2.09
REMARK 500   O    HOH A   528     O    HOH A   553              2.12
REMARK 500   O    HOH A   441     O    HOH A   732              2.13
REMARK 500   NH1  ARG A   107     O    HOH A   401              2.16
REMARK 500   O    HOH A   702     O    HOH A   706              2.16
REMARK 500   O    HOH A   738     O    HOH A   769              2.16
REMARK 500   O    HOH A   760     O    HOH A   834              2.17
REMARK 500   O    HOH A   571     O    HOH A   707              2.18
REMARK 500   O    HOH A   777     O    HOH A   794              2.18
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O    HOH A   599     O    HOH A   599     4556     1.94
REMARK 500   O    HOH A   618     O    HOH A   781     6554     2.17
REMARK 500   O    HOH A   545     O    HOH A   687     4556     2.18
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    THR A  96       -3.80     71.35
REMARK 500    SER A 165     -122.52     64.27
REMARK 500    THR A 188       58.28     33.65
REMARK 500    TYR A 218      -82.35   -124.24
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 837        DISTANCE =  5.82 ANGSTROMS
REMARK 525    HOH A 838        DISTANCE =  5.85 ANGSTROMS
REMARK 525    HOH A 839        DISTANCE =  5.93 ANGSTROMS
REMARK 525    HOH A 840        DISTANCE =  5.95 ANGSTROMS
REMARK 525    HOH A 841        DISTANCE =  6.27 ANGSTROMS
REMARK 525    HOH A 842        DISTANCE =  6.28 ANGSTROMS
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              CA A 303  CA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ARG A  65   O
REMARK 620 2 VAL A  68   O    83.0
REMARK 620 3 PHE A  71   O   106.8  96.4
REMARK 620 4 HOH A 516   O   177.9  95.9  71.5
REMARK 620 5 HOH A 542   O   108.1 166.4  73.2  72.7
REMARK 620 6 HOH A 702   O   112.6  78.8 139.2  68.8 103.2
REMARK 620 7 HOH A 706   O    81.7 106.9 156.1 100.3  82.9  45.2
REMARK 620 N                    1     2     3     4     5     6
DBREF  8QRJ A    1   293  UNP    G9BY57   PETH_UNKP        1    293
SEQADV 8QRJ GLY A  127  UNP  G9BY57    TYR   127 CONFLICT
SEQADV 8QRJ TYR A  218  UNP  G9BY57    HIS   218 ENGINEERED MUTATION
SEQADV 8QRJ CYS A  238  UNP  G9BY57    ASP   238 CONFLICT
SEQADV 8QRJ ILE A  243  UNP  G9BY57    PHE   243 CONFLICT
SEQADV 8QRJ CYS A  283  UNP  G9BY57    SER   283 CONFLICT
SEQRES   1 A  293  MET ASP GLY VAL LEU TRP ARG VAL ARG THR ALA ALA LEU
SEQRES   2 A  293  MET ALA ALA LEU LEU ALA LEU ALA ALA TRP ALA LEU VAL
SEQRES   3 A  293  TRP ALA SER PRO SER VAL GLU ALA GLN SER ASN PRO TYR
SEQRES   4 A  293  GLN ARG GLY PRO ASN PRO THR ARG SER ALA LEU THR ALA
SEQRES   5 A  293  ASP GLY PRO PHE SER VAL ALA THR TYR THR VAL SER ARG
SEQRES   6 A  293  LEU SER VAL SER GLY PHE GLY GLY GLY VAL ILE TYR TYR
SEQRES   7 A  293  PRO THR GLY THR SER LEU THR PHE GLY GLY ILE ALA MET
SEQRES   8 A  293  SER PRO GLY TYR THR ALA ASP ALA SER SER LEU ALA TRP
SEQRES   9 A  293  LEU GLY ARG ARG LEU ALA SER HIS GLY PHE VAL VAL LEU
SEQRES  10 A  293  VAL ILE ASN THR ASN SER ARG PHE ASP GLY PRO ASP SER
SEQRES  11 A  293  ARG ALA SER GLN LEU SER ALA ALA LEU ASN TYR LEU ARG
SEQRES  12 A  293  THR SER SER PRO SER ALA VAL ARG ALA ARG LEU ASP ALA
SEQRES  13 A  293  ASN ARG LEU ALA VAL ALA GLY HIS SER MET GLY GLY GLY
SEQRES  14 A  293  GLY THR LEU ARG ILE ALA GLU GLN ASN PRO SER LEU LYS
SEQRES  15 A  293  ALA ALA VAL PRO LEU THR PRO TRP HIS THR ASP LYS THR
SEQRES  16 A  293  PHE ASN THR SER VAL PRO VAL LEU ILE VAL GLY ALA GLU
