longtext: 8tru-pdb

content
HEADER    HYDROLASE                               10-AUG-23   8TRU
TITLE     CRYSTAL STRUCTURE OF A CE15 FROM FIBROBACTER SUCCINOGENES SUBSP.
TITLE    2 SUCCINOGENES S85
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: GLUCURONYL ESTERASE;
COMPND   3 CHAIN: A;
COMPND   4 FRAGMENT: RESIDUES 153-536;
COMPND   5 ENGINEERED: YES;
COMPND   6 OTHER_DETAILS: C-TERMINAL HIS-TAGGED CONSTRUCT
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: FIBROBACTER SUCCINOGENES SUBSP. SUCCINOGENES;
SOURCE   3 ORGANISM_TAXID: 59374;
SOURCE   4 STRAIN: ATCC 19169 / S85;
SOURCE   5 GENE: FSU_2898;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 469008
KEYWDS    CARBOHYDRATE, ESTERASE, GLUCURONYL ESTERASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    R.J.GRUNINGER,D.R.JONES
REVDAT   1   28-AUG-24 8TRU    0
JRNL        AUTH   R.J.GRUNINGER,M.KEVORKOVA,K.E.LOW,D.R.JONES,L.WORRALL,
JRNL        AUTH 2 T.A.MCALLISTER,D.W.ABBOTT
JRNL        TITL   STRUCTURAL, BIOCHEMICAL, AND PHYLOGENETIC ANALYSIS OF
JRNL        TITL 2 BACTERIAL AND FUNGAL CARBOHYDRATE ESTERASE FAMILY 15
JRNL        TITL 3 GLUCURONOYL ESTERASES IN THE RUMEN.
JRNL        REF    PROTEIN J.                                 2024
JRNL        REFN                   ISSN 1572-3887
JRNL        PMID   39153129
JRNL        DOI    10.1007/S10930-024-10221-0
REMARK   2
REMARK   2 RESOLUTION.    1.90 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.15.2_3472
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.90
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.94
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.340
REMARK   3   COMPLETENESS FOR RANGE        (%) : 100.0
REMARK   3   NUMBER OF REFLECTIONS             : 30274
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.210
REMARK   3   R VALUE            (WORKING SET) : 0.207
REMARK   3   FREE R VALUE                     : 0.256
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.990
REMARK   3   FREE R VALUE TEST SET COUNT      : 1512
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 19.9370 -  4.2135    1.00     2855   151  0.1588 0.1826
REMARK   3     2  4.2135 -  3.3495    1.00     2684   141  0.1653 0.2162
REMARK   3     3  3.3495 -  2.9276    1.00     2638   139  0.2077 0.2844
REMARK   3     4  2.9276 -  2.6606    1.00     2614   137  0.2284 0.2590
REMARK   3     5  2.6606 -  2.4703    1.00     2588   136  0.2337 0.2829
REMARK   3     6  2.4703 -  2.3249    1.00     2593   136  0.2269 0.2889
REMARK   3     7  2.3249 -  2.2086    1.00     2546   134  0.2359 0.3230
REMARK   3     8  2.2086 -  2.1126    1.00     2580   136  0.2490 0.3179
REMARK   3     9  2.1126 -  2.0313    1.00     2553   134  0.2761 0.3115
REMARK   3    10  2.0313 -  1.9613    1.00     2574   136  0.3069 0.3612
REMARK   3    11  1.9613 -  1.9000    1.00     2537   132  0.3441 0.3661
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.240
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 31.570
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 20.40
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 37.14
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.006           2919
REMARK   3   ANGLE     :  0.766           3942
REMARK   3   CHIRALITY :  0.049            413
REMARK   3   PLANARITY :  0.005            520
REMARK   3   DIHEDRAL  :  4.438           2378
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 6
