longtext: 8xiz-pdb

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HEADER    HYDROLASE                               20-DEC-23   8XIZ
TITLE     CRYSTAL STRUCTURE OF AN EPOXIDE HYDROLASE MUTANT A250IC/L344V FROM
TITLE    2 ASPERGILLUS USAMII E001 AT 2.17 ANGSTROMS RESOLUTION
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: MICROSOMAL EPOXIDE HYDDROLASE;
COMPND   3 CHAIN: A, B;
COMPND   4 EC: 3.3.2.9;
COMPND   5 ENGINEERED: YES;
COMPND   6 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: ASPERGILLUS USAMII;
SOURCE   3 ORGANISM_TAXID: 186680;
SOURCE   4 GENE: EH2;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI BL21(DE3);
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   8 EXPRESSION_SYSTEM_PLASMID: PET28A
KEYWDS    ALPHA AND BETA PROTEINS, ALPHA/BETA-HYDROLASES, STYRENE OXIDE,
KEYWDS   2 EPOXIDE HYDROLASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    B.C.HU,Z.Y.LU,C.D.TANG,D.HU
REVDAT   1   07-FEB-24 8XIZ    0
JRNL        AUTH   D.HU,Z.Y.LU
JRNL        TITL   DIRECTED EVOLUTION OF AN EPOXIDE HYDROLASE AND ITS
JRNL        TITL 2 STRUCTURAL MECHANISM FOR THE ENANTIOSELECTIVITY IMPROVEMENT
JRNL        TITL 3 TOWARD CHIRAL ORTHO-FLUOROSTYRENE OXIDE
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    2.18 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (1.13_2998: ???)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.18
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 44.60
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.360
REMARK   3   COMPLETENESS FOR RANGE        (%) : 98.1
REMARK   3   NUMBER OF REFLECTIONS             : 40335
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.165
REMARK   3   R VALUE            (WORKING SET) : 0.162
REMARK   3   FREE R VALUE                     : 0.222
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.330
REMARK   3   FREE R VALUE TEST SET COUNT      : 2148
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 44.6010 -  5.3614    0.98     2655   153  0.1584 0.1828
REMARK   3     2  5.3614 -  4.2565    0.99     2601   146  0.1360 0.1827
REMARK   3     3  4.2565 -  3.7188    0.99     2626   134  0.1426 0.2128
REMARK   3     4  3.7188 -  3.3789    0.99     2580   128  0.1565 0.2161
REMARK   3     5  3.3789 -  3.1368    0.99     2615   119  0.1752 0.2025
REMARK   3     6  3.1368 -  2.9519    0.99     2561   133  0.1814 0.2512
REMARK   3     7  2.9519 -  2.8041    0.99     2581   139  0.1745 0.2371
REMARK   3     8  2.8041 -  2.6820    0.99     2520   154  0.1729 0.2386
REMARK   3     9  2.6820 -  2.5788    0.99     2544   146  0.1756 0.2654
REMARK   3    10  2.5788 -  2.4898    0.98     2561   114  0.1801 0.2727
REMARK   3    11  2.4898 -  2.4120    0.98     2514   156  0.1733 0.2631
REMARK   3    12  2.4120 -  2.3430    0.98     2517   145  0.1771 0.2372
REMARK   3    13  2.3430 -  2.2814    0.98     2517   170  0.1737 0.2471
REMARK   3    14  2.2814 -  2.2257    0.98     2501   147  0.1733 0.2440
REMARK   3    15  2.2257 -  2.1751    0.90     2294   164  0.1745 0.2609
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.220
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 20.850
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 26.50
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.007           6502
REMARK   3   ANGLE     :  0.814           8886
REMARK   3   CHIRALITY :  0.051            948
REMARK   3   PLANARITY :  0.006           1146
REMARK   3   DIHEDRAL  :  6.558           3792
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 8XIZ COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 21-DEC-23.
