longtext: 9V2L-pdb

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HEADER    VIRAL PROTEIN/HYDROLASE                 20-MAY-25   9V2L
TITLE     COMPLEX STRUCTURE OF 2014-422 SPIKE RBD BOUND TO HUMAN DPP4
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: SPIKE GLYCOPROTEIN;
COMPND   3 CHAIN: A;
COMPND   4 SYNONYM: S GLYCOPROTEIN,E2,PEPLOMER PROTEIN;
COMPND   5 ENGINEERED: YES;
COMPND   6 MOL_ID: 2;
COMPND   7 MOLECULE: DIPEPTIDYL PEPTIDASE 4 SOLUBLE FORM;
COMPND   8 CHAIN: B, C;
COMPND   9 SYNONYM: DIPEPTIDYL PEPTIDASE IV SOLUBLE FORM;
COMPND  10 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: MIDDLE EAST RESPIRATORY SYNDROME-RELATED
SOURCE   3 CORONAVIRUS;
SOURCE   4 ORGANISM_TAXID: 1335626;
SOURCE   5 EXPRESSION_SYSTEM: TRICHOPLUSIA NI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 7111;
SOURCE   7 MOL_ID: 2;
SOURCE   8 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   9 ORGANISM_COMMON: HUMAN;
SOURCE  10 ORGANISM_TAXID: 9606;
SOURCE  11 GENE: DPP4, ADCP2, CD26;
SOURCE  12 EXPRESSION_SYSTEM: HOMO SAPIENS;
SOURCE  13 EXPRESSION_SYSTEM_TAXID: 9606
KEYWDS    2014-422 RBD, HUMAN DPP4, VIRAL PROTEIN, HYDROLASE, VIRAL
KEYWDS   2 PROTEIN/HYDROLASE, VIRAL PROTEIN-HYDROLASE COMPLEX
EXPDTA    ELECTRON MICROSCOPY
AUTHOR    X.WANG,Z.LIN
REVDAT   1   24-DEC-25 9V2L    0
JRNL        AUTH   X.WANG,Z.LIN
JRNL        TITL   COMPLEX STRUCTURE OF 2014-422 SPIKE RBD BOUND TO HUMAN DPP4
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    3.00 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   SOFTWARE PACKAGES      : PHENIX
REMARK   3   RECONSTRUCTION SCHEMA  : NULL
REMARK   3
REMARK   3 EM MAP-MODEL FITTING AND REFINEMENT
REMARK   3   PDB ENTRY                    : NULL
REMARK   3   REFINEMENT SPACE             : NULL
REMARK   3   REFINEMENT PROTOCOL          : NULL
REMARK   3   REFINEMENT TARGET            : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE  : NULL
REMARK   3
REMARK   3 FITTING PROCEDURE : NULL
REMARK   3
REMARK   3 EM IMAGE RECONSTRUCTION STATISTICS
REMARK   3   NOMINAL PIXEL SIZE (ANGSTROMS)    : NULL
REMARK   3   ACTUAL PIXEL SIZE  (ANGSTROMS)    : NULL
REMARK   3   EFFECTIVE RESOLUTION (ANGSTROMS)  : 3.000
REMARK   3   NUMBER OF PARTICLES               : 591107
REMARK   3   CTF CORRECTION METHOD             : NONE
REMARK   3
REMARK   3 EM RECONSTRUCTION MAGNIFICATION CALIBRATION: NULL
REMARK   3
REMARK   3 OTHER DETAILS: NULL
REMARK   4
REMARK   4 9V2L COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBC ON 23-MAY-25.
REMARK 100 THE DEPOSITION ID IS D_1300059647.
REMARK 245
REMARK 245 EXPERIMENTAL DETAILS
REMARK 245   RECONSTRUCTION METHOD          : SINGLE PARTICLE
REMARK 245   SPECIMEN TYPE                  : NULL
REMARK 245
REMARK 245 ELECTRON MICROSCOPE SAMPLE
REMARK 245   SAMPLE TYPE                    : PARTICLE
REMARK 245   PARTICLE TYPE                  : POINT
REMARK 245   NAME OF SAMPLE                 : COMPLEX STRUCTURE OF 2014-422
REMARK 245                                    SPIKE RBD BOUND TO HUMAN DPP4
REMARK 245   SAMPLE CONCENTRATION (MG ML-1) : NULL
REMARK 245   SAMPLE SUPPORT DETAILS         : NULL
REMARK 245   SAMPLE VITRIFICATION DETAILS   : NULL
REMARK 245   SAMPLE BUFFER                  : NULL
REMARK 245   PH                             : 8.00
REMARK 245   SAMPLE DETAILS                 : NULL
REMARK 245
REMARK 245 DATA ACQUISITION
REMARK 245   DATE OF EXPERIMENT                : NULL
REMARK 245   NUMBER OF MICROGRAPHS-IMAGES      : NULL
REMARK 245   TEMPERATURE (KELVIN)              : NULL
REMARK 245   MICROSCOPE MODEL                  : TFS KRIOS
REMARK 245   DETECTOR TYPE                     : GATAN K3 (6K X 4K)
REMARK 245   MINIMUM DEFOCUS (NM)              : 1200.00
REMARK 245   MAXIMUM DEFOCUS (NM)              : 1800.00
REMARK 245   MINIMUM TILT ANGLE (DEGREES)      : NULL
REMARK 245   MAXIMUM TILT ANGLE (DEGREES)      : NULL
REMARK 245   NOMINAL CS                        : NULL
REMARK 245   IMAGING MODE                      : BRIGHT FIELD
REMARK 245   ELECTRON DOSE (ELECTRONS NM**-2)  : 5000.00
REMARK 245   ILLUMINATION MODE                 : FLOOD BEAM
REMARK 245   NOMINAL MAGNIFICATION             : NULL
REMARK 245   CALIBRATED MAGNIFICATION          : NULL
REMARK 245   SOURCE                            : FIELD EMISSION GUN
REMARK 245   ACCELERATION VOLTAGE (KV)         : 300
REMARK 245   IMAGING DETAILS                   : NULL
REMARK 247
REMARK 247 ELECTRON MICROSCOPY
REMARK 247  THE COORDINATES IN THIS ENTRY WERE GENERATED FROM ELECTRON
REMARK 247  MICROSCOPY DATA. PROTEIN DATA BANK CONVENTIONS REQUIRE
REMARK 247  THAT CRYST1 AND SCALE RECORDS BE INCLUDED, BUT THE VALUES
REMARK 247  ON THESE RECORDS ARE MEANINGLESS EXCEPT FOR THE CALCULATION
REMARK 247  OF THE STRUCTURE FACTORS.