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 155 THROUGH 191 )
REMARK   3    ORIGIN FOR THE GROUP (A): -29.9885  24.6632  19.7542
REMARK   3    T TENSOR
REMARK   3      T11:   0.2754 T22:   0.2643
REMARK   3      T33:   0.4407 T12:  -0.0113
REMARK   3      T13:   0.0256 T23:  -0.0086
REMARK   3    L TENSOR
REMARK   3      L11:   1.8547 L22:   1.3366
REMARK   3      L33:   0.3484 L12:   0.1622
REMARK   3      L13:   0.2214 L23:   0.0534
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1117 S12:   0.0900 S13:   0.2724
REMARK   3      S21:  -0.0593 S22:   0.0722 S23:   0.0169
REMARK   3      S31:  -0.1633 S32:   0.0176 S33:   0.0454
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 192 THROUGH 238 )
REMARK   3    ORIGIN FOR THE GROUP (A): -14.1727  18.1653  16.0936
REMARK   3    T TENSOR
REMARK   3      T11:   0.2513 T22:   0.3083
REMARK   3      T33:   0.4702 T12:  -0.0686
REMARK   3      T13:   0.0920 T23:  -0.0922
REMARK   3    L TENSOR
REMARK   3      L11:   1.3361 L22:   0.4293
REMARK   3      L33:   0.5007 L12:   0.4461
REMARK   3      L13:   0.3517 L23:  -0.1149
REMARK   3    S TENSOR
REMARK   3      S11:  -0.2317 S12:   0.3186 S13:  -0.0053
REMARK   3      S21:  -0.1443 S22:   0.1938 S23:  -0.1863
REMARK   3      S31:   0.0277 S32:   0.0357 S33:   0.0121
REMARK   3   TLS GROUP : 3
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 239 THROUGH 274 )
REMARK   3    ORIGIN FOR THE GROUP (A):  -9.5435   3.6106   7.8839
REMARK   3    T TENSOR
REMARK   3      T11:   0.3830 T22:   0.5020
REMARK   3      T33:   0.6983 T12:  -0.1458
REMARK   3      T13:   0.1995 T23:  -0.2092
REMARK   3    L TENSOR
REMARK   3      L11:   0.1675 L22:   0.4476
REMARK   3      L33:   0.0580 L12:   0.2524
REMARK   3      L13:  -0.0717 L23:  -0.1579
REMARK   3    S TENSOR
REMARK   3      S11:  -0.3196 S12:   0.4020 S13:  -0.2868
REMARK   3      S21:  -0.1137 S22:   0.1586 S23:  -0.2199
REMARK   3      S31:   0.1617 S32:  -0.0483 S33:   0.0482
REMARK   3   TLS GROUP : 4
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 275 THROUGH 486 )
REMARK   3    ORIGIN FOR THE GROUP (A): -23.6809   6.5445  18.6242
REMARK   3    T TENSOR
REMARK   3      T11:   0.2434 T22:   0.3044
REMARK   3      T33:   0.4295 T12:  -0.0260
REMARK   3      T13:   0.0706 T23:  -0.0695
REMARK   3    L TENSOR
REMARK   3      L11:   0.9762 L22:   1.0260
REMARK   3      L33:   0.3027 L12:   0.4091
REMARK   3      L13:   0.0760 L23:   0.0445
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1229 S12:   0.1435 S13:  -0.1967
REMARK   3      S21:  -0.1025 S22:   0.1283 S23:  -0.1028
REMARK   3      S31:   0.0027 S32:   0.0586 S33:   0.0058
REMARK   3   TLS GROUP : 5
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 487 THROUGH 514 )
REMARK   3    ORIGIN FOR THE GROUP (A): -30.4503   0.8293   2.1076
REMARK   3    T TENSOR
REMARK   3      T11:   0.4174 T22:   0.4626
REMARK   3      T33:   0.4063 T12:  -0.1233
REMARK   3      T13:   0.0984 T23:  -0.1688
REMARK   3    L TENSOR
REMARK   3      L11:   0.8331 L22:   0.1665
REMARK   3      L33:   0.0986 L12:  -0.3393
REMARK   3      L13:  -0.1957 L23:   0.1116
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1904 S12:   0.2749 S13:  -0.3712
REMARK   3      S21:  -0.2806 S22:   0.1887 S23:   0.0701
REMARK   3      S31:   0.0839 S32:  -0.1008 S33:  -0.0046
REMARK   3   TLS GROUP : 6
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 515 THROUGH 538 )
REMARK   3    ORIGIN FOR THE GROUP (A): -42.8786  11.0316  17.1893