REMARK 100 THE DEPOSITION ID IS D_1300040916.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 20-JAN-21
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SSRF
REMARK 200  BEAMLINE                       : BL19U1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9791
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS3 6M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-3000
REMARK 200  DATA SCALING SOFTWARE          : HKL-3000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 40335
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.170
REMARK 200  RESOLUTION RANGE LOW       (A) : 47.890
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 97.8
REMARK 200  DATA REDUNDANCY                : 6.700
REMARK 200  R MERGE                    (I) : 0.07000
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 6.5500
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.17
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.18
REMARK 200  COMPLETENESS FOR SHELL     (%) : 97.8
REMARK 200  DATA REDUNDANCY IN SHELL       : 6.70
REMARK 200  R MERGE FOR SHELL          (I) : 0.07000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: AMPLE
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: TETRAGONAL CRYSTAL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 39.02
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.21
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1 M SODIUM CITRATE TRIBASIC
REMARK 280  DIHYDRATE,PH5.5,18%(W/V) POLYETHYLENE GLYCOL 3,350, VAPOR
REMARK 280  DIFFUSION, SITTING DROP, TEMPERATURE 291K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       25.64350
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A   -35
REMARK 465     GLY A   -34
REMARK 465     SER A   -33
REMARK 465     SER A   -32
REMARK 465     HIS A   -31
REMARK 465     HIS A   -30
REMARK 465     HIS A   -29
REMARK 465     HIS A   -28
REMARK 465     HIS A   -27
REMARK 465     HIS A   -26
REMARK 465     SER A   -25
REMARK 465     SER A   -24
REMARK 465     GLY A   -23
REMARK 465     LEU A   -22
REMARK 465     VAL A   -21
REMARK 465     PRO A   -20
REMARK 465     ARG A   -19
REMARK 465     GLY A   -18
REMARK 465     SER A   -17
REMARK 465     HIS A   -16
REMARK 465     MET A   -15
REMARK 465     ALA A   -14
REMARK 465     SER A   -13
REMARK 465     MET A   -12
REMARK 465     THR A   -11
REMARK 465     GLY A   -10
REMARK 465     GLY A    -9
REMARK 465     GLN A    -8
REMARK 465     GLN A    -7
REMARK 465     MET A    -6
REMARK 465     GLY A    -5
REMARK 465     ARG A    -4
REMARK 465     GLY A    -3
REMARK 465     SER A    -2
REMARK 465     GLU A    -1
REMARK 465     PHE A     0
REMARK 465     PRO A   224
REMARK 465     ASP A   225
REMARK 465     THR A   226
REMARK 465     ALA A   227
REMARK 465     ARG A   394
REMARK 465     LYS A   395
REMARK 465     MET B   -35
REMARK 465     GLY B   -34
REMARK 465     SER B   -33
REMARK 465     SER B   -32
REMARK 465     HIS B   -31
REMARK 465     HIS B   -30
REMARK 465     HIS B   -29
REMARK 465     HIS B   -28
REMARK 465     HIS B   -27
REMARK 465     HIS B   -26
REMARK 465     SER B   -25
REMARK 465     SER B   -24
REMARK 465     GLY B   -23
REMARK 465     LEU B   -22
REMARK 465     VAL B   -21
REMARK 465     PRO B   -20
REMARK 465     ARG B   -19
REMARK 465     GLY B   -18
REMARK 465     SER B   -17
REMARK 465     HIS B   -16
REMARK 465     MET B   -15
REMARK 465     ALA B   -14
REMARK 465     SER B   -13
REMARK 465     MET B   -12
REMARK 465     THR B   -11
REMARK 465     GLY B   -10
REMARK 465     GLY B    -9
REMARK 465     GLN B    -8
REMARK 465     GLN B    -7
REMARK 465     MET B    -6
REMARK 465     GLY B    -5
REMARK 465     ARG B    -4
REMARK 465     GLY B    -3
REMARK 465     SER B    -2
REMARK 465     GLU B    -1
REMARK 465     PHE B     0
REMARK 465     MET B     1
REMARK 465     ALA B     2
REMARK 465     PRO B   224
REMARK 465     THR B   226
REMARK 465     ALA B   227