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: TRIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: TRIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C, D, E, F
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLU A   371
REMARK 465     VAL A   372
REMARK 465     HIS A   373
REMARK 465     SER A   374
REMARK 465     ARG A   375
REMARK 465     GLY A   376
REMARK 465     GLN A   377
REMARK 465     PHE A   378
REMARK 465     ILE A   379
REMARK 465     GLU A   380
REMARK 465     GLN A   381
REMARK 465     PRO A   382
REMARK 465     ASN A   383
REMARK 465     SER A   384
REMARK 465     MET A   592
REMARK 465     ASP A   593
REMARK 465     LEU B   765
REMARK 465     PRO B   766
REMARK 465     LEU C   765
REMARK 465     PRO C   766
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OD2  ASP C   331     ND2  ASN C   338              2.09
REMARK 500   O    TYR C    43     OG1  THR C   570              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASN A 402       56.69    -91.97
REMARK 500    SER A 433       67.05     36.65
REMARK 500    PRO A 434       44.73    -82.32
REMARK 500    ASP A 435       -3.54     65.63
REMARK 500    ALA A 436      165.84    179.98
REMARK 500    ALA A 438       -0.75   -140.95
REMARK 500    ASN A 472      -60.30   -101.25
REMARK 500    GLN B  72     -161.67    -78.11
REMARK 500    ASN B  74       12.03   -142.55
REMARK 500    TRP B 124     -166.46   -129.33
REMARK 500    ILE B 193      -60.03   -124.74
REMARK 500    SER B 242      -13.94     75.78
REMARK 500    ASP B 243      164.40    178.98
REMARK 500    ALA B 306      -62.79    -93.72
REMARK 500    THR B 401       59.41    -94.78
REMARK 500    ASN B 450       79.67   -156.35
REMARK 500    SER B 473       31.84   -141.57
REMARK 500    ASN B 487       24.27   -140.33
REMARK 500    ASN B 520      -28.28     75.10
REMARK 500    GLU B 521      -19.93   -152.45
REMARK 500    SER B 630      -27.07     73.85
REMARK 500    ASP B 678      -71.75    -83.29
REMARK 500    ASN B 679       30.14   -147.30
REMARK 500    TYR C  58       77.14   -101.33
REMARK 500    SER C  64     -168.76   -160.43
REMARK 500    PHE C  98       -7.54     71.10
REMARK 500    GLN C 123      -66.39    -96.93
REMARK 500    TRP C 124     -167.76   -126.38
REMARK 500    LYS C 139      -84.36    -82.79
REMARK 500    ARG C 140        2.82   -159.88
REMARK 500    ASN C 151       53.05    -92.10
REMARK 500    ILE C 193      -59.95   -123.26
REMARK 500    TYR C 241      -79.99    -79.42
REMARK 500    SER C 242     -171.24    167.59
REMARK 500    TRP C 337       79.14   -103.38
REMARK 500    THR C 401       51.81    -91.56
REMARK 500    SER C 473       16.32   -142.22
REMARK 500    ASN C 520      -27.02     74.99
REMARK 500    GLU C 521      -19.20   -150.38
REMARK 500    ALA C 548       17.39     54.25
REMARK 500    SER C 630     -145.52   -159.89
REMARK 500    TYR C 631      -31.00    -38.89
REMARK 500    ASP C 678      -66.08    -94.48
REMARK 500    ILE C 742      109.53    -56.94
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 VAL B  546     TYR B  547                  148.20
REMARK 500 LYS C  139     ARG C  140                  145.11
REMARK 500 SER C  242     ASP C  243                  148.06
REMARK 500
REMARK 500 REMARK: NULL
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: EMD-64732   RELATED DB: EMDB
REMARK 900 COMPLEX STRUCTURE OF 2014-422 SPIKE RBD BOUND TO HUMAN DPP4
DBREF1 9V2L A  371   593  UNP                  A0A2R4KP93_MERS
DBREF2 9V2L A     A0A2R4KP93                        371         593
DBREF  9V2L B   40   766  UNP    P27487   DPP4_HUMAN      40    766
DBREF  9V2L C   40   766  UNP    P27487   DPP4_HUMAN      40    766
