longtext: 9iw9-pdb

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HEADER    HYDROLASE                               25-JUL-24   9IW9
TITLE     CRYSTAL STRUCTURE OF KBPETASE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: KBPETASE;
COMPND   3 CHAIN: A, B, C, D;
COMPND   4 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: KIBDELOSPORANGIUM BANGUIENSE;
SOURCE   3 ORGANISM_TAXID: 1365924;
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   5 EXPRESSION_SYSTEM_TAXID: 562
KEYWDS    PETASE, S9 FAMILY PEPTIDASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    B.H.WU
REVDAT   1   30-JUL-25 9IW9    0
JRNL        AUTH   B.WU,B.ZHONG,L.ZHENG,R.HUANG,S.JIANG,M.LI,L.HONG,P.TAN
JRNL        TITL   HARNESSING PROTEIN LANGUAGE MODEL FOR STRUCTURE-BASED
JRNL        TITL 2 DISCOVERY OF HIGHLY EFFICIENT AND ROBUST PET HYDROLASES.
JRNL        REF    NAT COMMUN                    V.  16  6211 2025
JRNL        REFN                   ESSN 2041-1723
JRNL        PMID   40617831
JRNL        DOI    10.1038/S41467-025-61599-Z
REMARK   2
REMARK   2 RESOLUTION.    1.75 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX 1.18.1_3865
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : GEOSTD + MONOMER LIBRARY + CDL V1.2
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.75
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 31.33
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.340
REMARK   3   COMPLETENESS FOR RANGE        (%) : 90.9
REMARK   3   NUMBER OF REFLECTIONS             : 164893
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.164
REMARK   3   R VALUE            (WORKING SET) : 0.164
REMARK   3   FREE R VALUE                     : 0.182
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 1.140
REMARK   3   FREE R VALUE TEST SET COUNT      : 1785
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 31.3300 -  4.1100    0.90    12352   141  0.1322 0.1316
REMARK   3     2  4.1100 -  3.2600    0.89    12236   131  0.1403 0.1608
REMARK   3     3  3.2600 -  2.8500    0.91    12474   141  0.1680 0.2270
REMARK   3     4  2.8500 -  2.5900    0.93    12821   172  0.1592 0.1879
REMARK   3     5  2.5900 -  2.4100    0.94    12912   152  0.1554 0.1799
REMARK   3     6  2.4100 -  2.2600    0.91    12503   140  0.1569 0.1773
REMARK   3     7  2.2600 -  2.1500    0.93    12751   148  0.1650 0.1495
REMARK   3     8  2.1500 -  2.0600    0.93    13020   129  0.1710 0.1815
REMARK   3     9  2.0600 -  1.9800    0.91    12459   150  0.1862 0.1800
REMARK   3    10  1.9800 -  1.9100    0.88    12101   148  0.2053 0.2349
REMARK   3    11  1.9100 -  1.8500    0.91    12532   133  0.2212 0.2280
REMARK   3    12  1.8500 -  1.8000    0.90    12387   161  0.2489 0.2891
REMARK   3    13  1.8000 -  1.7500    0.90    12467   132  0.2663 0.3014
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.187
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 18.942
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 17.88
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 20.82
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.014           7705
REMARK   3   ANGLE     :  1.598          10526
REMARK   3   CHIRALITY :  0.085           1115
REMARK   3   PLANARITY :  0.010           1398
REMARK   3   DIHEDRAL  : 13.739           2590
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 9IW9 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBC ON 30-JUL-24.