REMARK 465     ARG B   394
REMARK 465     LYS B   395
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    PRO A 117       47.39    -93.82
REMARK 500    THR A 152     -116.33     45.36
REMARK 500    ASN A 164     -166.07   -129.29
REMARK 500    ASP A 191     -135.09     51.39
REMARK 500    PRO B 117       48.42    -93.43
REMARK 500    THR B 152     -114.30     42.90
REMARK 500    ASP B 191     -134.18     51.08
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 634        DISTANCE =  6.22 ANGSTROMS
REMARK 525    HOH A 635        DISTANCE =  6.90 ANGSTROMS
REMARK 525    HOH A 636        DISTANCE =  8.63 ANGSTROMS
REMARK 525    HOH B 626        DISTANCE =  6.01 ANGSTROMS
REMARK 525    HOH B 627        DISTANCE =  6.87 ANGSTROMS
REMARK 525    HOH B 628        DISTANCE =  7.57 ANGSTROMS
REMARK 525    HOH B 629        DISTANCE =  7.60 ANGSTROMS
DBREF  8XIZ A    1   395  UNP    T2B4K5   T2B4K5_ASPUS     1    395
DBREF  8XIZ B    1   395  UNP    T2B4K5   T2B4K5_ASPUS     1    395
SEQADV 8XIZ MET A  -35  UNP  T2B4K5              INITIATING METHIONINE
SEQADV 8XIZ GLY A  -34  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER A  -33  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER A  -32  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS A  -31  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS A  -30  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS A  -29  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS A  -28  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS A  -27  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS A  -26  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER A  -25  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER A  -24  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY A  -23  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ LEU A  -22  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ VAL A  -21  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ PRO A  -20  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ ARG A  -19  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY A  -18  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER A  -17  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS A  -16  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ MET A  -15  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ ALA A  -14  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER A  -13  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ MET A  -12  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ THR A  -11  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY A  -10  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY A   -9  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLN A   -8  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLN A   -7  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ MET A   -6  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY A   -5  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ ARG A   -4  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY A   -3  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER A   -2  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLU A   -1  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ PHE A    0  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ ILE A  250  UNP  T2B4K5    ALA   250 ENGINEERED MUTATION
SEQADV 8XIZ VAL A  344  UNP  T2B4K5    LEU   344 ENGINEERED MUTATION
SEQADV 8XIZ MET B  -35  UNP  T2B4K5              INITIATING METHIONINE