SEQRES   1 A  223  GLU VAL HIS SER ARG GLY GLN PHE ILE GLU GLN PRO ASN
SEQRES   2 A  223  SER VAL GLU CYS ASP PHE THR LYS LEU LEU SER GLY THR
SEQRES   3 A  223  PRO PRO GLN VAL TYR ASN PHE ASN ARG LEU VAL PHE THR
SEQRES   4 A  223  ASN CYS ASN TYR ASN LEU THR LYS LEU LEU SER LEU PHE
SEQRES   5 A  223  MET VAL ASN GLU PHE SER CYS ASP GLY ILE SER PRO ASP
SEQRES   6 A  223  ALA ILE ALA ARG GLY CYS TYR SER SER LEU THR VAL ASP
SEQRES   7 A  223  TYR PHE ALA TYR PRO LEU SER MET LYS SER TYR MET GLN
SEQRES   8 A  223  PRO GLY SER ALA GLY VAL ILE SER GLN TYR ASN TYR LYS
SEQRES   9 A  223  GLN SER PHE ALA ASN PRO THR CYS ARG ILE PHE ALA THR
SEQRES  10 A  223  ALA PRO ALA ASN LEU THR ILE THR LYS PRO SER SER TYR
SEQRES  11 A  223  SER PHE ILE SER LYS CYS SER ARG LEU THR GLY ASP ASN
SEQRES  12 A  223  SER HIS ILE GLU THR PRO ILE VAL ILE ASN PRO GLY GLU
SEQRES  13 A  223  TYR SER ILE CYS LYS ASN PHE ALA PRO ASN GLY PHE SER
SEQRES  14 A  223  GLN ASP GLY ASP TYR PHE THR ARG GLN LEU SER GLN LEU
SEQRES  15 A  223  GLU GLY GLY GLY ILE LEU VAL GLY VAL GLY SER VAL THR
SEQRES  16 A  223  PRO MET THR ASP THR LEU GLN MET GLY PHE ILE ILE SER
SEQRES  17 A  223  VAL GLN TYR GLY THR ASP THR ASN SER VAL CYS PRO MET
SEQRES  18 A  223  MET ASP
SEQRES   1 B  727  ARG LYS THR TYR THR LEU THR ASP TYR LEU LYS ASN THR
SEQRES   2 B  727  TYR ARG LEU LYS LEU TYR SER LEU ARG TRP ILE SER ASP
SEQRES   3 B  727  HIS GLU TYR LEU TYR LYS GLN GLU ASN ASN ILE LEU VAL
SEQRES   4 B  727  PHE ASN ALA GLU TYR GLY ASN SER SER VAL PHE LEU GLU
SEQRES   5 B  727  ASN SER THR PHE ASP GLU PHE GLY HIS SER ILE ASN ASP
SEQRES   6 B  727  TYR SER ILE SER PRO ASP GLY GLN PHE ILE LEU LEU GLU
SEQRES   7 B  727  TYR ASN TYR VAL LYS GLN TRP ARG HIS SER TYR THR ALA
SEQRES   8 B  727  SER TYR ASP ILE TYR ASP LEU ASN LYS ARG GLN LEU ILE
SEQRES   9 B  727  THR GLU GLU ARG ILE PRO ASN ASN THR GLN TRP VAL THR
SEQRES  10 B  727  TRP SER PRO VAL GLY HIS LYS LEU ALA TYR VAL TRP ASN
SEQRES  11 B  727  ASN ASP ILE TYR VAL LYS ILE GLU PRO ASN LEU PRO SER
SEQRES  12 B  727  TYR ARG ILE THR TRP THR GLY LYS GLU ASP ILE ILE TYR
SEQRES  13 B  727  ASN GLY ILE THR ASP TRP VAL TYR GLU GLU GLU VAL PHE
SEQRES  14 B  727  SER ALA TYR SER ALA LEU TRP TRP SER PRO ASN GLY THR
SEQRES  15 B  727  PHE LEU ALA TYR ALA GLN PHE ASN ASP THR GLU VAL PRO
SEQRES  16 B  727  LEU ILE GLU TYR SER PHE TYR SER ASP GLU SER LEU GLN
SEQRES  17 B  727  TYR PRO LYS THR VAL ARG VAL PRO TYR PRO LYS ALA GLY
SEQRES  18 B  727  ALA VAL ASN PRO THR VAL LYS PHE PHE VAL VAL ASN THR
SEQRES  19 B  727  ASP SER LEU SER SER VAL THR ASN ALA THR SER ILE GLN
SEQRES  20 B  727  ILE THR ALA PRO ALA SER MET LEU ILE GLY ASP HIS TYR
SEQRES  21 B  727  LEU CYS ASP VAL THR TRP ALA THR GLN GLU ARG ILE SER
SEQRES  22 B  727  LEU GLN TRP LEU ARG ARG ILE GLN ASN TYR SER VAL MET
SEQRES  23 B  727  ASP ILE CYS ASP TYR ASP GLU SER SER GLY ARG TRP ASN
SEQRES  24 B  727  CYS LEU VAL ALA ARG GLN HIS ILE GLU MET SER THR THR
SEQRES  25 B  727  GLY TRP VAL GLY ARG PHE ARG PRO SER GLU PRO HIS PHE
SEQRES  26 B  727  THR LEU ASP GLY ASN SER PHE TYR LYS ILE ILE SER ASN
SEQRES  27 B  727  GLU GLU GLY TYR ARG HIS ILE CYS TYR PHE GLN ILE ASP
SEQRES  28 B  727  LYS LYS ASP CYS THR PHE ILE THR LYS GLY THR TRP GLU
SEQRES  29 B  727  VAL ILE GLY ILE GLU ALA LEU THR SER ASP TYR LEU TYR
SEQRES  30 B  727  TYR ILE SER ASN GLU TYR LYS GLY MET PRO GLY GLY ARG
SEQRES  31 B  727  ASN LEU TYR LYS ILE GLN LEU SER ASP TYR THR LYS VAL
SEQRES  32 B  727  THR CYS LEU SER CYS GLU LEU ASN PRO GLU ARG CYS GLN
SEQRES  33 B  727  TYR TYR SER VAL SER PHE SER LYS GLU ALA LYS TYR TYR
SEQRES  34 B  727  GLN LEU ARG CYS SER GLY PRO GLY LEU PRO LEU TYR THR
