longtext: 9lbt-pdb

content
HEADER    PEPTIDE BINDING PROTEIN                 03-JAN-25   9LBT
TITLE     DPPIV-VAMP
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: DIPEPTIDYL PEPTIDASE 4 SOLUBLE FORM;
COMPND   3 CHAIN: A, B;
COMPND   4 SYNONYM: DIPEPTIDYL PEPTIDASE IV SOLUBLE FORM;
COMPND   5 ENGINEERED: YES;
COMPND   6 MOL_ID: 2;
COMPND   7 MOLECULE: VAL-ALA-MET-PRO;
COMPND   8 CHAIN: C, D;
COMPND   9 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: DPP4, ADCP2, CD26;
SOURCE   6 EXPRESSION_SYSTEM: UNIDENTIFIED BACULOVIRUS;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 10469;
SOURCE   8 MOL_ID: 2;
SOURCE   9 SYNTHETIC: YES;
SOURCE  10 ORGANISM_SCIENTIFIC: CANNABIS SATIVA;
SOURCE  11 ORGANISM_TAXID: 3483
KEYWDS    DPP-IV, VAMP, PEPTIDE BINDING PROTEIN
EXPDTA    X-RAY DIFFRACTION
AUTHOR    H.H.CHEN,X.H.XING
REVDAT   1   23-JUL-25 9LBT    0
JRNL        AUTH   H.H.CHEN,X.H.XING
JRNL        TITL   DISCOVERY OF AN ORAL-ADMISTRATED BIOPEPTIDE VAMP: A NOVEL
JRNL        TITL 2 GLUCOSE HOMEOSTASIS MODULATOR WITH BIFUNCTIONALITIES OF
JRNL        TITL 3 TARGETING INTESTINAL DPP-IV AND MICROBIOTA
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    1.99 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (1.20.1_4487: ???)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.99
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 32.41
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.360
REMARK   3   COMPLETENESS FOR RANGE        (%) : 98.7
REMARK   3   NUMBER OF REFLECTIONS             : 138313
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.169
REMARK   3   R VALUE            (WORKING SET) : 0.169
REMARK   3   FREE R VALUE                     : 0.195
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 1.450
REMARK   3   FREE R VALUE TEST SET COUNT      : 2003
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1  3.8000 -  3.3200    1.00     9934   140  0.1560 0.1777
REMARK   3     2  3.3200 -  3.0200    1.00     9861   147  0.1639 0.2005
REMARK   3     3  3.0200 -  2.8000    1.00     9851   145  0.1717 0.1954
REMARK   3     4  2.8000 -  2.6400    0.99     9792   139  0.1697 0.2251
REMARK   3     5  2.6400 -  2.5100    0.99     9735   143  0.1698 0.2258
REMARK   3     6  2.5100 -  2.4000    0.99     9713   150  0.1644 0.1962
REMARK   3     7  2.4000 -  2.3100    0.99     9710   140  0.1665 0.2025
REMARK   3     8  2.3100 -  2.2300    0.99     9663   147  0.1764 0.2025
REMARK   3     9  2.2300 -  2.1600    0.98     9648   133  0.1789 0.2278
REMARK   3    10  2.1600 -  2.0900    0.98     9608   146  0.1885 0.2435
REMARK   3    11  2.0900 -  2.0400    0.98     9582   156  0.2127 0.2492
REMARK   3    12  2.0400 -  1.9900    0.91     8920   120  0.2530 0.3453
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.10
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.210
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 18.030
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :   NULL           NULL
REMARK   3   ANGLE     :   NULL           NULL
REMARK   3   CHIRALITY :  0.083           1756
REMARK   3   PLANARITY :  0.013           2110
REMARK   3   DIHEDRAL  :  6.862           1618
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 11
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 1 THROUGH 59 )
REMARK   3    ORIGIN FOR THE GROUP (A):  -8.0528  -6.2919 -55.0338
REMARK   3    T TENSOR
REMARK   3      T11:   0.3441 T22:   0.2614
REMARK   3      T33:   0.2985 T12:  -0.0153
REMARK   3      T13:  -0.0051 T23:   0.0103
REMARK   3    L TENSOR
REMARK   3      L11:   0.6127 L22:   0.2841
REMARK   3      L33:   0.3936 L12:  -0.2388
REMARK   3      L13:   0.0305 L23:   0.2390
REMARK   3    S TENSOR
REMARK   3      S11:   0.1636 S12:   0.1535 S13:  -0.2890
REMARK   3      S21:  -0.0462 S22:  -0.0839 S23:   0.0845
REMARK   3      S31:   0.3106 S32:   0.0079 S33:  -0.0293
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 60 THROUGH 255 )
REMARK   3    ORIGIN FOR THE GROUP (A): -14.3526  -8.1162 -28.4658
REMARK   3    T TENSOR
REMARK   3      T11:   0.2331 T22:   0.1747
REMARK   3      T33:   0.2833 T12:  -0.0059
REMARK   3      T13:  -0.0413 T23:   0.0844
REMARK   3    L TENSOR
REMARK   3      L11:   1.1272 L22:   0.3026
REMARK   3      L33:   0.5154 L12:  -0.0199
REMARK   3      L13:   0.0971 L23:   0.0327
REMARK   3    S TENSOR
REMARK   3      S11:   0.0873 S12:  -0.2077 S13:  -0.3582
REMARK   3      S21:   0.0448 S22:  -0.0386 S23:  -0.0741
REMARK   3      S31:   0.1487 S32:   0.0198 S33:  -0.0095
REMARK   3   TLS GROUP : 3
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 256 THROUGH 387 )
REMARK   3    ORIGIN FOR THE GROUP (A):   1.1993  13.4422 -28.6936
REMARK   3    T TENSOR
REMARK   3      T11:   0.1794 T22:   0.1912
REMARK   3      T33:   0.1806 T12:  -0.0069
REMARK   3      T13:  -0.0312 T23:   0.0116
REMARK   3    L TENSOR
REMARK   3      L11:   1.4764 L22:   0.5627
REMARK   3      L33:   0.6712 L12:   0.1910
REMARK   3      L13:   0.1510 L23:   0.1012
REMARK   3    S TENSOR
REMARK   3      S11:   0.0163 S12:  -0.2627 S13:  -0.0046
REMARK   3      S21:   0.1277 S22:  -0.0435 S23:  -0.0619
REMARK   3      S31:  -0.0557 S32:   0.1242 S33:  -0.0069
REMARK   3   TLS GROUP : 4
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 388 THROUGH 499 )
REMARK   3    ORIGIN FOR THE GROUP (A):  -4.0985  15.7446 -55.4596