REMARK 100 THE DEPOSITION ID IS D_1300049815.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 25-MAR-24
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5-7.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SSRF
REMARK 200  BEAMLINE                       : BL02U1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9792
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS EIGER2 S 9M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XDS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 164893
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.750
REMARK 200  RESOLUTION RANGE LOW       (A) : 31.330
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 97.0
REMARK 200  DATA REDUNDANCY                : 2.900
REMARK 200  R MERGE                    (I) : 0.06559
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 13.4200
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.75
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.81
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : 0.54430
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 3.540
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 43.73
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.19
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 28-34% POLYETHYLENE GLYCOL 3000, 0.1 M
REMARK 280  HEPES, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 291K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       52.39900
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2, 3, 4
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 3
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 4
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     ALA A     2
REMARK 465     MET B     1
REMARK 465     ALA B     2
REMARK 465     MET C     1
REMARK 465     ALA C     2
REMARK 465     PHE C     3
REMARK 465     ALA C     4
REMARK 465     MET D     1
REMARK 465     ALA D     2
REMARK 465     PHE D     3
REMARK 465     ALA D     4
REMARK 480
REMARK 480 ZERO OCCUPANCY ATOM
REMARK 480 THE FOLLOWING RESIDUES HAVE ATOMS MODELED WITH ZERO
REMARK 480 OCCUPANCY. THE LOCATION AND PROPERTIES OF THESE ATOMS
REMARK 480 MAY NOT BE RELIABLE. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 480 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 480   M RES C SSEQI ATOMS
REMARK 480     CYS A  239   N    CA   C    O    CB   SG
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    HOH B   401     O    HOH B   466              2.10
REMARK 500   OG   SER A    27     O    HOH A   301              2.12
REMARK 500   OD2  ASP D   245     O    HOH D   301              2.13
REMARK 500   O    HOH A   308     O    HOH A   426              2.15
REMARK 500   O    HOH C   417     O    HOH D   450              2.16
REMARK 500   O    HOH C   312     O    HOH C   472              2.16
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O    HOH A   331     O    HOH D   453     1655     2.17
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    SER A 108      -21.84   -141.78
REMARK 500    SER A 128     -124.95     65.59
REMARK 500    HIS A 184      -84.55   -117.81