SEQRES  35 B  727  LEU HIS SER SER VAL ASN ASP LYS GLY LEU ARG VAL LEU
SEQRES  36 B  727  GLU ASP ASN SER ALA LEU ASP LYS MET LEU GLN ASN VAL
SEQRES  37 B  727  GLN MET PRO SER LYS LYS LEU ASP PHE ILE ILE LEU ASN
SEQRES  38 B  727  GLU THR LYS PHE TRP TYR GLN MET ILE LEU PRO PRO HIS
SEQRES  39 B  727  PHE ASP LYS SER LYS LYS TYR PRO LEU LEU LEU ASP VAL
SEQRES  40 B  727  TYR ALA GLY PRO CYS SER GLN LYS ALA ASP THR VAL PHE
SEQRES  41 B  727  ARG LEU ASN TRP ALA THR TYR LEU ALA SER THR GLU ASN
SEQRES  42 B  727  ILE ILE VAL ALA SER PHE ASP GLY ARG GLY SER GLY TYR
SEQRES  43 B  727  GLN GLY ASP LYS ILE MET HIS ALA ILE ASN ARG ARG LEU
SEQRES  44 B  727  GLY THR PHE GLU VAL GLU ASP GLN ILE GLU ALA ALA ARG
SEQRES  45 B  727  GLN PHE SER LYS MET GLY PHE VAL ASP ASN LYS ARG ILE
SEQRES  46 B  727  ALA ILE TRP GLY TRP SER TYR GLY GLY TYR VAL THR SER
SEQRES  47 B  727  MET VAL LEU GLY SER GLY SER GLY VAL PHE LYS CYS GLY
SEQRES  48 B  727  ILE ALA VAL ALA PRO VAL SER ARG TRP GLU TYR TYR ASP
SEQRES  49 B  727  SER VAL TYR THR GLU ARG TYR MET GLY LEU PRO THR PRO
SEQRES  50 B  727  GLU ASP ASN LEU ASP HIS TYR ARG ASN SER THR VAL MET
SEQRES  51 B  727  SER ARG ALA GLU ASN PHE LYS GLN VAL GLU TYR LEU LEU
SEQRES  52 B  727  ILE HIS GLY THR ALA ASP ASP ASN VAL HIS PHE GLN GLN
SEQRES  53 B  727  SER ALA GLN ILE SER LYS ALA LEU VAL ASP VAL GLY VAL
SEQRES  54 B  727  ASP PHE GLN ALA MET TRP TYR THR ASP GLU ASP HIS GLY
SEQRES  55 B  727  ILE ALA SER SER THR ALA HIS GLN HIS ILE TYR THR HIS
SEQRES  56 B  727  MET SER HIS PHE ILE LYS GLN CYS PHE SER LEU PRO
SEQRES   1 C  727  ARG LYS THR TYR THR LEU THR ASP TYR LEU LYS ASN THR
SEQRES   2 C  727  TYR ARG LEU LYS LEU TYR SER LEU ARG TRP ILE SER ASP
SEQRES   3 C  727  HIS GLU TYR LEU TYR LYS GLN GLU ASN ASN ILE LEU VAL
SEQRES   4 C  727  PHE ASN ALA GLU TYR GLY ASN SER SER VAL PHE LEU GLU
SEQRES   5 C  727  ASN SER THR PHE ASP GLU PHE GLY HIS SER ILE ASN ASP
SEQRES   6 C  727  TYR SER ILE SER PRO ASP GLY GLN PHE ILE LEU LEU GLU
SEQRES   7 C  727  TYR ASN TYR VAL LYS GLN TRP ARG HIS SER TYR THR ALA
SEQRES   8 C  727  SER TYR ASP ILE TYR ASP LEU ASN LYS ARG GLN LEU ILE
SEQRES   9 C  727  THR GLU GLU ARG ILE PRO ASN ASN THR GLN TRP VAL THR
SEQRES  10 C  727  TRP SER PRO VAL GLY HIS LYS LEU ALA TYR VAL TRP ASN
SEQRES  11 C  727  ASN ASP ILE TYR VAL LYS ILE GLU PRO ASN LEU PRO SER
SEQRES  12 C  727  TYR ARG ILE THR TRP THR GLY LYS GLU ASP ILE ILE TYR
SEQRES  13 C  727  ASN GLY ILE THR ASP TRP VAL TYR GLU GLU GLU VAL PHE
SEQRES  14 C  727  SER ALA TYR SER ALA LEU TRP TRP SER PRO ASN GLY THR
SEQRES  15 C  727  PHE LEU ALA TYR ALA GLN PHE ASN ASP THR GLU VAL PRO
SEQRES  16 C  727  LEU ILE GLU TYR SER PHE TYR SER ASP GLU SER LEU GLN
SEQRES  17 C  727  TYR PRO LYS THR VAL ARG VAL PRO TYR PRO LYS ALA GLY
SEQRES  18 C  727  ALA VAL ASN PRO THR VAL LYS PHE PHE VAL VAL ASN THR
SEQRES  19 C  727  ASP SER LEU SER SER VAL THR ASN ALA THR SER ILE GLN
SEQRES  20 C  727  ILE THR ALA PRO ALA SER MET LEU ILE GLY ASP HIS TYR
SEQRES  21 C  727  LEU CYS ASP VAL THR TRP ALA THR GLN GLU ARG ILE SER
SEQRES  22 C  727  LEU GLN TRP LEU ARG ARG ILE GLN ASN TYR SER VAL MET
SEQRES  23 C  727  ASP ILE CYS ASP TYR ASP GLU SER SER GLY ARG TRP ASN
SEQRES  24 C  727  CYS LEU VAL ALA ARG GLN HIS ILE GLU MET SER THR THR
SEQRES  25 C  727  GLY TRP VAL GLY ARG PHE ARG PRO SER GLU PRO HIS PHE
SEQRES  26 C  727  THR LEU ASP GLY ASN SER PHE TYR LYS ILE ILE SER ASN
SEQRES  27 C  727  GLU GLU GLY TYR ARG HIS ILE CYS TYR PHE GLN ILE ASP
SEQRES  28 C  727  LYS LYS ASP CYS THR PHE ILE THR LYS GLY THR TRP GLU
SEQRES  29 C  727  VAL ILE GLY ILE GLU ALA LEU THR SER ASP TYR LEU TYR
SEQRES  30 C  727  TYR ILE SER ASN GLU TYR LYS GLY MET PRO GLY GLY ARG