REMARK   3    T TENSOR
REMARK   3      T11:   0.2088 T22:   0.1993
REMARK   3      T33:   0.1669 T12:  -0.0348
REMARK   3      T13:   0.0040 T23:   0.0421
REMARK   3    L TENSOR
REMARK   3      L11:   1.4660 L22:   0.4443
REMARK   3      L33:   0.8581 L12:  -0.2372
REMARK   3      L13:   0.2411 L23:  -0.1282
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0164 S12:   0.2307 S13:   0.1721
REMARK   3      S21:  -0.0269 S22:  -0.0383 S23:  -0.0627
REMARK   3      S31:  -0.1047 S32:   0.0982 S33:   0.0146
REMARK   3   TLS GROUP : 5
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 500 THROUGH 616 )
REMARK   3    ORIGIN FOR THE GROUP (A): -23.7815  24.4714 -46.7476
REMARK   3    T TENSOR
REMARK   3      T11:   0.2044 T22:   0.1507
REMARK   3      T33:   0.2072 T12:   0.0139
REMARK   3      T13:  -0.0023 T23:   0.0294
REMARK   3    L TENSOR
REMARK   3      L11:   0.9865 L22:   0.3910
REMARK   3      L33:   0.5635 L12:   0.0974
REMARK   3      L13:   0.0672 L23:   0.1000
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0305 S12:   0.0944 S13:   0.2200
REMARK   3      S21:  -0.0138 S22:   0.0306 S23:   0.0112
REMARK   3      S31:  -0.1730 S32:  -0.0426 S33:   0.0117
REMARK   3   TLS GROUP : 6
REMARK   3    SELECTION: CHAIN 'A' AND (RESID 617 THROUGH 723 )
REMARK   3    ORIGIN FOR THE GROUP (A): -33.7352  16.4834 -38.8160
REMARK   3    T TENSOR
REMARK   3      T11:   0.1422 T22:   0.1163
REMARK   3      T33:   0.1478 T12:   0.0180
REMARK   3      T13:   0.0103 T23:   0.0165
REMARK   3    L TENSOR
REMARK   3      L11:   1.1673 L22:   0.6824
REMARK   3      L33:   0.6912 L12:   0.1705
REMARK   3      L13:   0.0403 L23:   0.0548
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0011 S12:  -0.0091 S13:   0.1255
REMARK   3      S21:   0.0131 S22:   0.0016 S23:   0.0712
REMARK   3      S31:  -0.0825 S32:  -0.0781 S33:   0.0053
REMARK   3   TLS GROUP : 7
REMARK   3    SELECTION: CHAIN 'B' AND (RESID 1 THROUGH 59 )
REMARK   3    ORIGIN FOR THE GROUP (A): -75.9412  22.9238 -24.3147
REMARK   3    T TENSOR
REMARK   3      T11:   0.3172 T22:   0.2425
REMARK   3      T33:   0.3603 T12:  -0.0340
REMARK   3      T13:  -0.0468 T23:  -0.0159
REMARK   3    L TENSOR
REMARK   3      L11:   0.7972 L22:   0.4161
REMARK   3      L33:   0.4461 L12:  -0.0322
REMARK   3      L13:  -0.0860 L23:  -0.1910
REMARK   3    S TENSOR
REMARK   3      S11:   0.2022 S12:  -0.2786 S13:   0.3298
REMARK   3      S21:   0.2939 S22:  -0.1200 S23:  -0.2167
REMARK   3      S31:  -0.3841 S32:   0.2154 S33:   0.0266
REMARK   3   TLS GROUP : 8
REMARK   3    SELECTION: CHAIN 'B' AND (RESID 60 THROUGH 255 )
REMARK   3    ORIGIN FOR THE GROUP (A): -62.7160   1.0759 -13.3780
REMARK   3    T TENSOR
REMARK   3      T11:   0.1911 T22:   0.1689
REMARK   3      T33:   0.1610 T12:   0.0011
REMARK   3      T13:  -0.0231 T23:   0.0100
REMARK   3    L TENSOR
REMARK   3      L11:   0.9539 L22:   0.3951
REMARK   3      L33:   0.9401 L12:   0.3496
REMARK   3      L13:  -0.4626 L23:  -0.3353
REMARK   3    S TENSOR
REMARK   3      S11:   0.0738 S12:  -0.0834 S13:   0.0944
REMARK   3      S21:   0.1050 S22:  -0.0423 S23:  -0.0230
REMARK   3      S31:  -0.1420 S32:  -0.0608 S33:   0.0407
REMARK   3   TLS GROUP : 9
REMARK   3    SELECTION: CHAIN 'B' AND (RESID 256 THROUGH 723 )
REMARK   3    ORIGIN FOR THE GROUP (A): -71.1664   0.7886 -40.6957
REMARK   3    T TENSOR
REMARK   3      T11:   0.1345 T22:   0.1720
REMARK   3      T33:   0.1341 T12:  -0.0038
REMARK   3      T13:  -0.0267 T23:   0.0089
REMARK   3    L TENSOR
REMARK   3      L11:   1.0145 L22:   0.3148
REMARK   3      L33:   0.2932 L12:   0.1173
REMARK   3      L13:  -0.1973 L23:  -0.0399
REMARK   3    S TENSOR
REMARK   3      S11:   0.0008 S12:   0.1877 S13:   0.0149
REMARK   3      S21:  -0.0576 S22:   0.0139 S23:   0.0283
REMARK   3      S31:   0.0355 S32:  -0.0776 S33:  -0.0087
REMARK   3   TLS GROUP : 10
REMARK   3    SELECTION: CHAIN 'C' AND (RESID 1 THROUGH 4 )
REMARK   3    ORIGIN FOR THE GROUP (A): -21.8455  11.9388 -39.5546
REMARK   3    T TENSOR
REMARK   3      T11:   0.4421 T22:   0.9406
REMARK   3      T33:   0.5561 T12:  -0.0119
REMARK   3      T13:   0.0878 T23:  -0.0767
REMARK   3    L TENSOR
REMARK   3      L11:   0.1420 L22:   0.3136
REMARK   3      L33:   0.1363 L12:  -0.0761
REMARK   3      L13:   0.1320 L23:  -0.0094
REMARK   3    S TENSOR
REMARK   3      S11:   0.0496 S12:   0.0126 S13:  -0.1394
REMARK   3      S21:  -0.0873 S22:   0.0546 S23:   0.0141
REMARK   3      S31:   0.0899 S32:   0.0035 S33:  -0.0616
REMARK   3   TLS GROUP : 11
REMARK   3    SELECTION: CHAIN 'D' AND (RESID 1 THROUGH 4 )
REMARK   3    ORIGIN FOR THE GROUP (A): -62.4434   2.4361 -37.0719
REMARK   3    T TENSOR
REMARK   3      T11:   0.3086 T22:   0.3696
REMARK   3      T33:   0.5491 T12:   0.0398
REMARK   3      T13:   0.1036 T23:   0.0865
REMARK   3    L TENSOR
REMARK   3      L11:   1.1715 L22:   4.3888
REMARK   3      L33:   0.6255 L12:   1.2511
REMARK   3      L13:  -0.6880 L23:   0.0877
REMARK   3    S TENSOR
REMARK   3      S11:   0.0717 S12:   0.1249 S13:   0.1001
REMARK   3      S21:  -0.0926 S22:   0.0075 S23:   0.2216
REMARK   3      S31:  -0.0694 S32:  -0.1322 S33:  -0.0614
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 9LBT COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBC ON 11-JAN-25.