SEQADV 8XIZ GLY B  -34  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER B  -33  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER B  -32  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS B  -31  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS B  -30  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS B  -29  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS B  -28  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS B  -27  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS B  -26  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER B  -25  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER B  -24  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY B  -23  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ LEU B  -22  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ VAL B  -21  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ PRO B  -20  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ ARG B  -19  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY B  -18  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER B  -17  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ HIS B  -16  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ MET B  -15  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ ALA B  -14  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER B  -13  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ MET B  -12  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ THR B  -11  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY B  -10  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY B   -9  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLN B   -8  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLN B   -7  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ MET B   -6  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY B   -5  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ ARG B   -4  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLY B   -3  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ SER B   -2  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ GLU B   -1  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ PHE B    0  UNP  T2B4K5              EXPRESSION TAG
SEQADV 8XIZ ILE B  250  UNP  T2B4K5    ALA   250 ENGINEERED MUTATION
SEQADV 8XIZ VAL B  344  UNP  T2B4K5    LEU   344 ENGINEERED MUTATION
SEQRES   1 A  431  MET GLY SER SER HIS HIS HIS HIS HIS HIS SER SER GLY
SEQRES   2 A  431  LEU VAL PRO ARG GLY SER HIS MET ALA SER MET THR GLY
SEQRES   3 A  431  GLY GLN GLN MET GLY ARG GLY SER GLU PHE MET ALA LEU
SEQRES   4 A  431  ALA TYR SER ASN ILE PRO LEU GLY ALA THR VAL ILE PRO
SEQRES   5 A  431  SER PRO PHE GLN VAL HIS ILE SER ASP GLU GLN ILE GLU
SEQRES   6 A  431  GLU LEU GLN LEU LEU VAL LYS LEU SER LYS LEU ALA PRO
SEQRES   7 A  431  PRO THR TYR GLU GLY LEU GLN GLN ASP ARG ARG TYR GLY
SEQRES   8 A  431  ILE THR ASN GLU TRP LEU ALA ASN ALA LYS GLU ALA TRP
SEQRES   9 A  431  LYS SER PHE ASP TRP ARG PRO ALA GLU SER ARG ILE ASN
SEQRES  10 A  431  SER PHE PRO GLN PHE THR TYR ASP ILE GLU GLY LEU THR
SEQRES  11 A  431  ILE HIS PHE VAL ALA LEU PHE SER GLU LYS LYS ASP ALA
SEQRES  12 A  431  ILE PRO ILE VAL LEU LEU HIS GLY TRP PRO GLY SER PHE
SEQRES  13 A  431  LEU GLU PHE LEU PRO VAL LEU THR SER ILE ARG ASP LYS
SEQRES  14 A  431  TYR SER PRO GLU THR LEU PRO TYR HIS ILE VAL VAL PRO
SEQRES  15 A  431  SER LEU PRO GLY TYR THR PHE SER SER GLY PRO PRO LEU
SEQRES  16 A  431  ASP VAL ASN PHE ASN GLY GLU ASP THR ALA ARG VAL ILE
SEQRES  17 A  431  ASN LYS VAL MET LEU ASN LEU GLY PHE GLU ASP GLY TYR
SEQRES  18 A  431  VAL ALA GLN GLY GLY ASP ILE GLY SER LYS ILE GLY ARG
SEQRES  19 A  431  ILE LEU ALA VAL ASP HIS ASP ALA CYS LYS ALA VAL HIS
SEQRES  20 A  431  LEU ASN ALA CYS TYR MET GLY LYS PRO SER SER ILE PRO
SEQRES  21 A  431  ASP THR ALA ILE THR GLU GLU ASP LYS ARG ALA LEU ALA
SEQRES  22 A  431  ARG ALA GLN TRP PHE ALA THR PHE GLY SER GLY TYR ILE
SEQRES  23 A  431  VAL GLU HIS GLY THR ARG PRO SER THR ILE GLY ASN ALA
SEQRES  24 A  431  LEU SER THR SER PRO VAL ALA LEU LEU SER TRP ILE GLY
SEQRES  25 A  431  GLU LYS PHE LEU ASP TRP ALA GLY GLU THR ILE PRO LEU
SEQRES  26 A  431  GLU THR ILE LEU GLU SER VAL THR LEU TYR TRP PHE THR