SEQRES  31 C  727  ASN LEU TYR LYS ILE GLN LEU SER ASP TYR THR LYS VAL
SEQRES  32 C  727  THR CYS LEU SER CYS GLU LEU ASN PRO GLU ARG CYS GLN
SEQRES  33 C  727  TYR TYR SER VAL SER PHE SER LYS GLU ALA LYS TYR TYR
SEQRES  34 C  727  GLN LEU ARG CYS SER GLY PRO GLY LEU PRO LEU TYR THR
SEQRES  35 C  727  LEU HIS SER SER VAL ASN ASP LYS GLY LEU ARG VAL LEU
SEQRES  36 C  727  GLU ASP ASN SER ALA LEU ASP LYS MET LEU GLN ASN VAL
SEQRES  37 C  727  GLN MET PRO SER LYS LYS LEU ASP PHE ILE ILE LEU ASN
SEQRES  38 C  727  GLU THR LYS PHE TRP TYR GLN MET ILE LEU PRO PRO HIS
SEQRES  39 C  727  PHE ASP LYS SER LYS LYS TYR PRO LEU LEU LEU ASP VAL
SEQRES  40 C  727  TYR ALA GLY PRO CYS SER GLN LYS ALA ASP THR VAL PHE
SEQRES  41 C  727  ARG LEU ASN TRP ALA THR TYR LEU ALA SER THR GLU ASN
SEQRES  42 C  727  ILE ILE VAL ALA SER PHE ASP GLY ARG GLY SER GLY TYR
SEQRES  43 C  727  GLN GLY ASP LYS ILE MET HIS ALA ILE ASN ARG ARG LEU
SEQRES  44 C  727  GLY THR PHE GLU VAL GLU ASP GLN ILE GLU ALA ALA ARG
SEQRES  45 C  727  GLN PHE SER LYS MET GLY PHE VAL ASP ASN LYS ARG ILE
SEQRES  46 C  727  ALA ILE TRP GLY TRP SER TYR GLY GLY TYR VAL THR SER
SEQRES  47 C  727  MET VAL LEU GLY SER GLY SER GLY VAL PHE LYS CYS GLY
SEQRES  48 C  727  ILE ALA VAL ALA PRO VAL SER ARG TRP GLU TYR TYR ASP
SEQRES  49 C  727  SER VAL TYR THR GLU ARG TYR MET GLY LEU PRO THR PRO
SEQRES  50 C  727  GLU ASP ASN LEU ASP HIS TYR ARG ASN SER THR VAL MET
SEQRES  51 C  727  SER ARG ALA GLU ASN PHE LYS GLN VAL GLU TYR LEU LEU
SEQRES  52 C  727  ILE HIS GLY THR ALA ASP ASP ASN VAL HIS PHE GLN GLN
SEQRES  53 C  727  SER ALA GLN ILE SER LYS ALA LEU VAL ASP VAL GLY VAL
SEQRES  54 C  727  ASP PHE GLN ALA MET TRP TYR THR ASP GLU ASP HIS GLY
SEQRES  55 C  727  ILE ALA SER SER THR ALA HIS GLN HIS ILE TYR THR HIS
SEQRES  56 C  727  MET SER HIS PHE ILE LYS GLN CYS PHE SER LEU PRO
HET    NAG  D   1      14
HET    NAG  D   2      14
HET    NAG  E   1      14
HET    NAG  E   2      14
HET    NAG  F   1      14
HET    NAG  F   2      14
HET    NAG  B 801      14
HET    NAG  B 802      14
HET    NAG  B 803      14
HET    NAG  B 804      14
HET    NAG  C 801      14
HET    NAG  C 802      14
HET    NAG  C 803      14
HETNAM     NAG 2-ACETAMIDO-2-DEOXY-BETA-D-GLUCOPYRANOSE
HETSYN     NAG N-ACETYL-BETA-D-GLUCOSAMINE; 2-ACETAMIDO-2-DEOXY-BETA-
HETSYN   2 NAG  D-GLUCOSE; 2-ACETAMIDO-2-DEOXY-D-GLUCOSE; 2-ACETAMIDO-
HETSYN   3 NAG  2-DEOXY-GLUCOSE; N-ACETYL-D-GLUCOSAMINE
FORMUL   4  NAG    13(C8 H15 N O6)
HELIX    1 AA1 ASN A  414  PHE A  422  1                                   9
HELIX    2 AA2 PRO A  453  GLN A  461  5                                   9
HELIX    3 AA3 GLY A  466  ASN A  472  1                                   7
HELIX    4 AA4 GLY A  511  HIS A  515  5                                   5
HELIX    5 AA5 SER A  528  ALA A  534  5                                   7
HELIX    6 AA6 THR B   44  LYS B   50  1                                   7
HELIX    7 AA7 ASP B  200  GLU B  206  1                                   7
HELIX    8 AA8 ASP B  274  LEU B  276  5                                   3
HELIX    9 AA9 PRO B  290  ILE B  295  1                                   6
HELIX   10 AB1 GLU B  421  MET B  425  5                                   5
HELIX   11 AB2 LYS B  463  ALA B  465  5                                   3
HELIX   12 AB3 ASN B  497  GLN B  505  1                                   9
HELIX   13 AB4 ASN B  562  GLU B  571  1                                  10
HELIX   14 AB5 ILE B  590  ASN B  595  5                                   6
HELIX   15 AB6 THR B  600  MET B  616  1                                  17
HELIX   16 AB7 SER B  630  GLY B  641  1                                  12