REMARK 100 THE DEPOSITION ID IS D_1300055307.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 07-MAY-24
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SSRF
REMARK 200  BEAMLINE                       : BL02U1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9791
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS EIGER2 S 9M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 138319
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.990
REMARK 200  RESOLUTION RANGE LOW       (A) : 32.410
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.7
REMARK 200  DATA REDUNDANCY                : 11.60
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 14.9300
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.99
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.06
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHENIX
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 59.33
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.02
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.25 M AMMONIUM ACETATE, 0.1 M TRIS PH
REMARK 280  8.5, 21% W/V POLYETHYLENE GLYCOL 3350, VAPOR DIFFUSION, SITTING
REMARK 280  DROP, TEMPERATURE 296K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       58.91350
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       68.43350
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       63.18500
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       68.43350
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       58.91350
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       63.18500
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: TETRAMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: TETRAMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 5800 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 56880 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -27.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLN A    32A
REMARK 465     GLU A    32B
REMARK 465     ASN A    32C
REMARK 465     GLN B    32A
REMARK 465     GLU B    32B
REMARK 465     ASN B    32C
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     ASN A  33    N
REMARK 470     ASN B  33    N
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    HOH A  1305     O    HOH A  1437              2.05
REMARK 500   OE2  GLU B   104     O    HOH B   801              2.14
REMARK 500   O    SER B   233     N    SER B   235              2.16
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   CG1  VAL B   237     O    HOH A   840     2455     1.89
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    PRO C   4   C   -  N   -  CA  ANGL. DEV. = -13.6 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    SER A  25     -169.22   -165.22
REMARK 500    GLN A  81      -90.06   -105.56
REMARK 500    TRP A  82     -151.34    -99.77
REMARK 500    HIS A 120       32.63   -150.61
REMARK 500    GLU A 149      127.19    -39.84
REMARK 500    ILE A 151      -62.33   -125.79
REMARK 500    SER A 200     -164.71     66.88
REMARK 500    GLN A 278       31.89    -86.31
REMARK 500    ASN A 408       67.99   -163.94
REMARK 500    GLU A 479       16.69     57.77
REMARK 500    ARG A 555       42.96   -143.91
REMARK 500    THR A 558      -76.87   -119.71
REMARK 500    SER A 588     -116.87     67.44
REMARK 500    ALA A 612       55.86     39.14
REMARK 500    ASP A 636      -81.35   -113.41
REMARK 500    ASN A 637       22.59   -140.39
REMARK 500    ASN A 668      -67.48    -99.82
REMARK 500    ILE A 700       57.60     36.73
REMARK 500    SER B  64      116.14   -160.91
REMARK 500    GLN B  81      -90.16   -108.03
REMARK 500    TRP B  82     -150.67   -102.38
REMARK 500    HIS B 120       28.50   -148.56
REMARK 500    ILE B 151      -56.35   -128.02
REMARK 500    SER B 200     -166.19     67.62
REMARK 500    LEU B 234       55.93    -51.21
REMARK 500    SER B 235       19.39   -147.00
REMARK 500    VAL B 237       44.41     -1.08
REMARK 500    THR B 238        2.19     46.59
REMARK 500    GLN B 278       43.34    -85.85
REMARK 500    LYS B 381       19.71     55.51
REMARK 500    ASP B 396       96.86   -160.29
REMARK 500    ASN B 408       81.27   -153.76