REMARK   3    T TENSOR
REMARK   3      T11:   0.2288 T22:   0.3329
REMARK   3      T33:   0.4352 T12:  -0.0206
REMARK   3      T13:   0.0433 T23:  -0.0014
REMARK   3    L TENSOR
REMARK   3      L11:   1.0548 L22:   2.4147
REMARK   3      L33:   0.4563 L12:  -0.3149
REMARK   3      L13:   0.2880 L23:   0.2350
REMARK   3    S TENSOR
REMARK   3      S11:  -0.2247 S12:   0.2474 S13:   0.1280
REMARK   3      S21:   0.1423 S22:   0.1710 S23:   0.4339
REMARK   3      S31:  -0.0006 S32:  -0.1328 S33:   0.0816
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 8TRU COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 22-AUG-23.
REMARK 100 THE DEPOSITION ID IS D_1000276536.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 10-FEB-19
REMARK 200  TEMPERATURE           (KELVIN) : 80
REMARK 200  PH                             : 5.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : CLSI
REMARK 200  BEAMLINE                       : 08ID-1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0334
REMARK 200  MONOCHROMATOR                  : DOUBLE CRYSTAL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XDS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 30281
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.900
REMARK 200  RESOLUTION RANGE LOW       (A) : 19.940
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 100.0
REMARK 200  DATA REDUNDANCY                : 12.50
REMARK 200  R MERGE                    (I) : 0.17200
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 10.6000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.90
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.97
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 12.90
REMARK 200  R MERGE FOR SHELL          (I) : 1.13400
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.200
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHENIX
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 39.73
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.21
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 20% PEG6000, 100 MM SODIUM ACETATE, PH
REMARK 280  5.0, 10 MM ZINC CHLORIDE, VAPOR DIFFUSION, SITTING DROP,
REMARK 280  TEMPERATURE 298K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 41 21 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z+1/2
REMARK 290       3555   -Y+1/2,X+1/2,Z+1/4
REMARK 290       4555   Y+1/2,-X+1/2,Z+3/4
REMARK 290       5555   -X+1/2,Y+1/2,-Z+1/4
REMARK 290       6555   X+1/2,-Y+1/2,-Z+3/4
REMARK 290       7555   Y,X,-Z
REMARK 290       8555   -Y,-X,-Z+1/2
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      111.08000
REMARK 290   SMTRY1   3  0.000000 -1.000000  0.000000       28.64000
REMARK 290   SMTRY2   3  1.000000  0.000000  0.000000       28.64000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       55.54000
REMARK 290   SMTRY1   4  0.000000  1.000000  0.000000       28.64000
REMARK 290   SMTRY2   4 -1.000000  0.000000  0.000000       28.64000
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000      166.62000
REMARK 290   SMTRY1   5 -1.000000  0.000000  0.000000       28.64000
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       28.64000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       55.54000
REMARK 290   SMTRY1   6  1.000000  0.000000  0.000000       28.64000
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       28.64000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000      166.62000