REMARK 500    SER B 128     -125.09     65.20
REMARK 500    ALA B 179       62.42   -119.96
REMARK 500    HIS B 184      -85.23   -120.08
REMARK 500    SER C 128     -125.27     65.57
REMARK 500    HIS C 184      -79.85   -125.55
REMARK 500    SER D 108      -25.21   -143.66
REMARK 500    SER D 128     -122.07     67.70
REMARK 500    HIS D 184      -86.19   -129.78
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 492        DISTANCE =  5.84 ANGSTROMS
REMARK 525    HOH C 519        DISTANCE =  6.12 ANGSTROMS
REMARK 525    HOH C 520        DISTANCE =  6.30 ANGSTROMS
REMARK 525    HOH C 521        DISTANCE =  6.32 ANGSTROMS
REMARK 525    HOH D 518        DISTANCE =  6.42 ANGSTROMS
REMARK 525    HOH D 519        DISTANCE =  6.74 ANGSTROMS
DBREF  9IW9 A    1   254  PDB    9IW9     9IW9             1    254
DBREF  9IW9 B    1   254  PDB    9IW9     9IW9             1    254
DBREF  9IW9 C    1   254  PDB    9IW9     9IW9             1    254
DBREF  9IW9 D    1   254  PDB    9IW9     9IW9             1    254
SEQRES   1 A  254  MET ALA PHE ALA ASP GLU ILE GLY GLN ALA PRO THR ALA
SEQRES   2 A  254  ALA ASN ILE THR GLY ASP GLY SER PHE SER THR SER SER
SEQRES   3 A  254  SER ALA ILE PHE GLY GLN SER GLY PHE GLY GLY GLY THR
SEQRES   4 A  254  ALA TYR TYR PRO ASN THR ALA GLY LYS TYR PRO VAL VAL
SEQRES   5 A  254  ALA PHE ALA PRO GLY PHE LEU SER ASP TRP ASN ALA LEU
SEQRES   6 A  254  ASN TRP LEU GLY PRO ARG VAL ALA SER TRP GLY PHE VAL
SEQRES   7 A  254  VAL VAL GLY VAL ASN THR ASN THR PRO PHE ASP PHE PRO
SEQRES   8 A  254  ASP ALA ARG GLY ASP GLN LEU LEU ALA ALA LEU ASN TRP
SEQRES   9 A  254  ALA VAL ASN SER ALA PRO ALA ALA VAL ARG ASP LYS ALA
SEQRES  10 A  254  ASP GLY SER ARG ARG GLY VAL SER GLY TRP SER MET GLY
SEQRES  11 A  254  GLY GLY GLY THR LEU GLU ALA LEU ALA LYS ASP THR THR
SEQRES  12 A  254  GLY THR VAL LYS ALA GLY VAL PRO LEU ALA PRO TRP HIS
SEQRES  13 A  254  THR ASN LYS THR TRP SER LYS VAL THR GLU PRO VAL PHE
SEQRES  14 A  254  ILE ILE GLY GLY GLN ASN ASP SER VAL ALA ALA PRO ALA
SEQRES  15 A  254  SER HIS ALA ILE PRO PHE TYR ASN SER LEU GLY GLY PRO
SEQRES  16 A  254  LYS SER TYR LEU GLU ARG ALA GLY ALA ASP HIS PHE PHE
SEQRES  17 A  254  PRO THR LYS SER ASN GLY THR VAL SER ARG ALA VAL VAL
SEQRES  18 A  254  SER PHE PHE LYS ARG HIS VAL SER ALA ASP THR ARG PHE
SEQRES  19 A  254  THR PRO PHE LEU CYS GLY PHE SER GLY LEU ASP VAL SER
SEQRES  20 A  254  ASN PHE ARG SER ASN SER CYS
SEQRES   1 B  254  MET ALA PHE ALA ASP GLU ILE GLY GLN ALA PRO THR ALA
SEQRES   2 B  254  ALA ASN ILE THR GLY ASP GLY SER PHE SER THR SER SER
SEQRES   3 B  254  SER ALA ILE PHE GLY GLN SER GLY PHE GLY GLY GLY THR
SEQRES   4 B  254  ALA TYR TYR PRO ASN THR ALA GLY LYS TYR PRO VAL VAL
SEQRES   5 B  254  ALA PHE ALA PRO GLY PHE LEU SER ASP TRP ASN ALA LEU
SEQRES   6 B  254  ASN TRP LEU GLY PRO ARG VAL ALA SER TRP GLY PHE VAL
SEQRES   7 B  254  VAL VAL GLY VAL ASN THR ASN THR PRO PHE ASP PHE PRO
SEQRES   8 B  254  ASP ALA ARG GLY ASP GLN LEU LEU ALA ALA LEU ASN TRP
SEQRES   9 B  254  ALA VAL ASN SER ALA PRO ALA ALA VAL ARG ASP LYS ALA
SEQRES  10 B  254  ASP GLY SER ARG ARG GLY VAL SER GLY TRP SER MET GLY