REMARK 500    ASN B 445       24.17   -144.89
REMARK 500    TYR B 505      -65.08   -120.39
REMARK 500    ARG B 555       44.97   -142.13
REMARK 500    THR B 558      -81.52   -120.57
REMARK 500    SER B 588     -117.32     67.18
REMARK 500    ALA B 612       56.50     36.03
REMARK 500    ASP B 636      -85.13   -111.90
REMARK 500    ASN B 668      -66.56   -101.60
REMARK 500    ASP B 697     -161.37   -100.82
REMARK 500    ILE B 700       56.33     34.37
REMARK 500    ALA C   2       55.02     72.42
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: PLANAR GROUPS
REMARK 500
REMARK 500 PLANAR GROUPS IN THE FOLLOWING RESIDUES HAVE A TOTAL
REMARK 500 RMS DISTANCE OF ALL ATOMS FROM THE BEST-FIT PLANE
REMARK 500 BY MORE THAN AN EXPECTED VALUE OF 6*RMSD, WITH AN
REMARK 500 RMSD 0.02 ANGSTROMS, OR AT LEAST ONE ATOM HAS
REMARK 500 AN RMSD GREATER THAN THIS VALUE
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500  M RES CSSEQI        RMS     TYPE
REMARK 500    ARG A 294         0.13    SIDE CHAIN
REMARK 500    ARG A 340         0.10    SIDE CHAIN
REMARK 500    ARG B 340         0.09    SIDE CHAIN
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A1564        DISTANCE =  5.83 ANGSTROMS
REMARK 525    HOH A1565        DISTANCE =  5.92 ANGSTROMS
REMARK 525    HOH A1566        DISTANCE =  6.16 ANGSTROMS
REMARK 525    HOH A1567        DISTANCE =  6.47 ANGSTROMS
REMARK 525    HOH A1568        DISTANCE =  6.48 ANGSTROMS
REMARK 525    HOH A1569        DISTANCE =  6.52 ANGSTROMS
REMARK 525    HOH A1570        DISTANCE =  6.94 ANGSTROMS
REMARK 525    HOH A1571        DISTANCE =  7.43 ANGSTROMS
REMARK 525    HOH A1572        DISTANCE =  8.65 ANGSTROMS
REMARK 525    HOH A1573        DISTANCE =  9.06 ANGSTROMS
REMARK 525    HOH B1563        DISTANCE =  5.84 ANGSTROMS
REMARK 525    HOH B1564        DISTANCE =  6.65 ANGSTROMS
REMARK 525    HOH B1565        DISTANCE =  8.70 ANGSTROMS
REMARK 525    HOH B1566        DISTANCE =  8.77 ANGSTROMS
DBREF  9LBT A    2   723  UNP    P27487   DPP4_HUMAN      41    765
DBREF  9LBT B    2   723  UNP    P27487   DPP4_HUMAN      41    765
DBREF  9LBT C    1     4  PDB    9LBT     9LBT             1      4
DBREF  9LBT D    1     4  PDB    9LBT     9LBT             1      4
SEQADV 9LBT ALA A    1  UNP  P27487              EXPRESSION TAG
SEQADV 9LBT ALA B    1  UNP  P27487              EXPRESSION TAG
SEQRES   1 A  726  ALA LYS THR TYR THR LEU THR ASP TYR LEU LYS ASN THR
SEQRES   2 A  726  TYR ARG LEU LYS LEU TYR SER LEU ARG TRP ILE SER ASP
SEQRES   3 A  726  HIS GLU TYR LEU TYR LYS GLN GLU ASN ASN ILE LEU VAL
SEQRES   4 A  726  PHE ASN ALA GLU TYR GLY ASN SER SER VAL PHE LEU GLU
SEQRES   5 A  726  ASN SER THR PHE ASP GLU PHE GLY HIS SER ILE ASN ASP
SEQRES   6 A  726  TYR SER ILE SER PRO ASP GLY GLN PHE ILE LEU LEU GLU
SEQRES   7 A  726  TYR ASN TYR VAL LYS GLN TRP ARG HIS SER TYR THR ALA
SEQRES   8 A  726  SER TYR ASP ILE TYR ASP LEU ASN LYS ARG GLN LEU ILE
SEQRES   9 A  726  THR GLU GLU ARG ILE PRO ASN ASN THR GLN TRP VAL THR
SEQRES  10 A  726  TRP SER PRO VAL GLY HIS LYS LEU ALA TYR VAL TRP ASN
SEQRES  11 A  726  ASN ASP ILE TYR VAL LYS ILE GLU PRO ASN LEU PRO SER
SEQRES  12 A  726  TYR ARG ILE THR TRP THR GLY LYS GLU ASP ILE ILE TYR
SEQRES  13 A  726  ASN GLY ILE THR ASP TRP VAL TYR GLU GLU GLU VAL PHE
SEQRES  14 A  726  SER ALA TYR SER ALA LEU TRP TRP SER PRO ASN GLY THR
SEQRES  15 A  726  PHE LEU ALA TYR ALA GLN PHE ASN ASP THR GLU VAL PRO
SEQRES  16 A  726  LEU ILE GLU TYR SER PHE TYR SER ASP GLU SER LEU GLN
SEQRES  17 A  726  TYR PRO LYS THR VAL ARG VAL PRO TYR PRO LYS ALA GLY
SEQRES  18 A  726  ALA VAL ASN PRO THR VAL LYS PHE PHE VAL VAL ASN THR
SEQRES  19 A  726  ASP SER LEU SER SER VAL THR ASN ALA THR SER ILE GLN
SEQRES  20 A  726  ILE THR ALA PRO ALA SER MET LEU ILE GLY ASP HIS TYR
SEQRES  21 A  726  LEU CYS ASP VAL THR TRP ALA THR GLN GLU ARG ILE SER
SEQRES  22 A  726  LEU GLN TRP LEU ARG ARG ILE GLN ASN TYR SER VAL MET
SEQRES  23 A  726  ASP ILE CYS ASP TYR ASP GLU SER SER GLY ARG TRP ASN
SEQRES  24 A  726  CYS LEU VAL ALA ARG GLN HIS ILE GLU MET SER THR THR
SEQRES  25 A  726  GLY TRP VAL GLY ARG PHE ARG PRO SER GLU PRO HIS PHE
SEQRES  26 A  726  THR LEU ASP GLY ASN SER PHE TYR LYS ILE ILE SER ASN