SEQRES  27 A  431  GLU THR PHE PRO ARG SER ILE TYR HIS TYR ARG GLU ASN
SEQRES  28 A  431  PHE PRO PRO PRO LYS LEU ARG HIS THR GLU ASP PRO ARG
SEQRES  29 A  431  TRP TYR ILE ARG LYS PRO PHE GLY PHE SER TYR TYR PRO
SEQRES  30 A  431  MET GLU VAL VAL PRO THR PRO ARG ALA TRP VAL GLU THR
SEQRES  31 A  431  THR GLY ASN LEU VAL PHE TRP GLN ALA HIS GLU LYS GLY
SEQRES  32 A  431  GLY HIS PHE ALA ALA LEU GLU ARG PRO GLN ASP TYR LEU
SEQRES  33 A  431  ASP ASP LEU THR ALA PHE CYS GLU GLN VAL TRP ALA GLY
SEQRES  34 A  431  ARG LYS
SEQRES   1 B  431  MET GLY SER SER HIS HIS HIS HIS HIS HIS SER SER GLY
SEQRES   2 B  431  LEU VAL PRO ARG GLY SER HIS MET ALA SER MET THR GLY
SEQRES   3 B  431  GLY GLN GLN MET GLY ARG GLY SER GLU PHE MET ALA LEU
SEQRES   4 B  431  ALA TYR SER ASN ILE PRO LEU GLY ALA THR VAL ILE PRO
SEQRES   5 B  431  SER PRO PHE GLN VAL HIS ILE SER ASP GLU GLN ILE GLU
SEQRES   6 B  431  GLU LEU GLN LEU LEU VAL LYS LEU SER LYS LEU ALA PRO
SEQRES   7 B  431  PRO THR TYR GLU GLY LEU GLN GLN ASP ARG ARG TYR GLY
SEQRES   8 B  431  ILE THR ASN GLU TRP LEU ALA ASN ALA LYS GLU ALA TRP
SEQRES   9 B  431  LYS SER PHE ASP TRP ARG PRO ALA GLU SER ARG ILE ASN
SEQRES  10 B  431  SER PHE PRO GLN PHE THR TYR ASP ILE GLU GLY LEU THR
SEQRES  11 B  431  ILE HIS PHE VAL ALA LEU PHE SER GLU LYS LYS ASP ALA
SEQRES  12 B  431  ILE PRO ILE VAL LEU LEU HIS GLY TRP PRO GLY SER PHE
SEQRES  13 B  431  LEU GLU PHE LEU PRO VAL LEU THR SER ILE ARG ASP LYS
SEQRES  14 B  431  TYR SER PRO GLU THR LEU PRO TYR HIS ILE VAL VAL PRO
SEQRES  15 B  431  SER LEU PRO GLY TYR THR PHE SER SER GLY PRO PRO LEU
SEQRES  16 B  431  ASP VAL ASN PHE ASN GLY GLU ASP THR ALA ARG VAL ILE
SEQRES  17 B  431  ASN LYS VAL MET LEU ASN LEU GLY PHE GLU ASP GLY TYR
SEQRES  18 B  431  VAL ALA GLN GLY GLY ASP ILE GLY SER LYS ILE GLY ARG
SEQRES  19 B  431  ILE LEU ALA VAL ASP HIS ASP ALA CYS LYS ALA VAL HIS
SEQRES  20 B  431  LEU ASN ALA CYS TYR MET GLY LYS PRO SER SER ILE PRO
SEQRES  21 B  431  ASP THR ALA ILE THR GLU GLU ASP LYS ARG ALA LEU ALA
SEQRES  22 B  431  ARG ALA GLN TRP PHE ALA THR PHE GLY SER GLY TYR ILE
SEQRES  23 B  431  VAL GLU HIS GLY THR ARG PRO SER THR ILE GLY ASN ALA
SEQRES  24 B  431  LEU SER THR SER PRO VAL ALA LEU LEU SER TRP ILE GLY
SEQRES  25 B  431  GLU LYS PHE LEU ASP TRP ALA GLY GLU THR ILE PRO LEU
SEQRES  26 B  431  GLU THR ILE LEU GLU SER VAL THR LEU TYR TRP PHE THR
SEQRES  27 B  431  GLU THR PHE PRO ARG SER ILE TYR HIS TYR ARG GLU ASN
SEQRES  28 B  431  PHE PRO PRO PRO LYS LEU ARG HIS THR GLU ASP PRO ARG
SEQRES  29 B  431  TRP TYR ILE ARG LYS PRO PHE GLY PHE SER TYR TYR PRO
SEQRES  30 B  431  MET GLU VAL VAL PRO THR PRO ARG ALA TRP VAL GLU THR
SEQRES  31 B  431  THR GLY ASN LEU VAL PHE TRP GLN ALA HIS GLU LYS GLY
SEQRES  32 B  431  GLY HIS PHE ALA ALA LEU GLU ARG PRO GLN ASP TYR LEU
SEQRES  33 B  431  ASP ASP LEU THR ALA PHE CYS GLU GLN VAL TRP ALA GLY
SEQRES  34 B  431  ARG LYS
FORMUL   3  HOH   *465(H2 O)
HELIX    1 AA1 SER A   24  SER A   38  1                                  15
HELIX    2 AA2 TYR A   45  GLN A   49  5                                   5
HELIX    3 AA3 THR A   57  SER A   70  1                                  14
HELIX    4 AA4 ASP A   72  SER A   82  1                                  11
HELIX    5 AA5 SER A  119  GLU A  122  5                                   4
HELIX    6 AA6 PHE A  123  TYR A  134  1                                  12
HELIX    7 AA7 ASN A  164  LEU A  179  1                                  16
HELIX    8 AA8 ASP A  191  HIS A  204  1                                  14
HELIX    9 AA9 THR A  229  GLY A  246  1                                  18
HELIX   10 AB1 SER A  247  ARG A  256  1                                  10
HELIX   11 AB2 ARG A  256  THR A  266  1                                  11
HELIX   12 AB3 SER A  267  ALA A  283  1                                  17
HELIX   13 AB4 PRO A  288  THR A  302  1                                  15