REMARK 290   SMTRY1   7  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2   7  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2   8 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000      111.08000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH A 904  LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A   152
REMARK 465     ASN A   153
REMARK 465     GLU A   154
REMARK 465     THR A   252
REMARK 465     LYS A   253
REMARK 465     GLY A   266
REMARK 465     MET A   267
REMARK 465     MET A   268
REMARK 465     GLY A   269
REMARK 465     ALA A   539
REMARK 465     PHE A   540
REMARK 465     LEU A   541
REMARK 465     TYR A   542
REMARK 465     LYS A   543
REMARK 465     PRO A   544
REMARK 465     ALA A   545
REMARK 465     PHE A   546
REMARK 465     LEU A   547
REMARK 465     TYR A   548
REMARK 465     LYS A   549
REMARK 465     VAL A   550
REMARK 465     VAL A   551
REMARK 465     ILE A   552
REMARK 465     MET A   553
REMARK 465     HIS A   554
REMARK 465     HIS A   555
REMARK 465     HIS A   556
REMARK 465     HIS A   557
REMARK 465     HIS A   558
REMARK 465     HIS A   559
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     LYS A 187    CG   CD   CE   NZ
REMARK 470     LYS A 246    CG   CD   CE   NZ
REMARK 470     ASP A 254    CG   OD1  OD2
REMARK 470     LYS A 255    CG   CD   CE   NZ
REMARK 470     LYS A 338    CG   CD   CE   NZ
REMARK 470     GLU A 470    CG   CD   OE1  OE2
REMARK 470     LYS A 523    CD   CE   NZ
REMARK 470     SER A 534    OG
REMARK 470     LYS A 536    CG   CD   CE   NZ
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    MET A 210      -54.19   -124.12
REMARK 500    ASP A 236      147.07   -172.79
REMARK 500    ASN A 249      173.17     67.98
REMARK 500    ALA A 250     -149.73   -163.29
REMARK 500    ALA A 294       90.84   -160.40
REMARK 500    ASN A 307       39.42   -153.37
REMARK 500    SER A 346     -124.48     62.74
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 945        DISTANCE =  6.44 ANGSTROMS
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              ZN A 602  ZN
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HIS A 159   ND1
REMARK 620 2 GLU A 360   OE1  97.0
REMARK 620 3 GLU A 360   OE2 103.0  56.4
REMARK 620 4 HOH A 718   O   111.3 132.0  79.4
REMARK 620 5 HOH A 864   O   118.4  81.7 124.1 114.1
REMARK 620 N                    1     2     3     4
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              ZN A 601  ZN
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 GLY A 488   O
REMARK 620 2 ASP A 491   OD1  86.5
REMARK 620 3 ASP A 491   OD2 143.1  58.6
REMARK 620 4 GLU A 492   OE2 112.6 109.1  91.6
REMARK 620 5 HOH A 778   O    71.6 158.1 142.3  81.4
REMARK 620 N                    1     2     3     4
DBREF  8TRU A  153   536  UNP    C9RKY6   C9RKY6_FIBSS   153    536
SEQADV 8TRU MET A  152  UNP  C9RKY6              INITIATING METHIONINE
SEQADV 8TRU ASP A  537  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU PRO A  538  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU ALA A  539  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU PHE A  540  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU LEU A  541  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU TYR A  542  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU LYS A  543  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU PRO A  544  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU ALA A  545  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU PHE A  546  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU LEU A  547  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU TYR A  548  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU LYS A  549  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU VAL A  550  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU VAL A  551  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU ILE A  552  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU MET A  553  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU HIS A  554  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU HIS A  555  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU HIS A  556  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU HIS A  557  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU HIS A  558  UNP  C9RKY6              EXPRESSION TAG
SEQADV 8TRU HIS A  559  UNP  C9RKY6              EXPRESSION TAG
SEQRES   1 A  408  MET ASN GLU PHE VAL GLU ASP HIS ARG SER GLU CYS GLN
SEQRES   2 A  408  ILE GLY ASN ILE PRO SER SER VAL ASN ASN ALA LYS LEU
SEQRES   3 A  408  PRO ASP PRO PHE MET GLY LEU ASP GLY LYS ARG ILE SER
SEQRES   4 A  408  SER LYS ALA ASP TRP LYS CYS ARG ARG GLU GLU ILE GLY
SEQRES   5 A  408  ALA MET TYR GLU LYS LEU MET PHE GLY THR LYS PRO ARG
SEQRES   6 A  408  ASN PRO GLU LYS VAL GLU GLY SER TYR SER GLY GLY LYS
SEQRES   7 A  408  LEU THR ILE LYS VAL THR ASP LYS GLY LYS SER GLY SER
SEQRES   8 A  408  PHE SER VAL LYS ILE SER ASN ALA GLY THR LYS ASP LYS
SEQRES   9 A  408  PRO LYS PRO ALA MET ILE GLY PHE GLY GLY GLY MET MET
SEQRES  10 A  408  GLY GLY CYS GLY SER LEU GLY ASN ALA THR ASN GLY LEU
SEQRES  11 A  408  ASP ILE ALA GLN ILE THR PHE ASN PRO ASP ASP VAL ALA
SEQRES  12 A  408  PRO GLU SER GLY GLY GLY MET PHE PHE GLN LEU TYR ASN
SEQRES  13 A  408  GLN GLY GLN GLY THR ILE ILE ALA TRP ALA TRP GLY VAL
SEQRES  14 A  408  SER ARG ILE ILE ASP ALA LEU GLU LYS THR PRO GLU ALA
SEQRES  15 A  408  GLY ILE ASP VAL LYS HIS LEU ALA MET THR GLY CYS SER
SEQRES  16 A  408  ARG TRP GLY LYS GLY THR LEU ALA VAL GLY ALA PHE ASP
SEQRES  17 A  408  GLU ARG ILE ALA LEU THR ILE PRO GLN GLU SER GLY SER
SEQRES  18 A  408  GLY GLY ALA SER LEU TRP ARG VAL GLY ALA GLN VAL ASN
SEQRES  19 A  408  LYS GLN LYS GLY LYS GLN PHE VAL GLN GLY LEU SER SER
SEQRES  20 A  408  ALA GLY THR GLU GLY LYS TRP MET ILE SER SER PHE LYS
SEQRES  21 A  408  ASN TYR ASP GLY LYS GLU ASN THR LEU PRO PHE ASP GLN
SEQRES  22 A  408  HIS MET LEU VAL ALA MET VAL ALA PRO ARG ALA LEU LEU
SEQRES  23 A  408  ILE LEU ASP ASN ALA GLY GLN GLU TRP LEU GLY GLU VAL
SEQRES  24 A  408  PRO SER ASN TYR CYS GLY GLN ALA SER LYS GLU VAL TYR
SEQRES  25 A  408  ASP ALA LEU GLY ALA THR GLU ASN TYR THR TYR SER GLN
SEQRES  26 A  408  GLU GLY GLY HIS GLY HIS CYS GLN LEU PRO ASN GLY GLN
SEQRES  27 A  408  PHE ASP GLU VAL LYS ASP PHE MET ASN LYS PHE LEU LEU
SEQRES  28 A  408  GLY LYS ASP ALA LYS THR GLY LYS ILE ASP TYR THR LYS
SEQRES  29 A  408  ASN THR GLN THR ILE ASN PHE LYS LYS SER GLU TRP ILE
SEQRES  30 A  408  ASP TRP GLU THR PRO SER LEU LYS ASP PRO ALA PHE LEU
SEQRES  31 A  408  TYR LYS PRO ALA PHE LEU TYR LYS VAL VAL ILE MET HIS
SEQRES  32 A  408  HIS HIS HIS HIS HIS
HET     ZN  A 601       1
HET     ZN  A 602       1
HETNAM      ZN ZINC ION
FORMUL   2   ZN    2(ZN 2+)
FORMUL   4  HOH   *245(H2 O)
HELIX    1 AA1 HIS A  159  CYS A  163  5                                   5