SEQRES  27 A  726  GLU GLU GLY TYR ARG HIS ILE CYS TYR PHE GLN ILE ASP
SEQRES  28 A  726  LYS LYS ASP CYS THR PHE ILE THR LYS GLY THR TRP GLU
SEQRES  29 A  726  VAL ILE GLY ILE GLU ALA LEU THR SER ASP TYR LEU TYR
SEQRES  30 A  726  TYR ILE SER ASN GLU TYR LYS GLY MET PRO GLY GLY ARG
SEQRES  31 A  726  ASN LEU TYR LYS ILE GLN LEU SER ASP TYR THR LYS VAL
SEQRES  32 A  726  THR CYS LEU SER CYS GLU LEU ASN PRO GLU ARG CYS GLN
SEQRES  33 A  726  TYR TYR SER VAL SER PHE SER LYS GLU ALA LYS TYR TYR
SEQRES  34 A  726  GLN LEU ARG CYS SER GLY PRO GLY LEU PRO LEU TYR THR
SEQRES  35 A  726  LEU HIS SER SER VAL ASN ASP LYS GLY LEU ARG VAL LEU
SEQRES  36 A  726  GLU ASP ASN SER ALA LEU ASP LYS MET LEU GLN ASN VAL
SEQRES  37 A  726  GLN MET PRO SER LYS LYS LEU ASP PHE ILE ILE LEU ASN
SEQRES  38 A  726  GLU THR LYS PHE TRP TYR GLN MET ILE LEU PRO PRO HIS
SEQRES  39 A  726  PHE ASP LYS SER LYS LYS TYR PRO LEU LEU LEU ASP VAL
SEQRES  40 A  726  TYR ALA GLY PRO CYS SER GLN LYS ALA ASP THR VAL PHE
SEQRES  41 A  726  ARG LEU ASN TRP ALA THR TYR LEU ALA SER THR GLU ASN
SEQRES  42 A  726  ILE ILE VAL ALA SER PHE ASP GLY ARG GLY SER GLY TYR
SEQRES  43 A  726  GLN GLY ASP LYS ILE MET HIS ALA ILE ASN ARG ARG LEU
SEQRES  44 A  726  GLY THR PHE GLU VAL GLU ASP GLN ILE GLU ALA ALA ARG
SEQRES  45 A  726  GLN PHE SER LYS MET GLY PHE VAL ASP ASN LYS ARG ILE
SEQRES  46 A  726  ALA ILE TRP GLY TRP SER TYR GLY GLY TYR VAL THR SER
SEQRES  47 A  726  MET VAL LEU GLY SER GLY SER GLY VAL PHE LYS CYS GLY
SEQRES  48 A  726  ILE ALA VAL ALA PRO VAL SER ARG TRP GLU TYR TYR ASP
SEQRES  49 A  726  SER VAL TYR THR GLU ARG TYR MET GLY LEU PRO THR PRO
SEQRES  50 A  726  GLU ASP ASN LEU ASP HIS TYR ARG ASN SER THR VAL MET
SEQRES  51 A  726  SER ARG ALA GLU ASN PHE LYS GLN VAL GLU TYR LEU LEU
SEQRES  52 A  726  ILE HIS GLY THR ALA ASP ASP ASN VAL HIS PHE GLN GLN
SEQRES  53 A  726  SER ALA GLN ILE SER LYS ALA LEU VAL ASP VAL GLY VAL
SEQRES  54 A  726  ASP PHE GLN ALA MET TRP TYR THR ASP GLU ASP HIS GLY
SEQRES  55 A  726  ILE ALA SER SER THR ALA HIS GLN HIS ILE TYR THR HIS
SEQRES  56 A  726  MET SER HIS PHE ILE LYS GLN CYS PHE SER LEU
SEQRES   1 B  726  ALA LYS THR TYR THR LEU THR ASP TYR LEU LYS ASN THR
SEQRES   2 B  726  TYR ARG LEU LYS LEU TYR SER LEU ARG TRP ILE SER ASP
SEQRES   3 B  726  HIS GLU TYR LEU TYR LYS GLN GLU ASN ASN ILE LEU VAL
SEQRES   4 B  726  PHE ASN ALA GLU TYR GLY ASN SER SER VAL PHE LEU GLU
SEQRES   5 B  726  ASN SER THR PHE ASP GLU PHE GLY HIS SER ILE ASN ASP
SEQRES   6 B  726  TYR SER ILE SER PRO ASP GLY GLN PHE ILE LEU LEU GLU
SEQRES   7 B  726  TYR ASN TYR VAL LYS GLN TRP ARG HIS SER TYR THR ALA
SEQRES   8 B  726  SER TYR ASP ILE TYR ASP LEU ASN LYS ARG GLN LEU ILE
SEQRES   9 B  726  THR GLU GLU ARG ILE PRO ASN ASN THR GLN TRP VAL THR
SEQRES  10 B  726  TRP SER PRO VAL GLY HIS LYS LEU ALA TYR VAL TRP ASN
SEQRES  11 B  726  ASN ASP ILE TYR VAL LYS ILE GLU PRO ASN LEU PRO SER
SEQRES  12 B  726  TYR ARG ILE THR TRP THR GLY LYS GLU ASP ILE ILE TYR
SEQRES  13 B  726  ASN GLY ILE THR ASP TRP VAL TYR GLU GLU GLU VAL PHE
SEQRES  14 B  726  SER ALA TYR SER ALA LEU TRP TRP SER PRO ASN GLY THR
SEQRES  15 B  726  PHE LEU ALA TYR ALA GLN PHE ASN ASP THR GLU VAL PRO
SEQRES  16 B  726  LEU ILE GLU TYR SER PHE TYR SER ASP GLU SER LEU GLN
SEQRES  17 B  726  TYR PRO LYS THR VAL ARG VAL PRO TYR PRO LYS ALA GLY
SEQRES  18 B  726  ALA VAL ASN PRO THR VAL LYS PHE PHE VAL VAL ASN THR
SEQRES  19 B  726  ASP SER LEU SER SER VAL THR ASN ALA THR SER ILE GLN
SEQRES  20 B  726  ILE THR ALA PRO ALA SER MET LEU ILE GLY ASP HIS TYR
SEQRES  21 B  726  LEU CYS ASP VAL THR TRP ALA THR GLN GLU ARG ILE SER
SEQRES  22 B  726  LEU GLN TRP LEU ARG ARG ILE GLN ASN TYR SER VAL MET
SEQRES  23 B  726  ASP ILE CYS ASP TYR ASP GLU SER SER GLY ARG TRP ASN
SEQRES  24 B  726  CYS LEU VAL ALA ARG GLN HIS ILE GLU MET SER THR THR
SEQRES  25 B  726  GLY TRP VAL GLY ARG PHE ARG PRO SER GLU PRO HIS PHE
SEQRES  26 B  726  THR LEU ASP GLY ASN SER PHE TYR LYS ILE ILE SER ASN
SEQRES  27 B  726  GLU GLU GLY TYR ARG HIS ILE CYS TYR PHE GLN ILE ASP
SEQRES  28 B  726  LYS LYS ASP CYS THR PHE ILE THR LYS GLY THR TRP GLU
SEQRES  29 B  726  VAL ILE GLY ILE GLU ALA LEU THR SER ASP TYR LEU TYR