HELIX   17 AB8 ASP B  663  GLY B  672  1                                  10
HELIX   18 AB9 ASN B  679  ASN B  685  1                                   7
HELIX   19 AC1 VAL B  688  VAL B  698  5                                  11
HELIX   20 AC2 HIS B  712  ASP B  725  1                                  14
HELIX   21 AC3 SER B  744  PHE B  763  1                                  20
HELIX   22 AC4 THR C   44  LYS C   50  1                                   7
HELIX   23 AC5 ASP C  200  GLU C  206  1                                   7
HELIX   24 AC6 PRO C  290  ILE C  295  1                                   6
HELIX   25 AC7 GLU C  421  MET C  425  5                                   5
HELIX   26 AC8 LYS C  463  ALA C  465  5                                   3
HELIX   27 AC9 ASN C  497  LEU C  504  1                                   8
HELIX   28 AD1 ASN C  562  GLU C  571  1                                  10
HELIX   29 AD2 GLY C  587  HIS C  592  1                                   6
HELIX   30 AD3 ALA C  593  ASN C  595  5                                   3
HELIX   31 AD4 THR C  600  LYS C  615  1                                  16
HELIX   32 AD5 SER C  630  GLY C  641  1                                  12
HELIX   33 AD6 ASP C  663  MET C  671  1                                   9
HELIX   34 AD7 LEU C  680  SER C  686  1                                   7
HELIX   35 AD8 THR C  687  VAL C  698  5                                  12
HELIX   36 AD9 PHE C  713  GLY C  727  1                                  15
HELIX   37 AE1 SER C  744  SER C  764  1                                  21
SHEET    1 AA1 2 ASN A 412  TYR A 413  0
SHEET    2 AA1 2 CYS A 589  PRO A 590  1  O  CYS A 589   N  TYR A 413
SHEET    1 AA2 4 MET A 423  PHE A 427  0
SHEET    2 AA2 4 THR A 481  ALA A 488 -1  O  THR A 487   N  MET A 423
SHEET    3 AA2 4 LEU A 571  GLN A 580 -1  O  LEU A 571   N  ALA A 488
SHEET    4 AA2 4 SER A 444  VAL A 447 -1  N  THR A 446   O  SER A 578
SHEET    1 AA3 4 ILE A 516  GLU A 517  0
SHEET    2 AA3 4 TYR A 500  THR A 510 -1  N  THR A 510   O  ILE A 516
SHEET    3 AA3 4 ILE A 557  THR A 565 -1  O  SER A 563   N  PHE A 502
SHEET    4 AA3 4 ASP A 543  TYR A 544 -1  N  ASP A 543   O  GLY A 562
SHEET    1 AA4 4 ILE A 516  GLU A 517  0
SHEET    2 AA4 4 TYR A 500  THR A 510 -1  N  THR A 510   O  ILE A 516
SHEET    3 AA4 4 ILE A 557  THR A 565 -1  O  SER A 563   N  PHE A 502
SHEET    4 AA4 4 ARG A 547  GLN A 548 -1  N  ARG A 547   O  LEU A 558
SHEET    1 AA5 2 LYS B  41  THR B  42  0
SHEET    2 AA5 2 VAL B 507  GLN B 508  1  O  GLN B 508   N  LYS B  41
SHEET    1 AA6 3 GLU B  67  LYS B  71  0
SHEET    2 AA6 3 ILE B  76  ASN B  80 -1  O  PHE B  79   N  TYR B  68
SHEET    3 AA6 3 SER B  87  LEU B  90 -1  O  PHE B  89   N  ILE B  76
SHEET    1 AA7 3 ASP B 104  ILE B 107  0
SHEET    2 AA7 3 PHE B 113  LYS B 122 -1  O  GLU B 117   N  ASP B 104
SHEET    3 AA7 3 TYR B 128  ASP B 136 -1  O  SER B 131   N  TYR B 118
SHEET    1 AA8 4 TRP B 154  TRP B 157  0
SHEET    2 AA8 4 LEU B 164  VAL B 167 -1  O  VAL B 167   N  TRP B 154
SHEET    3 AA8 4 ILE B 172  LYS B 175 -1  O  LYS B 175   N  LEU B 164
SHEET    4 AA8 4 TYR B 183  ARG B 184 -1  O  TYR B 183   N  VAL B 174
SHEET    1 AA9 3 ILE B 194  ASN B 196  0
SHEET    2 AA9 3 PHE B 222  ASN B 229 -1  O  PHE B 228   N  TYR B 195
SHEET    3 AA9 3 LEU B 214  TRP B 216 -1  N  TRP B 215   O  ALA B 224
SHEET    1 AB1 4 ILE B 194  ASN B 196  0
SHEET    2 AB1 4 PHE B 222  ASN B 229 -1  O  PHE B 228   N  TYR B 195
SHEET    3 AB1 4 THR B 265  ASN B 272 -1  O  LYS B 267   N  GLN B 227
SHEET    4 AB1 4 SER B 284  ILE B 287 -1  O  ILE B 287   N  PHE B 268