HELIX    2 AA2 SER A  191  ALA A  193  5                                   3
HELIX    3 AA3 ASP A  194  MET A  210  1                                  17
HELIX    4 AA4 LEU A  274  ASN A  279  5                                   6
HELIX    5 AA5 ASN A  289  VAL A  293  5                                   5
HELIX    6 AA6 MET A  301  TYR A  306  1                                   6
HELIX    7 AA7 GLY A  311  LYS A  329  1                                  19
HELIX    8 AA8 THR A  330  ALA A  333  5                                   4
HELIX    9 AA9 SER A  346  ASP A  359  1                                  14
HELIX   10 AB1 LEU A  377  GLY A  389  1                                  13
HELIX   11 AB2 GLY A  395  GLY A  400  1                                   6
HELIX   12 AB3 THR A  401  TRP A  405  5                                   5
HELIX   13 AB4 SER A  409  LEU A  420  5                                  12
HELIX   14 AB5 GLN A  424  ALA A  432  1                                   9
HELIX   15 AB6 GLY A  448  LEU A  466  1                                  19
HELIX   16 AB7 ALA A  468  GLU A  470  5                                   3
HELIX   17 AB8 PRO A  486  GLY A  488  5                                   3
HELIX   18 AB9 GLN A  489  LEU A  501  1                                  13
HELIX   19 AC1 LYS A  523  ILE A  528  1                                   6
SHEET    1 AA110 LYS A 220  SER A 226  0
SHEET    2 AA110 LYS A 229  ASP A 236 -1  O  LYS A 233   N  GLU A 222
SHEET    3 AA110 LYS A 239  SER A 248 -1  O  PHE A 243   N  ILE A 232
SHEET    4 AA110 ALA A 284  PHE A 288 -1  O  GLN A 285   N  SER A 248
SHEET    5 AA110 LYS A 257  PHE A 263  1  N  PRO A 258   O  ALA A 284
SHEET    6 AA110 ILE A 335  CYS A 345  1  O  ALA A 341   N  ILE A 261
SHEET    7 AA110 LEU A 364  GLN A 368  1  O  ILE A 366   N  MET A 342
SHEET    8 AA110 ALA A 435  ASP A 440  1  O  LEU A 437   N  THR A 365
SHEET    9 AA110 TYR A 472  GLU A 477  1  O  SER A 475   N  ILE A 438
SHEET   10 AA110 LYS A 510  THR A 514  1  O  ASP A 512   N  GLN A 476
SSBOND   1 CYS A  163    CYS A  197                          1555   1555  2.02
SSBOND   2 CYS A  345    CYS A  483                          1555   1555  2.04
LINK         ND1 HIS A 159                ZN    ZN A 602     1555   1555  2.05
LINK         OE1 GLU A 360                ZN    ZN A 602     1555   1555  2.52
LINK         OE2 GLU A 360                ZN    ZN A 602     1555   1555  1.99
LINK         O   GLY A 488                ZN    ZN A 601     1555   1555  2.03
LINK         OD1 ASP A 491                ZN    ZN A 601     1555   1555  2.06
LINK         OD2 ASP A 491                ZN    ZN A 601     1555   1555  2.39
LINK         OE2 GLU A 492                ZN    ZN A 601     1555   1555  2.14
LINK        ZN    ZN A 601                 O   HOH A 778     1555   1555  2.62
LINK        ZN    ZN A 602                 O   HOH A 718     1555   1555  1.88
LINK        ZN    ZN A 602                 O   HOH A 864     1555   1555  2.21
CISPEP   1 ALA A  432    PRO A  433          0         2.41
CRYST1   57.280   57.280  222.160  90.00  90.00  90.00 P 41 21 2     8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.017458  0.000000  0.000000        0.00000
SCALE2      0.000000  0.017458  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004501        0.00000
TER    2857      PRO A 538
MASTER      425    0    2   19   10    0    0    6 3098    1   18   32
END