SEQRES  30 B  726  TYR ILE SER ASN GLU TYR LYS GLY MET PRO GLY GLY ARG
SEQRES  31 B  726  ASN LEU TYR LYS ILE GLN LEU SER ASP TYR THR LYS VAL
SEQRES  32 B  726  THR CYS LEU SER CYS GLU LEU ASN PRO GLU ARG CYS GLN
SEQRES  33 B  726  TYR TYR SER VAL SER PHE SER LYS GLU ALA LYS TYR TYR
SEQRES  34 B  726  GLN LEU ARG CYS SER GLY PRO GLY LEU PRO LEU TYR THR
SEQRES  35 B  726  LEU HIS SER SER VAL ASN ASP LYS GLY LEU ARG VAL LEU
SEQRES  36 B  726  GLU ASP ASN SER ALA LEU ASP LYS MET LEU GLN ASN VAL
SEQRES  37 B  726  GLN MET PRO SER LYS LYS LEU ASP PHE ILE ILE LEU ASN
SEQRES  38 B  726  GLU THR LYS PHE TRP TYR GLN MET ILE LEU PRO PRO HIS
SEQRES  39 B  726  PHE ASP LYS SER LYS LYS TYR PRO LEU LEU LEU ASP VAL
SEQRES  40 B  726  TYR ALA GLY PRO CYS SER GLN LYS ALA ASP THR VAL PHE
SEQRES  41 B  726  ARG LEU ASN TRP ALA THR TYR LEU ALA SER THR GLU ASN
SEQRES  42 B  726  ILE ILE VAL ALA SER PHE ASP GLY ARG GLY SER GLY TYR
SEQRES  43 B  726  GLN GLY ASP LYS ILE MET HIS ALA ILE ASN ARG ARG LEU
SEQRES  44 B  726  GLY THR PHE GLU VAL GLU ASP GLN ILE GLU ALA ALA ARG
SEQRES  45 B  726  GLN PHE SER LYS MET GLY PHE VAL ASP ASN LYS ARG ILE
SEQRES  46 B  726  ALA ILE TRP GLY TRP SER TYR GLY GLY TYR VAL THR SER
SEQRES  47 B  726  MET VAL LEU GLY SER GLY SER GLY VAL PHE LYS CYS GLY
SEQRES  48 B  726  ILE ALA VAL ALA PRO VAL SER ARG TRP GLU TYR TYR ASP
SEQRES  49 B  726  SER VAL TYR THR GLU ARG TYR MET GLY LEU PRO THR PRO
SEQRES  50 B  726  GLU ASP ASN LEU ASP HIS TYR ARG ASN SER THR VAL MET
SEQRES  51 B  726  SER ARG ALA GLU ASN PHE LYS GLN VAL GLU TYR LEU LEU
SEQRES  52 B  726  ILE HIS GLY THR ALA ASP ASP ASN VAL HIS PHE GLN GLN
SEQRES  53 B  726  SER ALA GLN ILE SER LYS ALA LEU VAL ASP VAL GLY VAL
SEQRES  54 B  726  ASP PHE GLN ALA MET TRP TYR THR ASP GLU ASP HIS GLY
SEQRES  55 B  726  ILE ALA SER SER THR ALA HIS GLN HIS ILE TYR THR HIS
SEQRES  56 B  726  MET SER HIS PHE ILE LYS GLN CYS PHE SER LEU
SEQRES   1 C    4  VAL ALA MET PRO
SEQRES   1 D    4  VAL ALA MET PRO
FORMUL   5  HOH   *1543(H2 O)
HELIX    1 AA1 THR A    5  ASN A   12  1                                   8
HELIX    2 AA2 ASP A  158  VAL A  165  1                                   8
HELIX    3 AA3 PRO A  248  ILE A  253  1                                   6
HELIX    4 AA4 VAL A  299  GLN A  302  5                                   4
HELIX    5 AA5 GLU A  379  MET A  383  5                                   5
HELIX    6 AA6 ASN A  455  ASN A  464  1                                  10
HELIX    7 AA7 ASN A  520  THR A  528  1                                   9
HELIX    8 AA8 GLY A  545  HIS A  550  1                                   6
HELIX    9 AA9 ALA A  551  ASN A  553  5                                   3
HELIX   10 AB1 THR A  558  SER A  572  1                                  15
HELIX   11 AB2 SER A  588  GLY A  599  1                                  12
HELIX   12 AB3 ARG A  616  TYR A  620  5                                   5
HELIX   13 AB4 ASP A  621  GLY A  630  1                                  10
HELIX   14 AB5 ASN A  637  SER A  644  1                                   8
HELIX   15 AB6 VAL A  646  VAL A  656  5                                  11
HELIX   16 AB7 HIS A  670  GLY A  685  1                                  16
HELIX   17 AB8 SER A  702  PHE A  721  1                                  20
HELIX   18 AB9 THR B    5  ASN B   12  1                                   8
HELIX   19 AC1 PHE B   53  GLY B   57  5                                   5
HELIX   20 AC2 ASP B  158  VAL B  165  1                                   8
HELIX   21 AC3 ASP B  232  LEU B  234  5                                   3
HELIX   22 AC4 PRO B  248  ILE B  253  1                                   6
HELIX   23 AC5 VAL B  299  GLN B  302  5                                   4
HELIX   24 AC6 GLU B  379  MET B  383  5                                   5
HELIX   25 AC7 ASN B  455  ASN B  464  1                                  10
HELIX   26 AC8 ASN B  520  THR B  528  1                                   9
HELIX   27 AC9 GLY B  545  HIS B  550  1                                   6
HELIX   28 AD1 ALA B  551  ASN B  553  5                                   3