SEQRES  11 B  254  GLY GLY GLY THR LEU GLU ALA LEU ALA LYS ASP THR THR
SEQRES  12 B  254  GLY THR VAL LYS ALA GLY VAL PRO LEU ALA PRO TRP HIS
SEQRES  13 B  254  THR ASN LYS THR TRP SER LYS VAL THR GLU PRO VAL PHE
SEQRES  14 B  254  ILE ILE GLY GLY GLN ASN ASP SER VAL ALA ALA PRO ALA
SEQRES  15 B  254  SER HIS ALA ILE PRO PHE TYR ASN SER LEU GLY GLY PRO
SEQRES  16 B  254  LYS SER TYR LEU GLU ARG ALA GLY ALA ASP HIS PHE PHE
SEQRES  17 B  254  PRO THR LYS SER ASN GLY THR VAL SER ARG ALA VAL VAL
SEQRES  18 B  254  SER PHE PHE LYS ARG HIS VAL SER ALA ASP THR ARG PHE
SEQRES  19 B  254  THR PRO PHE LEU CYS GLY PHE SER GLY LEU ASP VAL SER
SEQRES  20 B  254  ASN PHE ARG SER ASN SER CYS
SEQRES   1 C  254  MET ALA PHE ALA ASP GLU ILE GLY GLN ALA PRO THR ALA
SEQRES   2 C  254  ALA ASN ILE THR GLY ASP GLY SER PHE SER THR SER SER
SEQRES   3 C  254  SER ALA ILE PHE GLY GLN SER GLY PHE GLY GLY GLY THR
SEQRES   4 C  254  ALA TYR TYR PRO ASN THR ALA GLY LYS TYR PRO VAL VAL
SEQRES   5 C  254  ALA PHE ALA PRO GLY PHE LEU SER ASP TRP ASN ALA LEU
SEQRES   6 C  254  ASN TRP LEU GLY PRO ARG VAL ALA SER TRP GLY PHE VAL
SEQRES   7 C  254  VAL VAL GLY VAL ASN THR ASN THR PRO PHE ASP PHE PRO
SEQRES   8 C  254  ASP ALA ARG GLY ASP GLN LEU LEU ALA ALA LEU ASN TRP
SEQRES   9 C  254  ALA VAL ASN SER ALA PRO ALA ALA VAL ARG ASP LYS ALA
SEQRES  10 C  254  ASP GLY SER ARG ARG GLY VAL SER GLY TRP SER MET GLY
SEQRES  11 C  254  GLY GLY GLY THR LEU GLU ALA LEU ALA LYS ASP THR THR
SEQRES  12 C  254  GLY THR VAL LYS ALA GLY VAL PRO LEU ALA PRO TRP HIS
SEQRES  13 C  254  THR ASN LYS THR TRP SER LYS VAL THR GLU PRO VAL PHE
SEQRES  14 C  254  ILE ILE GLY GLY GLN ASN ASP SER VAL ALA ALA PRO ALA
SEQRES  15 C  254  SER HIS ALA ILE PRO PHE TYR ASN SER LEU GLY GLY PRO
SEQRES  16 C  254  LYS SER TYR LEU GLU ARG ALA GLY ALA ASP HIS PHE PHE
SEQRES  17 C  254  PRO THR LYS SER ASN GLY THR VAL SER ARG ALA VAL VAL
SEQRES  18 C  254  SER PHE PHE LYS ARG HIS VAL SER ALA ASP THR ARG PHE
SEQRES  19 C  254  THR PRO PHE LEU CYS GLY PHE SER GLY LEU ASP VAL SER
SEQRES  20 C  254  ASN PHE ARG SER ASN SER CYS
SEQRES   1 D  254  MET ALA PHE ALA ASP GLU ILE GLY GLN ALA PRO THR ALA
SEQRES   2 D  254  ALA ASN ILE THR GLY ASP GLY SER PHE SER THR SER SER
SEQRES   3 D  254  SER ALA ILE PHE GLY GLN SER GLY PHE GLY GLY GLY THR
SEQRES   4 D  254  ALA TYR TYR PRO ASN THR ALA GLY LYS TYR PRO VAL VAL
SEQRES   5 D  254  ALA PHE ALA PRO GLY PHE LEU SER ASP TRP ASN ALA LEU
SEQRES   6 D  254  ASN TRP LEU GLY PRO ARG VAL ALA SER TRP GLY PHE VAL
SEQRES   7 D  254  VAL VAL GLY VAL ASN THR ASN THR PRO PHE ASP PHE PRO
SEQRES   8 D  254  ASP ALA ARG GLY ASP GLN LEU LEU ALA ALA LEU ASN TRP
SEQRES   9 D  254  ALA VAL ASN SER ALA PRO ALA ALA VAL ARG ASP LYS ALA
SEQRES  10 D  254  ASP GLY SER ARG ARG GLY VAL SER GLY TRP SER MET GLY
SEQRES  11 D  254  GLY GLY GLY THR LEU GLU ALA LEU ALA LYS ASP THR THR
SEQRES  12 D  254  GLY THR VAL LYS ALA GLY VAL PRO LEU ALA PRO TRP HIS
SEQRES  13 D  254  THR ASN LYS THR TRP SER LYS VAL THR GLU PRO VAL PHE
SEQRES  14 D  254  ILE ILE GLY GLY GLN ASN ASP SER VAL ALA ALA PRO ALA
SEQRES  15 D  254  SER HIS ALA ILE PRO PHE TYR ASN SER LEU GLY GLY PRO