HELIX   14 AB5 GLU A  303  SER A  308  1                                   6
HELIX   15 AB6 ILE A  309  PHE A  316  5                                   8
HELIX   16 AB7 ARG A  322  TYR A  330  5                                   9
HELIX   17 AB8 PRO A  348  THR A  354  1                                   7
HELIX   18 AB9 PHE A  370  ARG A  375  1                                   6
HELIX   19 AC1 ARG A  375  TRP A  391  1                                  17
HELIX   20 AC2 SER B   24  SER B   38  1                                  15
HELIX   21 AC3 TYR B   45  GLN B   49  5                                   5
HELIX   22 AC4 THR B   57  SER B   70  1                                  14
HELIX   23 AC5 ASP B   72  SER B   82  1                                  11
HELIX   24 AC6 SER B  119  GLU B  122  5                                   4
HELIX   25 AC7 PHE B  123  TYR B  134  1                                  12
HELIX   26 AC8 ASN B  164  LEU B  179  1                                  16
HELIX   27 AC9 ASP B  191  HIS B  204  1                                  14
HELIX   28 AD1 THR B  229  GLY B  246  1                                  18
HELIX   29 AD2 SER B  247  ARG B  256  1                                  10
HELIX   30 AD3 ARG B  256  THR B  266  1                                  11
HELIX   31 AD4 SER B  267  ALA B  283  1                                  17
HELIX   32 AD5 PRO B  288  THR B  302  1                                  15
HELIX   33 AD6 GLU B  303  ILE B  309  1                                   7
HELIX   34 AD7 TYR B  310  PHE B  316  5                                   7
HELIX   35 AD8 ARG B  322  TYR B  330  5                                   9
HELIX   36 AD9 PRO B  348  GLU B  353  1                                   6
HELIX   37 AE1 PHE B  370  ARG B  375  1                                   6
HELIX   38 AE2 ARG B  375  TRP B  391  1                                  17
SHEET    1 AA1 9 SER A  17  PRO A  18  0
SHEET    2 AA1 9 GLN A  85  ILE A  90 -1  O  THR A  87   N  SER A  17
SHEET    3 AA1 9 LEU A  93  LEU A 100 -1  O  ILE A  95   N  TYR A  88
SHEET    4 AA1 9 TYR A 141  PRO A 146 -1  O  VAL A 145   N  VAL A  98
SHEET    5 AA1 9 ILE A 108  LEU A 113  1  N  ILE A 110   O  VAL A 144
SHEET    6 AA1 9 TYR A 185  GLY A 189  1  O  VAL A 186   N  VAL A 111
SHEET    7 AA1 9 CYS A 207  LEU A 212  1  O  LYS A 208   N  TYR A 185
SHEET    8 AA1 9 ILE A 331  TYR A 339  1  O  GLY A 336   N  LEU A 212
SHEET    9 AA1 9 GLY A 356  ALA A 363  1  O  ASN A 357   N  ILE A 331
SHEET    1 AA2 9 SER B  17  PRO B  18  0
SHEET    2 AA2 9 GLN B  85  ILE B  90 -1  O  THR B  87   N  SER B  17
SHEET    3 AA2 9 LEU B  93  LEU B 100 -1  O  ILE B  95   N  TYR B  88
SHEET    4 AA2 9 TYR B 141  PRO B 146 -1  O  VAL B 145   N  VAL B  98
SHEET    5 AA2 9 ILE B 108  LEU B 113  1  N  ILE B 110   O  HIS B 142
SHEET    6 AA2 9 TYR B 185  GLY B 189  1  O  GLN B 188   N  LEU B 113
SHEET    7 AA2 9 CYS B 207  LEU B 212  1  O  HIS B 211   N  ALA B 187
SHEET    8 AA2 9 ILE B 331  TYR B 339  1  O  GLY B 336   N  LEU B 212
SHEET    9 AA2 9 GLY B 356  ALA B 363  1  O  GLN B 362   N  TYR B 339
CISPEP   1 TRP A  116    PRO A  117          0         1.71
CISPEP   2 GLY A  156    PRO A  157          0        -0.85
CISPEP   3 PHE A  316    PRO A  317          0        -2.39
CISPEP   4 TRP B  116    PRO B  117          0         2.13
CISPEP   5 GLY B  156    PRO B  157          0        -1.32
CISPEP   6 PHE B  316    PRO B  317          0         0.24
CRYST1   57.049   51.287  133.812  90.00  90.52  90.00 P 1 21 1      4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.017529  0.000000  0.000160        0.00000
SCALE2      0.000000  0.019498  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007473        0.00000
TER    3120      GLY A 393
TER    6235      GLY B 393
MASTER      331    0    0   38   18    0    0    6 6698    2    0   68
END