HELIX   29 AD2 THR B  558  MET B  574  1                                  17
HELIX   30 AD3 SER B  588  GLY B  599  1                                  12
HELIX   31 AD4 ARG B  616  TYR B  620  5                                   5
HELIX   32 AD5 ASP B  621  GLY B  630  1                                  10
HELIX   33 AD6 ASN B  637  SER B  644  1                                   8
HELIX   34 AD7 VAL B  646  VAL B  656  5                                  11
HELIX   35 AD8 HIS B  670  VAL B  684  1                                  15
HELIX   36 AD9 SER B  702  PHE B  721  1                                  20
SHEET    1 AA1 2 LYS A   2  THR A   3  0
SHEET    2 AA1 2 VAL A 465  GLN A 466  1  O  GLN A 466   N  LYS A   2
SHEET    1 AA2 4 ARG A  22  TRP A  23  0
SHEET    2 AA2 4 GLU A  28  TYR A  31 -1  O  LEU A  30   N  ARG A  22
SHEET    3 AA2 4 ILE A  34  ASN A  38 -1  O  LEU A  35   N  TYR A  31
SHEET    4 AA2 4 SER A  44  LEU A  48 -1  O  PHE A  47   N  ILE A  34
SHEET    1 AA3 4 ILE A  60  ILE A  65  0
SHEET    2 AA3 4 PHE A  71  LYS A  80 -1  O  LEU A  73   N  SER A  64
SHEET    3 AA3 4 TYR A  86  ASP A  94 -1  O  ASP A  91   N  LEU A  74
SHEET    4 AA3 4 GLN A  99  LEU A 100 -1  O  GLN A  99   N  ASP A  94
SHEET    1 AA4 4 TRP A 112  TRP A 115  0
SHEET    2 AA4 4 LEU A 122  TRP A 126 -1  O  VAL A 125   N  TRP A 112
SHEET    3 AA4 4 ASP A 129  LYS A 133 -1  O  TYR A 131   N  TYR A 124
SHEET    4 AA4 4 TYR A 141  ARG A 142 -1  O  TYR A 141   N  VAL A 132
SHEET    1 AA5 3 ILE A 152  ASN A 154  0
SHEET    2 AA5 3 PHE A 180  ASN A 187 -1  O  PHE A 186   N  TYR A 153
SHEET    3 AA5 3 LEU A 172  TRP A 174 -1  N  TRP A 173   O  ALA A 182
SHEET    1 AA6 4 ILE A 152  ASN A 154  0
SHEET    2 AA6 4 PHE A 180  ASN A 187 -1  O  PHE A 186   N  TYR A 153
SHEET    3 AA6 4 THR A 223  ASN A 230 -1  O  PHE A 227   N  TYR A 183
SHEET    4 AA6 4 ILE A 243  ILE A 245 -1  O  ILE A 243   N  VAL A 228
SHEET    1 AA7 2 LEU A 193  PHE A 198  0
SHEET    2 AA7 2 LYS A 208  PRO A 213 -1  O  VAL A 210   N  TYR A 196
SHEET    1 AA8 4 HIS A 256  THR A 265  0
SHEET    2 AA8 4 ARG A 268  ARG A 275 -1  O  ARG A 268   N  ALA A 264
SHEET    3 AA8 4 TYR A 280  ASP A 289 -1  O  CYS A 286   N  ILE A 269
SHEET    4 AA8 4 ARG A 294  CYS A 297 -1  O  ARG A 294   N  ASP A 289
SHEET    1 AA9 4 HIS A 256  THR A 265  0
SHEET    2 AA9 4 ARG A 268  ARG A 275 -1  O  ARG A 268   N  ALA A 264
SHEET    3 AA9 4 TYR A 280  ASP A 289 -1  O  CYS A 286   N  ILE A 269
SHEET    4 AA9 4 HIS A 303  MET A 306 -1  O  HIS A 303   N  MET A 283
SHEET    1 AB1 4 HIS A 321  PHE A 322  0
SHEET    2 AB1 4 SER A 328  SER A 334 -1  O  TYR A 330   N  HIS A 321
SHEET    3 AB1 4 ARG A 340  GLN A 346 -1  O  PHE A 345   N  PHE A 329
SHEET    4 AB1 4 THR A 353  PHE A 354 -1  O  THR A 353   N  TYR A 344
SHEET    1 AB2 4 VAL A 362  LEU A 368  0
SHEET    2 AB2 4 TYR A 372  SER A 377 -1  O  TYR A 374   N  ALA A 367
SHEET    3 AB2 4 ASN A 388  GLN A 393 -1  O  TYR A 390   N  TYR A 375
SHEET    4 AB2 4 VAL A 400  CYS A 402 -1  O  THR A 401   N  LYS A 391
SHEET    1 AB3 4 CYS A 412  PHE A 419  0
SHEET    2 AB3 4 TYR A 425  PRO A 433 -1  O  GLY A 432   N  GLN A 413
SHEET    3 AB3 4 LEU A 437  SER A 442 -1  O  HIS A 441   N  TYR A 426
SHEET    4 AB3 4 LYS A 447  GLU A 453 -1  O  GLU A 453   N  TYR A 438
SHEET    1 AB4 8 SER A 469  LEU A 477  0
SHEET    2 AB4 8 THR A 480  LEU A 488 -1  O  PHE A 482   N  ILE A 475
SHEET    3 AB4 8 ILE A 532  PHE A 536 -1  O  SER A 535   N  GLN A 485
SHEET    4 AB4 8 TYR A 498  VAL A 504  1  N  LEU A 501   O  ILE A 532
SHEET    5 AB4 8 VAL A 577  TRP A 587  1  O  ASP A 578   N  TYR A 498
SHEET    6 AB4 8 CYS A 607  VAL A 611  1  O  VAL A 611   N  GLY A 586
SHEET    7 AB4 8 GLU A 657  GLY A 663  1  O  ILE A 661   N  ALA A 610
SHEET    8 AB4 8 GLN A 689  TYR A 693  1  O  GLN A 689   N  TYR A 658
SHEET    1 AB5 2 LYS B   2  THR B   3  0
SHEET    2 AB5 2 VAL B 465  GLN B 466  1  O  GLN B 466   N  LYS B   2
SHEET    1 AB6 4 ARG B  22  TRP B  23  0
SHEET    2 AB6 4 GLU B  28  TYR B  31 -1  O  LEU B  30   N  ARG B  22
SHEET    3 AB6 4 ILE B  34  ASN B  38 -1  O  PHE B  37   N  TYR B  29
SHEET    4 AB6 4 SER B  44  LEU B  48 -1  O  LEU B  48   N  ILE B  34