SEQRES  16 D  254  LYS SER TYR LEU GLU ARG ALA GLY ALA ASP HIS PHE PHE
SEQRES  17 D  254  PRO THR LYS SER ASN GLY THR VAL SER ARG ALA VAL VAL
SEQRES  18 D  254  SER PHE PHE LYS ARG HIS VAL SER ALA ASP THR ARG PHE
SEQRES  19 D  254  THR PRO PHE LEU CYS GLY PHE SER GLY LEU ASP VAL SER
SEQRES  20 D  254  ASN PHE ARG SER ASN SER CYS
FORMUL   5  HOH   *823(H2 O)
HELIX    1 AA1 ASP A   61  ASN A   66  5                                   6
HELIX    2 AA2 TRP A   67  SER A   74  1                                   8
HELIX    3 AA3 PHE A   90  SER A  108  1                                  19
HELIX    4 AA4 PRO A  110  ASP A  115  1                                   6
HELIX    5 AA5 MET A  129  ASP A  141  1                                  13
HELIX    6 AA6 HIS A  184  LEU A  192  1                                   9
HELIX    7 AA7 PHE A  207  LYS A  211  5                                   5
HELIX    8 AA8 ASN A  213  VAL A  228  1                                  16
HELIX    9 AA9 ASP A  231  CYS A  239  5                                   9
HELIX   10 AB1 ASP B   61  ASN B   66  5                                   6
HELIX   11 AB2 TRP B   67  SER B   74  1                                   8
HELIX   12 AB3 PHE B   90  SER B  108  1                                  19
HELIX   13 AB4 PRO B  110  ASP B  115  1                                   6
HELIX   14 AB5 SER B  128  ASP B  141  1                                  14
HELIX   15 AB6 HIS B  184  LEU B  192  1                                   9
HELIX   16 AB7 PHE B  207  LYS B  211  5                                   5
HELIX   17 AB8 ASN B  213  VAL B  228  1                                  16
HELIX   18 AB9 ASP B  231  CYS B  239  5                                   9
HELIX   19 AC1 ASP C   61  ASN C   66  5                                   6
HELIX   20 AC2 TRP C   67  SER C   74  1                                   8
HELIX   21 AC3 PHE C   90  SER C  108  1                                  19
HELIX   22 AC4 PRO C  110  ASP C  115  1                                   6
HELIX   23 AC5 SER C  128  ASP C  141  1                                  14
HELIX   24 AC6 HIS C  184  LEU C  192  1                                   9
HELIX   25 AC7 PHE C  207  LYS C  211  5                                   5
HELIX   26 AC8 ASN C  213  VAL C  228  1                                  16
HELIX   27 AC9 ASP C  231  CYS C  239  5                                   9
HELIX   28 AD1 ASP D   61  ASN D   66  5                                   6
HELIX   29 AD2 TRP D   67  SER D   74  1                                   8
HELIX   30 AD3 PHE D   90  SER D  108  1                                  19
HELIX   31 AD4 PRO D  110  ASP D  115  1                                   6
HELIX   32 AD5 SER D  128  ASP D  141  1                                  14
HELIX   33 AD6 HIS D  184  LEU D  192  1                                   9
HELIX   34 AD7 PHE D  207  LYS D  211  5                                   5
HELIX   35 AD8 ASN D  213  VAL D  228  1                                  16