SHEET    1 AB2 2 LEU B 235  PHE B 240  0
SHEET    2 AB2 2 LYS B 250  PRO B 255 -1  O  LYS B 250   N  PHE B 240
SHEET    1 AB3 4 HIS B 298  THR B 307  0
SHEET    2 AB3 4 ARG B 310  ARG B 317 -1  O  SER B 312   N  THR B 304
SHEET    3 AB3 4 TYR B 322  TYR B 330 -1  O  ASP B 326   N  LEU B 313
SHEET    4 AB3 4 TRP B 337  CYS B 339 -1  O  ASN B 338   N  ASP B 329
SHEET    1 AB4 4 HIS B 298  THR B 307  0
SHEET    2 AB4 4 ARG B 310  ARG B 317 -1  O  SER B 312   N  THR B 304
SHEET    3 AB4 4 TYR B 322  TYR B 330 -1  O  ASP B 326   N  LEU B 313
SHEET    4 AB4 4 HIS B 345  MET B 348 -1  O  HIS B 345   N  MET B 325
SHEET    1 AB5 4 HIS B 363  PHE B 364  0
SHEET    2 AB5 4 SER B 370  SER B 376 -1  O  TYR B 372   N  HIS B 363
SHEET    3 AB5 4 ARG B 382  GLN B 388 -1  O  CYS B 385   N  LYS B 373
SHEET    4 AB5 4 CYS B 394  PHE B 396 -1  O  THR B 395   N  TYR B 386
SHEET    1 AB6 4 VAL B 404  LEU B 410  0
SHEET    2 AB6 4 TYR B 414  SER B 419 -1  O  ILE B 418   N  ILE B 405
SHEET    3 AB6 4 ASN B 430  GLN B 435 -1  O  TYR B 432   N  TYR B 417
SHEET    4 AB6 4 VAL B 442  CYS B 444 -1  O  THR B 443   N  LYS B 433
SHEET    1 AB7 4 TYR B 457  PHE B 461  0
SHEET    2 AB7 4 TYR B 467  CYS B 472 -1  O  ARG B 471   N  SER B 458
SHEET    3 AB7 4 LEU B 479  SER B 484 -1  O  HIS B 483   N  TYR B 468
SHEET    4 AB7 4 LYS B 489  GLU B 495 -1  O  GLU B 495   N  TYR B 480
SHEET    1 AB8 8 LYS B 513  LEU B 519  0
SHEET    2 AB8 8 THR B 522  ILE B 529 -1  O  PHE B 524   N  ILE B 517
SHEET    3 AB8 8 ILE B 574  PHE B 578 -1  O  SER B 577   N  GLN B 527
SHEET    4 AB8 8 TYR B 540  VAL B 546  1  N  LEU B 543   O  ILE B 574
SHEET    5 AB8 8 VAL B 619  TRP B 629  1  O  ASP B 620   N  TYR B 540
SHEET    6 AB8 8 CYS B 649  VAL B 653  1  O  VAL B 653   N  GLY B 628
SHEET    7 AB8 8 GLU B 699  GLY B 705  1  O  LEU B 701   N  ALA B 652
SHEET    8 AB8 8 GLN B 731  TYR B 735  1  O  GLN B 731   N  TYR B 700
SHEET    1 AB9 2 LYS C  41  THR C  42  0
SHEET    2 AB9 2 VAL C 507  GLN C 508  1  O  GLN C 508   N  LYS C  41
SHEET    1 AC1 3 GLU C  67  LYS C  71  0
SHEET    2 AC1 3 ILE C  76  ASN C  80 -1  O  LEU C  77   N  TYR C  70
SHEET    3 AC1 3 SER C  86  LEU C  90 -1  O  SER C  87   N  VAL C  78
SHEET    1 AC2 4 TYR C 105  ILE C 107  0
SHEET    2 AC2 4 PHE C 113  LYS C 122 -1  O  LEU C 115   N  SER C 106
SHEET    3 AC2 4 TYR C 128  ASP C 136 -1  O  SER C 131   N  TYR C 118
SHEET    4 AC2 4 LEU C 142  ILE C 143 -1  O  ILE C 143   N  ILE C 134
SHEET    1 AC3 4 THR C 152  TRP C 157  0
SHEET    2 AC3 4 LEU C 164  TRP C 168 -1  O  VAL C 167   N  GLN C 153
SHEET    3 AC3 4 ILE C 172  LYS C 175 -1  O  LYS C 175   N  LEU C 164
SHEET    4 AC3 4 TYR C 183  ARG C 184 -1  O  TYR C 183   N  VAL C 174
SHEET    1 AC4 3 ILE C 194  ASN C 196  0
SHEET    2 AC4 3 PHE C 222  ASN C 229 -1  O  PHE C 228   N  TYR C 195
SHEET    3 AC4 3 LEU C 214  TRP C 216 -1  N  TRP C 215   O  ALA C 224
SHEET    1 AC5 4 ILE C 194  ASN C 196  0
SHEET    2 AC5 4 PHE C 222  ASN C 229 -1  O  PHE C 228   N  TYR C 195
SHEET    3 AC5 4 THR C 265  ASN C 272 -1  O  PHE C 269   N  TYR C 225
SHEET    4 AC5 4 SER C 284  GLN C 286 -1  O  ILE C 285   N  VAL C 270
SHEET    1 AC6 2 LEU C 235  PHE C 240  0
SHEET    2 AC6 2 LYS C 250  PRO C 255 -1  O  VAL C 252   N  TYR C 238
SHEET    1 AC7 4 HIS C 298  ASP C 302  0
SHEET    2 AC7 4 ILE C 311  ARG C 317 -1  O  LEU C 316   N  TYR C 299
SHEET    3 AC7 4 TYR C 322  CYS C 328 -1  O  ASP C 326   N  LEU C 313
SHEET    4 AC7 4 GLN C 344  MET C 348 -1  O  GLU C 347   N  SER C 323
SHEET    1 AC8 4 HIS C 363  PHE C 364  0
SHEET    2 AC8 4 SER C 370  SER C 376 -1  O  TYR C 372   N  HIS C 363
SHEET    3 AC8 4 ARG C 382  GLN C 388 -1  O  CYS C 385   N  LYS C 373