SHEET    1 AB7 4 ILE B  60  ILE B  65  0
SHEET    2 AB7 4 PHE B  71  LYS B  80 -1  O  LEU B  73   N  SER B  64
SHEET    3 AB7 4 TYR B  86  ASP B  94 -1  O  SER B  89   N  TYR B  76
SHEET    4 AB7 4 GLN B  99  LEU B 100 -1  O  GLN B  99   N  ASP B  94
SHEET    1 AB8 4 THR B 110  TRP B 115  0
SHEET    2 AB8 4 LEU B 122  TRP B 126 -1  O  VAL B 125   N  GLN B 111
SHEET    3 AB8 4 ASP B 129  LYS B 133 -1  O  LYS B 133   N  LEU B 122
SHEET    4 AB8 4 TYR B 141  ARG B 142 -1  O  TYR B 141   N  VAL B 132
SHEET    1 AB9 3 ILE B 152  ASN B 154  0
SHEET    2 AB9 3 PHE B 180  ASN B 187 -1  O  PHE B 186   N  TYR B 153
SHEET    3 AB9 3 LEU B 172  TRP B 174 -1  N  TRP B 173   O  ALA B 182
SHEET    1 AC1 4 ILE B 152  ASN B 154  0
SHEET    2 AC1 4 PHE B 180  ASN B 187 -1  O  PHE B 186   N  TYR B 153
SHEET    3 AC1 4 THR B 223  ASN B 230 -1  O  PHE B 227   N  TYR B 183
SHEET    4 AC1 4 SER B 242  GLN B 244 -1  O  ILE B 243   N  VAL B 228
SHEET    1 AC2 2 LEU B 193  PHE B 198  0
SHEET    2 AC2 2 LYS B 208  PRO B 213 -1  O  LYS B 208   N  PHE B 198
SHEET    1 AC3 4 HIS B 256  THR B 265  0
SHEET    2 AC3 4 ARG B 268  ARG B 275 -1  O  ARG B 268   N  ALA B 264
SHEET    3 AC3 4 TYR B 280  ASP B 289 -1  O  CYS B 286   N  ILE B 269
SHEET    4 AC3 4 ARG B 294  CYS B 297 -1  O  ARG B 294   N  ASP B 289
SHEET    1 AC4 4 HIS B 256  THR B 265  0
SHEET    2 AC4 4 ARG B 268  ARG B 275 -1  O  ARG B 268   N  ALA B 264
SHEET    3 AC4 4 TYR B 280  ASP B 289 -1  O  CYS B 286   N  ILE B 269
SHEET    4 AC4 4 HIS B 303  MET B 306 -1  O  HIS B 303   N  MET B 283
SHEET    1 AC5 4 HIS B 321  PHE B 322  0
SHEET    2 AC5 4 SER B 328  SER B 334 -1  O  TYR B 330   N  HIS B 321
SHEET    3 AC5 4 ARG B 340  GLN B 346 -1  O  PHE B 345   N  PHE B 329
SHEET    4 AC5 4 THR B 353  PHE B 354 -1  O  THR B 353   N  TYR B 344
SHEET    1 AC6 4 VAL B 362  LEU B 368  0
SHEET    2 AC6 4 TYR B 372  SER B 377 -1  O  TYR B 374   N  ALA B 367
SHEET    3 AC6 4 ASN B 388  GLN B 393 -1  O  TYR B 390   N  TYR B 375
SHEET    4 AC6 4 ASP B 396  CYS B 402 -1  O  THR B 401   N  LYS B 391
SHEET    1 AC7 4 TYR B 415  PHE B 419  0
SHEET    2 AC7 4 TYR B 425  CYS B 430 -1  O  GLN B 427   N  SER B 418
SHEET    3 AC7 4 LEU B 437  SER B 442 -1  O  LEU B 437   N  CYS B 430
SHEET    4 AC7 4 GLY B 448  GLU B 453 -1  O  LEU B 449   N  LEU B 440
SHEET    1 AC8 8 SER B 469  LEU B 477  0
SHEET    2 AC8 8 THR B 480  LEU B 488 -1  O  THR B 480   N  LEU B 477
SHEET    3 AC8 8 ILE B 532  PHE B 536 -1  O  VAL B 533   N  ILE B 487
SHEET    4 AC8 8 TYR B 498  VAL B 504  1  N  ASP B 503   O  ALA B 534
SHEET    5 AC8 8 VAL B 577  TRP B 587  1  O  TRP B 585   N  VAL B 504
SHEET    6 AC8 8 CYS B 607  VAL B 611  1  O  VAL B 611   N  GLY B 586
SHEET    7 AC8 8 GLU B 657  GLY B 663  1  O  LEU B 659   N  ALA B 610
SHEET    8 AC8 8 GLN B 689  TYR B 693  1  O  GLN B 689   N  TYR B 658
SSBOND   1 CYS A  286    CYS A  297                          1555   1555  2.08
SSBOND   2 CYS A  343    CYS A  352                          1555   1555  2.09
SSBOND   3 CYS A  402    CYS A  405                          1555   1555  2.03
SSBOND   4 CYS A  412    CYS A  430                          1555   1555  2.09
SSBOND   5 CYS A  607    CYS A  720                          1555   1555  2.11
SSBOND   6 CYS B  286    CYS B  297                          1555   1555  2.09
SSBOND   7 CYS B  343    CYS B  352                          1555   1555  2.06
SSBOND   8 CYS B  402    CYS B  405                          1555   1555  2.06
SSBOND   9 CYS B  412    CYS B  430                          1555   1555  2.11
SSBOND  10 CYS B  607    CYS B  720                          1555   1555  2.08
CISPEP   1 GLY A  432    PRO A  433          0         8.38
CISPEP   2 GLY B  432    PRO B  433          0         6.16
CRYST1  117.827  126.370  136.867  90.00  90.00  90.00 P 21 21 21    8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008487  0.000000  0.000000        0.00000
SCALE2      0.000000  0.007913  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007306        0.00000
TER    5918      LEU A 723
TER   11836      LEU B 723
TER   11864      PRO C   4
TER   11892      PRO D   4
MASTER      544    0    0   36  102    0    0    613431    4   20  114
END