SEQRES  17 A  293  ALA ASP THR VAL ALA PRO VAL SER GLN TYR ALA ILE PRO
SEQRES  18 A  293  PHE TYR GLN ASN LEU PRO SER THR THR PRO LYS VAL TYR
SEQRES  19 A  293  VAL GLU LEU CYS ASN ALA SER HIS ILE ALA PRO ASN SER
SEQRES  20 A  293  ASN ASN ALA ALA ILE SER VAL TYR THR ILE SER TRP MET
SEQRES  21 A  293  LYS LEU TRP VAL ASP ASN ASP THR ARG TYR ARG GLN PHE
SEQRES  22 A  293  LEU CYS ASN VAL ASN ASP PRO ALA LEU CYS ASP PHE ARG
SEQRES  23 A  293  THR ASN ASN ARG HIS CYS GLN
HET    EDO  A 301       4
HET    P33  A 302      22
HET     CA  A 303       1
HETNAM     EDO 1,2-ETHANEDIOL
HETNAM     P33 3,6,9,12,15,18-HEXAOXAICOSANE-1,20-DIOL
HETNAM      CA CALCIUM ION
HETSYN     EDO ETHYLENE GLYCOL
HETSYN     P33 HEPTAETHYLENE GLYCOL; PEG330
FORMUL   2  EDO    C2 H6 O2
FORMUL   3  P33    C14 H30 O8
FORMUL   4   CA    CA 2+
FORMUL   5  HOH   *442(H2 O)
HELIX    1 AA1 ARG A   47  ALA A   52  5                                   6
HELIX    2 AA2 SER A   64  VAL A   68  5                                   5
HELIX    3 AA3 ASP A   98  SER A  101  5                                   4
HELIX    4 AA4 LEU A  102  HIS A  112  1                                  11
HELIX    5 AA5 GLY A  127  SER A  145  1                                  19
HELIX    6 AA6 PRO A  147  ARG A  153  1                                   7
HELIX    7 AA7 SER A  165  ASN A  178  1                                  14
HELIX    8 AA8 TYR A  218  LEU A  226  1                                   9
HELIX    9 AA9 ILE A  243  SER A  247  5                                   5
HELIX   10 AB1 ASN A  249  ASP A  265  1                                  17
HELIX   11 AB2 ASP A  267  LEU A  274  5                                   8
SHEET    1 AA1 6 SER A  57  VAL A  63  0
SHEET    2 AA1 6 GLY A  74  THR A  80 -1  O  ILE A  76   N  TYR A  61
SHEET    3 AA1 6 VAL A 115  ILE A 119 -1  O  VAL A 116   N  TYR A  77
SHEET    4 AA1 6 PHE A  86  SER A  92  1  N  ILE A  89   O  LEU A 117
SHEET    5 AA1 6 LEU A 154  HIS A 164  1  O  ASP A 155   N  PHE A  86
SHEET    6 AA1 6 ALA A 184  LEU A 187  1  O  LEU A 187   N  GLY A 163
SHEET    1 AA2 3 VAL A 202  ALA A 207  0
SHEET    2 AA2 3 LYS A 232  LEU A 237  1  O  VAL A 235   N  GLY A 206
SHEET    3 AA2 3 LEU A 282  THR A 287 -1  O  CYS A 283   N  GLU A 236
SSBOND   1 CYS A  238    CYS A  283                          1555   1555  2.07
SSBOND   2 CYS A  275    CYS A  292                          1555   1555  2.05
LINK         O   ARG A  65                CA    CA A 303     1555   1555  2.65
LINK         O   VAL A  68                CA    CA A 303     1555   1555  2.69
LINK         O   PHE A  71                CA    CA A 303     1555   1555  2.77
LINK        CA    CA A 303                 O   HOH A 516     1555   1555  2.92
LINK        CA    CA A 303                 O   HOH A 542     1555   1555  2.84
LINK        CA    CA A 303                 O   HOH A 702     1555   1555  2.80
LINK        CA    CA A 303                 O   HOH A 706     1555   1555  2.82
CRYST1   81.900   94.510   80.950  90.00  90.00  90.00 C 2 2 21      8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.012210  0.000000  0.000000        0.00000
SCALE2      0.000000  0.010581  0.000000        0.00000
SCALE3      0.000000  0.000000  0.012353        0.00000
TER    1967      CYS A 292
MASTER      480    0    3   11    9    0    0    6 2410    1   39   23
END