SHEET    4 AC8 4 THR C 395  PHE C 396 -1  O  THR C 395   N  TYR C 386
SHEET    1 AC9 4 VAL C 404  LEU C 410  0
SHEET    2 AC9 4 TYR C 414  SER C 419 -1  O  TYR C 416   N  ALA C 409
SHEET    3 AC9 4 ASN C 430  GLN C 435 -1  O  TYR C 432   N  TYR C 417
SHEET    4 AC9 4 VAL C 442  CYS C 444 -1  O  THR C 443   N  LYS C 433
SHEET    1 AD1 4 TYR C 457  PHE C 461  0
SHEET    2 AD1 4 TYR C 467  CYS C 472 -1  O  ARG C 471   N  SER C 458
SHEET    3 AD1 4 LEU C 479  SER C 484 -1  O  HIS C 483   N  TYR C 468
SHEET    4 AD1 4 LYS C 489  GLU C 495 -1  O  LEU C 491   N  LEU C 482
SHEET    1 AD2 8 LYS C 513  LEU C 519  0
SHEET    2 AD2 8 THR C 522  ILE C 529 -1  O  THR C 522   N  LEU C 519
SHEET    3 AD2 8 ILE C 574  PHE C 578 -1  O  SER C 577   N  GLN C 527
SHEET    4 AD2 8 TYR C 540  ASP C 545  1  N  LEU C 543   O  ILE C 574
SHEET    5 AD2 8 VAL C 619  GLY C 628  1  O  ASP C 620   N  TYR C 540
SHEET    6 AD2 8 CYS C 649  VAL C 653  1  O  VAL C 653   N  GLY C 628
SHEET    7 AD2 8 GLU C 699  GLY C 705  1  O  LEU C 701   N  GLY C 650
SHEET    8 AD2 8 GLN C 731  TYR C 735  1  O  GLN C 731   N  TYR C 700
SSBOND   1 CYS A  387    CYS A  411                          1555   1555  2.03
SSBOND   2 CYS A  429    CYS A  482                          1555   1555  2.04
SSBOND   3 CYS A  441    CYS A  589                          1555   1555  2.03
SSBOND   4 CYS A  506    CYS A  530                          1555   1555  2.03
SSBOND   5 CYS B  328    CYS B  339                          1555   1555  2.03
SSBOND   6 CYS B  385    CYS B  394                          1555   1555  2.02
SSBOND   7 CYS B  444    CYS B  447                          1555   1555  2.03
SSBOND   8 CYS B  454    CYS B  472                          1555   1555  2.03
SSBOND   9 CYS B  649    CYS B  762                          1555   1555  2.04
SSBOND  10 CYS C  328    CYS C  339                          1555   1555  2.04
SSBOND  11 CYS C  385    CYS C  394                          1555   1555  2.04
SSBOND  12 CYS C  444    CYS C  447                          1555   1555  2.03
SSBOND  13 CYS C  454    CYS C  472                          1555   1555  2.03
SSBOND  14 CYS C  649    CYS C  762                          1555   1555  2.04
LINK         ND2 ASN B  85                 C1  NAG B 801     1555   1555  1.44
LINK         ND2 ASN B  92                 C1  NAG B 804     1555   1555  1.44
LINK         ND2 ASN B 150                 C1  NAG B 802     1555   1555  1.44
LINK         ND2 ASN B 219                 C1  NAG D   1     1555   1555  1.43
LINK         ND2 ASN B 229                 C1  NAG E   1     1555   1555  1.44
LINK         ND2 ASN B 281                 C1  NAG F   1     1555   1555  1.45
LINK         ND2 ASN B 321                 C1  NAG B 803     1555   1555  1.44
LINK         ND2 ASN C 150                 C1  NAG C 801     1555   1555  1.44
LINK         ND2 ASN C 229                 C1  NAG C 802     1555   1555  1.44
LINK         ND2 ASN C 685                 C1  NAG C 803     1555   1555  1.45
LINK         O4  NAG D   1                 C1  NAG D   2     1555   1555  1.45
LINK         O4  NAG E   1                 C1  NAG E   2     1555   1555  1.44
LINK         O4  NAG F   1                 C1  NAG F   2     1555   1555  1.45
CRYST1    1.000    1.000    1.000  90.00  90.00  90.00 P 1
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      1.000000  0.000000  0.000000        0.00000
SCALE2      0.000000  1.000000  0.000000        0.00000
SCALE3      0.000000  0.000000  1.000000        0.00000
TER    1587      MET A 591
TER    7530      SER B 764
TER   13473      SER C 764
MASTER      212    0   13   37  109    0    0    613652    3  220  130
END