HELIX   36 AD9 ASP D  231  CYS D  239  5                                   9
SHEET    1 AA1 6 THR A  24  ILE A  29  0
SHEET    2 AA1 6 GLY A  37  PRO A  43 -1  O  ALA A  40   N  SER A  27
SHEET    3 AA1 6 VAL A  78  ASN A  83 -1  O  GLY A  81   N  THR A  39
SHEET    4 AA1 6 TYR A  49  ALA A  55  1  N  VAL A  52   O  VAL A  78
SHEET    5 AA1 6 ALA A 117  SER A 128  1  O  ASP A 118   N  TYR A  49
SHEET    6 AA1 6 VAL A 146  PRO A 154  1  O  LEU A 152   N  GLY A 126
SHEET    1 AA2 3 VAL A 168  GLY A 173  0
SHEET    2 AA2 3 LYS A 196  ARG A 201  1  O  SER A 197   N  ILE A 170
SHEET    3 AA2 3 VAL A 246  SER A 251 -1  O  ARG A 250   N  TYR A 198
SHEET    1 AA3 6 THR B  24  ALA B  28  0
SHEET    2 AA3 6 GLY B  37  PRO B  43 -1  O  ALA B  40   N  SER B  27
SHEET    3 AA3 6 VAL B  78  ASN B  83 -1  O  GLY B  81   N  THR B  39
SHEET    4 AA3 6 TYR B  49  ALA B  55  1  N  VAL B  52   O  VAL B  78
SHEET    5 AA3 6 ALA B 117  TRP B 127  1  O  ASP B 118   N  TYR B  49
SHEET    6 AA3 6 VAL B 146  LEU B 152  1  O  LEU B 152   N  GLY B 126
SHEET    1 AA4 3 VAL B 168  GLY B 173  0
SHEET    2 AA4 3 LYS B 196  ARG B 201  1  O  ARG B 201   N  GLY B 172
SHEET    3 AA4 3 VAL B 246  SER B 251 -1  O  ARG B 250   N  TYR B 198
SHEET    1 AA5 6 THR C  24  ALA C  28  0
SHEET    2 AA5 6 GLY C  37  PRO C  43 -1  O  ALA C  40   N  SER C  27
SHEET    3 AA5 6 VAL C  78  ASN C  83 -1  O  GLY C  81   N  THR C  39
SHEET    4 AA5 6 TYR C  49  ALA C  55  1  N  VAL C  52   O  VAL C  78
SHEET    5 AA5 6 ALA C 117  TRP C 127  1  O  ASP C 118   N  TYR C  49
SHEET    6 AA5 6 VAL C 146  LEU C 152  1  O  LEU C 152   N  GLY C 126
SHEET    1 AA6 3 VAL C 168  GLY C 173  0
SHEET    2 AA6 3 LYS C 196  ARG C 201  1  O  ARG C 201   N  GLY C 172
SHEET    3 AA6 3 VAL C 246  SER C 251 -1  O  ASN C 248   N  GLU C 200
SHEET    1 AA7 6 THR D  24  ILE D  29  0
SHEET    2 AA7 6 GLY D  37  PRO D  43 -1  O  ALA D  40   N  SER D  27
SHEET    3 AA7 6 VAL D  78  ASN D  83 -1  O  GLY D  81   N  THR D  39
SHEET    4 AA7 6 TYR D  49  ALA D  55  1  N  VAL D  52   O  VAL D  78
SHEET    5 AA7 6 ALA D 117  TRP D 127  1  O  ASP D 118   N  TYR D  49
SHEET    6 AA7 6 VAL D 146  LEU D 152  1  O  LEU D 152   N  GLY D 126
SHEET    1 AA8 3 VAL D 168  GLY D 173  0
SHEET    2 AA8 3 LYS D 196  ARG D 201  1  O  ARG D 201   N  GLY D 172
SHEET    3 AA8 3 VAL D 246  SER D 251 -1  O  ASN D 248   N  GLU D 200
SSBOND   1 CYS A  239    CYS A  254                          1555   1555  2.76
SSBOND   2 CYS B  239    CYS B  254                          1555   1555  2.08
SSBOND   3 CYS C  239    CYS C  254                          1555   1555  2.14
SSBOND   4 CYS D  239    CYS D  254                          1555   1555  2.12
CRYST1   62.496  104.798   71.020  90.00  93.83  90.00 P 1 21 1      8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.016001  0.000000  0.001071        0.00000
SCALE2      0.000000  0.009542  0.000000        0.00000
SCALE3      0.000000  0.000000  0.014112        0.00000
TER    1880      CYS A 254
TER    3753      CYS B 254
TER    5611      CYS C 254
TER    7477      CYS D 254
MASTER      319    0    0   36   36    0    0    6 8251    4    8   80
END